Professional Documents
Culture Documents
ISBN 971-22-0020-5
Foreword
Farmers everywhere search for ways to widen the narrow margin of profit
between production costs and crop returns. Weeds cause more yield losses in
ricefields than any other pest. Cost-effective methods for controlling weeds
could help preserve profits and increase yields.
The authors have designed this book to provide practical information on
controlling weeds in the many different rice cultural systems, using an inte-
grated management approach. It is exceptionally comprehensive, and can be
used as a textbook, as a field guide, and as a manual for making decisions in
crop management.
The handbook has been especially designed to facilitate inexpensive trans-
lation and copublication. Contact Communication and Publications Services,
IRRI Information Center, for permission and assistance, at no charge, in any
activities to extend its benefits to rice workers who read languages other than
English.
Klaus Lampe
Director General
Preface
Kwesi Ampong-Nyarko
S. K. De Datta
IRRI July 1990
Contents
Significance of weeds
in rice farming
Weeds are plants growing where Table 1.1. Cultivated species of rice evolved from weedy wild species (adapted from Chang
1976).
they are not wanted. A weed in one
place may be a useful food, feed, or South and Southeast Asia Tropical Africa
medicine in another. Thus, a plant
Wild perennial O. rufipogon Griff. O. longistaminata A. Chev. & Roehr.
species cannot be classified as a weed Wild annual O. nivara Sharma & Shastry O. barthii A. Chev.
under all conditions. Many plants, Cultivated annual O. sativa O. glaberrima
however, are classified as weeds
everywhere they occur, because they
commonly grow on regularly tilled Table 1.2. Yield losses due to uncontrolled after weed control. These losses can
weed growth in different types of rice culture in
areas such as ricefields. the Philippines (sources: IRRI 1977 to 1988).
amount to 46 million t (based on 1987
Many weeds co-evolved with world rough rice production). There
crops and, in some cases, were Yield Experi- is considerable variation in yield loss
Type of rice culture loss ments
ancestors of cultivated plant species. (mean %) (no.)
to weeds among countries.
For example, the wild rices Oryza There is a need to improve farm-
barthii and O. longistaminata are lrrigated ers’ weed control practices. Improved
Transplanted 48 42
ancestors of cultivated O. glaberrima Water seeded 44 1 weed management will contribute
in Africa. The wild rices O. rufipogon Direct seeded 55 28 significantly to future gains in rice
and O. nivara are ancestors of Rainfed lowland yield in many countries.
cultivated O. sativa in Asia Direct seeded (dry seeds) 74 11
Direct seeded on 61 7
(Table 1.1). puddled soil
Increase in rice production costs
The prevalent weeds in ricefields Transplanted in puddled 51 9 The cost of rice weed control, includ-
are often legacies of previous years’ soil ing herbicides, cultural and mechani-
crops—seeds, rhizomes, tubers, and Rainfed upland cal practices, and hand weeding, is
Broadcast or drilled 96 16
bulbs surviving in the soil. The weed estimated to be about 5% of world
flora in a ricefield is greatly influ- rice production and amount to
enced by the rice culture practiced. US$3.5 billion annually. When the
Continuous rice with an unchanged Effects of weeds 10% loss of rough rice grain yield is
cultural system encourages the Weed infestations primarily constrain added to this cost, the world’s total
buildup of weeds adapted to that rice production by reducing grain estimated cost for rice weeds and
system. In contrast, where crop yield. Yield reductions caused by their control amounts to 15% of total
rotation is practiced, a diverse weed uncontrolled weed growth through- annual production—valued at
flora will result. Perennial weeds out a crop season have been esti- US$10.5 billion (based on 1987
increase in nontilled ricefields. mated to be from 44 to 96%, depend- average export prices at Bangkok,
ing on the rice culture (Table 1.2). In Thailand, for 5% brokens white rice
practice, almost all farmers control US$230/t, IRRI 1988b).
weeds in their ricefields. Worldwide,
some 10% loss of rice yield can be
attributed just to weeds that grow
Significance of weeds 1
Table 1.3. Weeds as secondary hosts for diseases, insects, and nematodes of rice. weeding, by hand or with simple
Disease/insect Host weeds Reference tools, than on any other farming task.
Hand weeding 1 ha of rice requires
Rice dwarf disease (vlrus) Echinochloa crus-galli Ou (1985) from 100 to 780 labor-hours per crop,
Rice stripe disease (virus) E. crus-gall; Ou (1985)
Cynodon dactylon depending on the rice culture.
Setaria vindis
Digitaria adscendens Aquatic weed problems
Rice yellow dwarf (virus) Paspalum distichum Ou (1985)
Leptochloa chinensis Nutrient availability and favorable
Leersia hexandra temperatures throughout the year,
lmperata cylindrica especially in the tropics, allow luxuri-
Rice hoja blanca (virus) Leptochloa spp. Ou (1985)
Digitaria spp. ant aquatic weed growth in flooded
Bacterial leaf blight L. chinensis Ou (1985) ricefields. Noxious aquatic weeds
(Xanthomonas campestris pv. oryzae) have increasingly infested
Brown spot C. dactylon
(Cochliobolus miyabeanus) L. hexandra Ou (1985) impounded water in the tropics.
Digitaria sanguinalis Heavy aquatic weed infestation
Whlte tip S. viridis Ou (1985) causes excessive water loss through
(Aphelenchoides oryzae) Cyperus iria
I. cylindrica evaporation and impedes water flow
Meloidogyne (nematodes) Fimbristylis miliacea Ou (1985) in irrigation canals. In some cases,
Echinochloa colona aquatic weeds may be a health
Rlce grassy stunt virus L. hexandra IRRI (1988a)
(transmltted by brown C. dactylon hazard to persons living near
planthopper Nilaparvata lugens) Cyperus rotundus impounded water. For example, the
E. colona association of the weed Ceratophyllum
Monochoria vaginalis
Rice tungro assoclated viruses C. rotundus IRRI (1988a) demersum with the intermediate host
F. miliacea snail of schistosomiasis (bilharzia)
Oryza longistaminata Bulinus sp. is well documented.
Oryza barthii
Nymphula depunctalis (Avenee) L. chinensis IRRI (1986)
(caseworm) L. hexandra
lschaemum rugosum Rice-weed competition
Weeds interfere with rice growth by
competing for one or more growth-
Weeds as secondary Effects on harvesting and limiting resources, such as light,
hosts for pests grain quality nutrients, and water. Allelopathy
Weeds indirectly limit production by Weeds hamper rice harvesting and (chemical production by living or
serving as hosts for organisms that increase harvest costs through direct decaying weed plant tissues) may
adversely affect rice. Weeds provide interference with the harvesting also adversely affect the growth of a
food, shelter, and reproduction sites operation and by causing lodging. neighboring rice plant.
for insects, nematodes, pathogens, Weed seeds contaminate rough rice, Rice and rice weeds have similar
and rodents. Table 1.3 lists weeds thus reducing grain quality and requirements for growth and devel-
that serve as alternate hosts to rice market value. For example, the weed opment. Competition occurs when
pests. This indicates the importance red rice has a pigmented layer that one of the limiting resources falls
of recognizing weeds as secondary shatters easily and readily short of the combined requirements
hosts for pests and of removing contaminates rough rice. Removing of both. The degree of rice-weed
weeds from the margins of ricefields all traces of the pigmented layer competition depends on rainfall, rice
to prevent continued infection of the requires intense milling and results in variety, soil factors, weed density,
rice crop. decreased grain quality and lower duration of rice and weed growth,
milling rates. crop age when weeds started to
compete, and nutrient resources,
Social costs of weed control among other variables.
The drudgery of weeding and labor
shortages have made rice farming
unattractive. In most tropical coun-
tries, farmers spend more time on
~~
Significance of weeds 3
horizontal leaves. Tall plants have
an advantage over short plants.
Nutrients
Characteristics of
The three most common yield-
For example, when Rottboellia limiting nutrients are N, P, and K. successful weeds
cochinchinensis was allowed to grow Competition, however, may occur for Weeds and crops may have common
with rice, R. cochinchinensis was 150 any nutrient required for plant origins, and hence, may have similar
cm tall and rice was 50 cm tall at 8 wk growth. Many weed species have a characteristics. Weeds often have
after seeding. The amount of light nutrient uptake similar to that of rice, evolved adaptive traits that increase
received at 25 cm within the rice but have higher nutrient-use their persistence or competitiveness,
canopy was only 3% of the light at efficiency than rice. In general, factors or both. Traits contributing to
the top of the weed canopy. In this that give a plant a competitive successful weeds fall under three
situation, the weed clearly had an advantage in water uptake also give categories: 1) weed competitiveness
advantage. the plant a competitive advantage in and growth, 2) weed reproduction
nutrient uptake. and dispersal, and 3) similarities
Water between weed and rice.
Rice yield losses from water deficit Critical period of weed competition
depend on the severity and duration Competition between weed seedlings Weed competitiveness and growth
of the deficit, environmental condi- and direct seeded rice seedlings Some weeds have large seeds which
tions, cultivar, and growth stage usually begins as competition for allow for rapid seedling growth.
when drought stress occurs. Drought light. The effect of shading on rice Some weeds are climbers; others are
stress reduces photosynthesis by growth is most severe during the tall. Some weeds have higher rates of
reducing leaf expansion and causing seedling stage, but rice can recover photosynthesis, faster growth, larger
loss of leaf turgor, which leads to from weed competition if weeds are leaves, and deeper root systems than
stomatal closure. Transpiration and eliminated early in the season. Weeds rice and other crops. Deep root
photosynthesis by leaves are greatly will not further reduce the grain yield systems enable a weed to exploit
reduced when stomata close. of a mature rice crop ready for har- nutrients essential for crop growth.
Responses to drought stress include vesting. Weeds are highly adaptable to
wilting and increased leaf mortality. The so-called critical period lies changing environments (phenotypic
Rice and weeds differ in their between the seedling and harvest plasticity). Under favorable condi-
tolerance for drought because of stages, the 30-45 d when weed tions, weeds become large and
differences in their root distribution, competition is most damaging to rice. produce many seeds. Under unfavor-
root elongation rate, genetic tolerance To avoid grain yield losses, it is able conditions, weeds remain small
for low water availability in plant important to controI weeds through- and produce only a few seeds.
tissue, and control of water loss out this critical period. At 45- to
through transpiration. C4 weeds have 60-d-old, well-established, weed-free Reproduction and dispersal
lower water requirements than those rice plants are able to suppress later Many weeds produce large quantities
of C3 rice and are able to tolerate germinating weed seedlings. When of seed in favorable environments.
more drought stress than rice. herbicides are used, their persistence For example, Eckinochloa colona may
should be long enough to cover the produce 100,000 seeds/plant; Eleusine
critical period of competition. indica, 50,000 seeds/plant; Commelina
benghalensis, 1,600 seeds/plant;
Trianthema portulacastrum, 52,000
seeds/plant; and Amaranthus spp.,
196,000 seeds/plant. Production of a
large number of seeds ensure species
Significance of weeds 5
Chapter 2
National and regional methods of and agricultural importance are Stolons are horizontally growing
classifying rice cultures can vary described. Local names of weeds in stems with long slender internodes;
widely, depending on rice-growing the countries where they are of major adventitious roots form at the nodes
conditions and purposes of the classi- concern are provided to compliment when in contact with soil. Paspalum
fication system. A classification the photographs for weed identifica- distichum is a weed with stolons.
system may be based on general tion. (We know the list of local names A tuber is a specialized structure
surface hydrology, source of water, is not complete, and the spelling used that results from the swelling of the
landform and soil units, ecological for the local names may not be the one terminal portion of an underground
factors, or crop season. For weed preferred. Readers are invited to send stem or root; it contains stored food.
control, general surface hydrology the authors the correct local names.) Cyperus rotundus produces tubers.
and rice seeding method are more Weeds in rice are classified by their
important than other factors (De Datta life cycle, habitat, and morphological
1981). characteristics. Morphology
A classification based on surface Weeds are also classified as mono-
hydrology and seeding method is cotyledonous or dicotyledonous.
used here to discuss weed problems Life cycles
and the integrated methods available Weeds are classified as annual or Monocotyledons
for weed control in each type of rice perennial, or both. Where moisture or The seeds of monocotyledonous
culture. Rice culture is classified on temperature is not limiting and life weeds have a single cotyledon (seed
the basis of water management as cycle is short, an annual weed may leaf). A monocotyledon’s mature
lowland, upland, or deepwater; it may complete more than one life cycle in a leaves are long and narrow with
be irrigated or rainfed. This is further year. Annuals produce many seeds, parallel veins. The stem or culm is
subdivided on the basis of rice estab- some of which remain dormant and cylindrical and the growing point is
lishment method. buffer a species against weed control protected by a sheath. The root
This chapter covers weeds of measures. systems arise adventitiously and are
worldwide importance in lowland, Perennial weeds propagate by usually fibrous. Examples of families
upland, and deepwater ricefields. The vegetative structures such as bulbs, of monocotyledoneae are
weeds in each ecosystem are arranged corms, rhizomes, stolons, and tubers. Potamogetonaceae, Pontederiaceae,
alphabetically on the basis of family. A bulb is an underground bud. Cyperaceae, Poaceae, and
Photographs show seed, seedling, Rhizomes are underground shoots Commelinaceae. Sedges (Cyperaceae)
mature plant, and flower for each with short, thick internodes buried in resemble grasses but differ from
weed, and each weed’s characteristics the soil. They have specialized buds grasses in that their stems are
that can remain dormant. These unjointed, solid, and often triangular
shoots are rich in stored food and in cross section.
enable plants to survive from year to
year. Imperata cylindrica and Cynodon
dactylon are rhizomatous weeds.
Weeds worldwide 7
Dicotyledons For further information on the
The seeds of dicotyledonous weeds biology of the weeds listed in this
have two cotyledons. Mature leaves chapter, and on other weeds that may
are broad and usually net-veined. The be of interest, readers can refer to the
root systems have tap roots. The book The world's worst weeds - distribu-
dicotyledons have a branched growth tion and biology (1977) by L.G. Holm
form. Not all broadleaf weeds how- and colleagues. Other references
ever, are dicotyledonous. Commelina include Krantz et al (1977), Moody
benghalensis, Monochoria vaginalis, and (1981), and Moody et al (1984).
Eichhornia crassipes have broad leaves
but are monocotyledonous weeds. Approved computer
codes for weeds
Habitat Computer codes for weeds now
Weeds in rice can be grouped on the available (Bayer Agrochemicals
basis of their adaptation to submerged Division 1986) are useful for univer-
conditions as lowland or upland sal identification in bibliographies
weeds. Lowland weeds may be and data bases. An approved com-
semiaquatic or aquatic; upland weeds puter code is a five-letter abbrevia-
are adapted to dry sites. tion based on the scientific name of
Information on yield loss from each plant. In general, the first three
uncontrolled weeds from different letters refer to the genus and the last
experiments around the world is two denote the species, the subspe-
given when available to indicate the cies, or the variety. For example, the
weed's potential competitive ability. code for Echinochloa colona is ECHCO;
The losses will differ from one locality for Echinochloa crus-galli, ECHCG;
to another because they are affected and for Echinochloa crus-galli P.B. var.
by environmental factors such as nu- kasaharae Ohwi, ECHCK.
trient, moisture, temperature, time of
weed emergence, and density. The
yield loss information is not meant to
show the relative competitiveness
among the different weeds, and the
yield loss figures should not be used
to target particular weeds for control,
leaving others to compete with the
crop.
Seedling
Weeds worldwide 9
Lowland rice weeds / CYPERACEAE (sedge family)
Seedling Inflorescence
Mature plant
Seed, magnified
Inflorescence
Weeds worldwide 11
Lowland rice weeds / CYPERACEAE (sedge family)
Seed, magnified
Seedling
Synonyms: None
Local names:
County Common name
France scirpe maritime
Peru coco grande (Spanish)
Philippines apulid (Tagalog)
spreads by tubers.
Habitat: Grows in wet and flooded
soils.
Weedy nature: Tubers have
dormancy. Stems grow rapidly during
early rice growth and may severely
shade semidwarf rice cultivars during
the first 40 d. Difficult to control
because of apical bud dormancy and
capacity to produce numerous tubers.
Establishes, spreads, and becomes a
dominant weed within a short time.
Agricultural concern: Very competi-
tive in lowland rice. Rice yield losses of
60-80% can occur. Difficult to eradi-
cate because of dormant tubers and
buds.
Vegetative shoot
Weeds worldwide 13
Lowland rice weeds /MARSILEACEAE
Mature plant
Seedling
Seed, magnified
Seedling
Weeds worldwide 15
Lowland rice weeds /POACEAE (grass family)
Inflorescence
Seed, magnified
Weeds worldwide 17
Lowland rice weeds / POACEAE (grass family)
Seed, magnified
Seedling
Inflorescence
Weeds worldwide 19
Lowland rice weeds / POACEAE (grass family)
Mature plant
Seed, magnified
Seedling
Weeds worldwide 21
Lowland rice weeds / POACEAE (grass family)
Seed, magnified
Seedling
Weeds worldwide 23
Lowland rice weeds / SPHENOCLEACEAE (sphenoclea family)
Seedling
Inflorescence
Seedling Inflorescence
Weeds worldwide 25
Upland rice weed / AMARANTHACEAE (amaranth family)
Seedling Inflorescence
Weeds worldwide 27
Upland rice weeds / COMMELINACEAE (spiderwort family)
Seed, magnified
Mature plant
Weed name: Cyperus rotundus L.
Synonyms: None
Local names:
Country Common name
Argentina cebollita
Bangladesh motha (Bengali)
Brazil tiririca-comum
(Portuguese)
Cambodia smav kravanh chrouk
(Khmer)
France souchet rond
Ghana hyia-me-nonum (Fanti)
India nagar, motha (Orya)
korai (Tamil)
dila (Telegu)
Indonesia teki
Malaysia rumput haliyahitan Rhizomes and tubers
(Malay)
Mexico coquillo (Spanish)
Pakistan deela (Urdu)
Philippines mutha (Tagalog)
Senegal ndidan (Wolof)
Sri Lanka kalanthi (Sinhalese)
Thailand haew moo (Thai)
USA purple nutsedge
Life cycle: A perennial sedge that
can grow as tall as 75 cm. Propagates
mainly by tubers and produces few
seeds.
Habitat: Grows well in moist soils.
Weedy nature: Persistent and
difficult to control. Enormous amounts
of tubers may be produced. Tubers,
which have dormancy, can survive long
periods of environmental stress.
Difficult to control by cultivation
because cultivation breaks the
dormancy in the interconnected tuber
leading to more vigorous sprouting.
Agricultural concern: Limits rice
Inflorescence
production by competing for water and
nutrients. Can reduce rice yields 50%.
Seedling
Weeds worldwide 29
Upland rice weeds / EUPHORBIACEAE (spurge family)
Seedling Inflorescence
Weeds worldwide 31
Upland rice weeds / POACEAE (grass family)
Seedling Inflorescence
Seed, magnified
Weeds worldwide 33
Upland rice weeds / POACEAE (grass family)
Rhizomes
Seed, magnified
Weeds worldwide 35
Upland rice weeds / POACEAE (grass family)
Seed, magnified
Seedling
Weeds worldwide 37
Deepwater rice weeds
PONTEDERIACEAE (pickerel-weed family)
Flower
Weeds worldwide 39
Chapter 3
Weed control
Weeds have always reduced rice techniques. Each control method has Land prepration. Land preparation
yields. As a result, many different advantages and disadvantages, and a includes plowing, disking, harrow-
weed control methods have evolved. single method is rarely adequate for ing, soil puddling, and land leveling.
Farmers consider financial resources effective and economical control. A well-prepared field allows the rice
and availability of labor in deciding crop optimal early growth. Careful
what weed control method to use. Cultural methods land preparation primarily provides
Problems of input availability, avail- A basic principle of cultural control is weed-free conditions at planting.
ability of new technologies, specific to increase the competitive ability of In general, tillage practices most
weed problems, farm size, and rice and enable it to suppress weed affect plant growth during germina-
availability of family labor are basic growth. A vigorous rice crop com- tion, seedling emergence, and stand
management factors they take into petes more effectively with weeds establishment stages. Plowing buries
account in making weed control than does a less vigorous crop. weed seeds to depths from which
decisions. Cultural control methods include they cannot emerge, but it also brings
prevention of weed introduction, some weed seeds to the soil surface
Planning effective land preparation, crop rotation, where conditions favor germination.
cultivar selection, time of seeding, Thus, a new flush of weed seedlings
control planting method, plant population, occurs after each cultivation.
Planning is important in making fertilization, and water management. To destroy as many weeds as
appropriate decisions on weed Prevention of weed introduction. possible, the interval between succes-
control. Unfortunately, weed control Although weed seeds may be intro- sive cultivations should be long
often is not planned. The decision to duced in ways beyond a farmer's enough to allow many weed seeds to
control is not made until the problem control, many aspects of weed disper- germinate and be killed by later
has become serious, when control sal are controllable. Control involves harrowings. This can reduce a weed
may be uneconomical, ineffective, or preventing the introduction, estab- seed population about 50%. Tillage
even impossible. lishment, and spread of weeds, seeds, during the dry season is a practical
Advance knowledge of weed tubers, and rhizomes in a crop or method of controlling perennial
problems can be obtained by survey- between two crops. In rice, this is best grasses such as Paspalum distichum,
ing and recording the weed species in achieved by planting rice seeds free Cynodon dactylon, Oryza longistami-
a ricefield after rice emergence, at of weed seeds in weed-free seedbeds nata, and Imperata cylindrica; it
midseason, and at harvest. This and by seeding rice or transplanting desiccates the perennial structures. In
record is useful in planning weed seedlings in weed-free fields. Field temperate areas, tubers and rhizomes
control and crop rotation programs. borders not cropped or not kept clean brought to the soil surface are killed
are a constant source of weed seeds. during cold, dry periods.
Levees and irrigation canals also The type of land preparation
Control methods must be kept weed free. Weed seeds needed for rice depends on the water
Weed control methods can be can be introduced into a clean area by management system. Land prepara-
grouped into cultural, manual, machinery or tillage equipment that tion can be classified broadly as
mechanical, chemical, and biological are carrying weed seed-contaminated wetland tillage, dryland tillage, and
soil. limited tillage.
Weed control 41
Wetland tillage is common in Under limited tillage, perennial capacity of modern high-yielding
most tropical Asian countries. Tradi- weed problems increase over time. cultivars. Traditional varieties also
tionally, it involves plowing and pud- Where perennial weeds are con- have other attributes, such as suscep-
dling the soil. The processes involved trolled, limited tillage can be used for tibility to lodging, that make them
are both upland and lowland rice culture. undesirable.
flooding the field for about 7 d. Herbicide use is an integral part of Modern rice cultivars, although
plowing the soil to 10- to 20-cm a limited tillage system. However, if competitive against weeds, can have
depth, to bury the weed seeds and inappropriate herbicides are used, more weed problems than traditional
weed and rice stubble. perennial broadleaf weeds and rice cultivars. The erect leaves and
harrowing to break up big clods in grasses common in fallow in the short stature of modern rices allow
the soil. Two to three harrowings tropics are not controlled and signifi- light to reach the soil and stimulate
are done at 7- to 10-d intervals. cant yield losses can occur. Plowing weed seed germination. Within the
leveling the field. and harrowing more often does not crop canopy, small differences in
Puddling hastens crop establish- eliminate the need for direct weed crop height affect the quantity and
ment and tillering of transplanted control. Farmers still must follow quality of light received by weeds.
rice seedlings. This results in vigor- land preparation with other weed The high nitrogen rates used on
ous rice growth and increases the control methods. It is more cost- modern cultivars also aggravate
crop’s competitive ability against effective to reduce preplanting weed problems. This makes competi-
weeds. During transplanting, weed harrowings and combine those with tiveness with weeds a breeding
seedlings also are trampled and direct weed control methods. objective for modern rices. When
incorporated into the soil. Crop rotation. Weeds adapt to the available, such cultivars should be
Dryland tillage is the basis of growing conditions of rice. A ricefield used.
many rice culture systems. Almost all that is tilled regularly has more Time of seeding. When soil moisture
rice-growing areas in the United annual weeds than many undis- is not limiting, allowing weeds to
States and southern Australia, most turbed habitats, which have more germinate before tilling and seeding
of Latin America and West Africa, perennial weeds. Continuous crop- rice reduces weed populations. Rice
and parts of tropical Asia and Europe ping encourages the buildup of seeding, however, should not be
use dryland tillage. In dryland prepa- difficult weeds. delayed beyond the optimum time of
ration, soil clods are broken up so Each rice culture is associated with planting.
they do not interfere with seeding a characteristic weed problem, In rainfed rice crops, time of
and seedling emergence. The seedbed influenced by the cultural practices seeding is critical. Rice plants affected
is left rough because germination of used. Lowland rice has predomi- by drought become stunted and
weed seeds, which are usually much nantly water-tolerant weeds, upland cannot compete effectively with
smaller than rice, is encouraged by rice has mostly dryland weeds. weeds that are more tolerant of
fine tilth. Rice is seeded immediately Rotation of lowland rice with an drought. Dry weather after sowing
following the last tillage operation, to upland crop reduces infestation by can reduce the number of weeds that
give rice an even start with weeds. water-tolerant weeds in the rice and germinate near the soil surface, but
Limited tiIlage covers a range of by upland weeds in the upland crop. rice seed germination will also be
land preparation techniques, from Growing a broadleaf crop in rotation delayed. Perennial and large weed
zero tillage to elimination of one with rice allows the use of herbicides seeds from deeper soil depths are still
preseeding cultivation. Herbicides effective against grassy weeds, which able to germinate and compete
replace the omitted tillage operations. are difficult to control in rice. strongly with the rice crop.
Limited tillage has soil conservation Varietal selection. Improved rice Planting method. Planting method
advantages. cultivars resistant to diseases and affects the ability of rice to compete
Smallholders who use slash-and- insects are more competitive against with weeds. Rice may be trans-
burn land preparation are practicing weeds than are traditional rices. planted or drill seeded in straight
limited tillage. Rice is planted in a Traditional, tall varieties with droopy rows, or broadcast seeded. Straight-
dead mulch with only enough soil leaves are thought to be more com- row planting is necessary for hand
disturbance to cover the seed. petitive against weeds, but research weeding and for using mechanical
Minimum and zero tillage techniques has failed to measure competitive weeders. In a broadcast crop,
can result in timely land preparation differences that are due to cultivar. mechanical and hand weeding
and savings in labor, water, power, The height advantage of traditional damage the rice plants.
and capital. varieties is offset by the high-tillering
Weed control 43
leaf size), with the resultant shade Weed problems intensify in Burning. Burning, common under
helping to suppress late-germinating irrigated and rainfed lowland rice the slash-and-burn system of land
weeds. Phosphorus encourages rice when water supply is inadequate. preparation, kills weed seeds and
root development and increases Increased weed control benefits are weed seedlings, gets rid of unwanted
tillering. Vigorous rice root growth is obtained with improved water vegetation, and reduces the amount
advantageous in below-ground supply and control, high levels of of weed seeds returned to the soil. It
competition with weeds for moisture fertilizer, and improved cultivars also encourages germination of weed
and nutrients. used in concert. seeds near the soil surface. Thorough
Weeds take up nutrients in large Increasing water depth to control burning can keep the ricefield free of
quantities, however, and sometimes weeds as part of integrated weed weeds for the first 2-3 wk.
absorb fertilizer faster than rice. management is cost effective. When Burning saves labor, is low cost,
When weeds are not controlled combined with other direct weed and adds neutralizing ash to low-pH
effectively, however, fertilization is of control methods, increased water soils. On the other hand, widespread
little significance. There is little or no depth can give considerable savings uncontrolled burning leaves the soil
response to N by rice in shade, and P in production costs. bare, increasing soil erosion and loss
and N left on the soil surface will Timely and thorough drainage of of N and other nutrients. Careful
stimulate growth of shallow weed flooded fields reduces aquatic weed planning and rational use can
seeds. At high fertilization levels, it is problems. minimize the adverse effects of
not possible to produce high rice burning. Local regulations on
yields without weeding. Manual methods burning should be followed.
Early- to midseason N applications Manual weed control includes Hand pulling. Hand pulling
are beneficial to rice, but weeds must burning, hand pulling, and mechani- controls weed seedlings growing
be controlled to maximize the effect cal hand weeding. These labor- near and between rice plants where
of N on grain yield. Figure 3.1 shows intensive methods are the oldest and, implements are difficult to use. Hand
the best time to apply N in various in many cases, the farmer’s only pulling is not effective in dry soil,
rice cultures. means of controlling weeds in rice, where weed seedlings break and
Water management. Since ancient and are highly effective. resprout easily. Frequent hand
times, water has been used to manage Several hand tools are still the pulling is necessary for effective
weeds in ricefields. Many nonaquatic principal means of rice weed control weed control because very small
weeds do not survive in submerged in many developing countries. But weed seedlings that are not removed
environments and many aquatic manual methods are slow, grow quickly to reinfest the ricefield.
weeds do not survive in upland unattractive, and tedious, and are This method is the most labor inten-
environments. often carried out after the rice crop sive of all weed control measures,
Water control during the early rice has been severely damaged by and is best suited to small farms.
growth stages has a major effect on weeds. Hand weeding is often In some places, lowland rice
weed control. As weeds become ineffective on weeds with special farmers trample weeds instead of
established, controlling them through survival mechanisms such as the rice hand pulling.
water management is more difficult. mimics (e.g., I. rugosum) and deep- Mechanical weeding. Weeding using
Grassy weeds can be largely elimi- rooted perennial weeds. Rice mimics hand tools is common in almost all
nated by continuous flooding to are difficult to distinguish from rice tropical rice-growing areas. But the
15-cm depth maintained throughout at the early growth stage. Repeated degree of weeding and the problems
crop growth. The response of broad- hand weedings are necessary to associated with it largely depend on
leaf weeds and sedges to different effectively control all weeds. the type of rice culture. In many
water depths varies. After continuous countries, hand tools such as the hoe,
flooding, C3 weeds (see page 3) will narrow spade, Swiss hoe, knife,
be dominant. machete, and pointed sticks are
primarily used to remove weeds in
upland rice. Weeds within rows must
be removed by hand. The amount of
labor required to weed 1 ha ranges
from 10 to 30 d.
Weed control 45
Economics of control Table 3.1. Economic acceptability a of direct weed control methods in irrigated transplanted rice
in the Philippines (data for analysis obtained from Moody et al 1983).
The economic benefit of weed control
Grain Total Total Return above Marginal
must exceed the cost. The primary Control method yield variable return variable cost benefit-
aim of a rational farmer is to optimize (t/ha) cost (US$) (US$) (US$) cost ratio
profits. One way to achieve that is to
No weeding 1.5 0 263 263
reduce weed control costs. It is One hand weeding 3.6 50 630 580 6.3
logical, therefore, that where one or a Two hand weedings 3.7 90 648 558 3.3
combination of methods exists, and Two rotary weedings 2.9 44 508 463 4.5
2,4-D (0.8 kg/ha) 3.1 10.3 543 532 26
both are equally effective, the farmer Thiobencarb/2,4-D 3.3 19.0 578 559 16.0
will choose the least costly. (1.0 + 0.5 kg/ha)
Weed control costs include direct a Assumptions: Labor = US$2.09/d; First hand weeding = 24 d/ha: Second hand weeding = 19 d/ha: First rotary
costs (labor, herbicides, sprayers, etc.) weeding = 11 d/ha; Second rotary weeding = 10 d/ha: Herbicide = 2,4-D (0.8 kg/ha) = US$10.30. Thiobencarb/
and hidden costs. Management time 2,4-D = US$19.
Weed control 47
Chapter 4
Herbicide use 49
achieved by applying herbicides to the crus-galli. This is due to differences in The xylem is the plant system
soil surface, by band application, or by plant enzyme levels. Rice plants have a through which water and dissolved
using shields. Selectivity of soil- high level of aryl acylamidase, an mineral nutrients pass to the leaves.
applied herbicides may be lost if the enzyme that hydrolyzes propanil to Herbicides taken up by the root move
herbicides leach through the soil and nonphytotoxic 3,4-dichloroaniline and along this stream. Herbicide uptake is
come into contact with rice roots and propionic acid. E. crus-galli has a low therefore affected by factors affecting
shoots. level of this enzyme and is unable to transpiration rate, such as light, tem-
Plant growth stage. In general, plants hydrolyze propanil; therefore it is perature, wind speed, humidity, and
are most susceptible to herbicides at easily killed. This detoxification soil moisture. The triazine herbicides
the seedling stage. As plants grow, process can be inhibited by organo- are good examples of herbicides trans-
they become less susceptible. In direct phosphorus and carbamate insecti- ported up in the system.
seeded rice, there is no difference in cides. For a more detailed discussion of the
growth stage between rice and weeds, principles of herbicide action, refer to
and selectivity cannot be achieved
through plant growth stage. In trans-
Herbicide movement Weed science principles by W.P. Ander-
son (1983).
planted rice, however, differences in in plants
growth stage and position of growing
points of rice and weeds can be used to
Herbicides must enter the plant before Timing of herbicide
their toxic effect can be induced.
achieve selectivity. Plants growing Herbicides applied to leaf surfaces and application
under drought conditions are less buds penetrate the plant by diffusion. Weeds should be removed from rice as
affected by herbicides than plants Higher temperatures increase the rate early as possible. Thus, herbicides
growing with normal soil moisture. of penetration. Herbicide absorption should be applied during early crop
takes place in the guard cells of the growth stages. The time to apply a
Biological factors of selectivity stomata and through the cuticle. herbicide depends on the properties of
Biological factors of herbicide selectiv- In the soil, herbicides move in the the herbicide and the target weeds,
ity include differences in morphology, soil solution to the seed and roots, or weather, and cultural practices. Herbi-
physiology, and metabolism among are intercepted by the root tips. Herbi- cides can be applied at several periods
plant species. Leaf surfaces that are cides may penetrate the walls of root before and during the crop growing
waxy, smooth, or densely hairy are epidermal cells by mass flow. period. In general, herbicides are
wetted less readily by aqueous sprays Once in plants, herbicides move via applied at preplanting, preemergence,
than are surfaces that are less waxy or the phloem and the xylem systems to or postemergence.
moderately hairy. Vertical leaves retain cells and tissues remote from the site of
less spray than do horizontal leaves. uptake. The phloem conveys sugar Preplanting herbicide application
Once absorbed by a plant cell, the from the green tissues of the plant A preplanting herbicide application is
herbicide may be immobilized within (where sugar is manufactured) to made before the rice crop is sown. This
the cell; that also contributes to herbi- storage tissues. Very young leaves do application helps in land preparation
cide selectivity. Selectivity among not export sugar, so herbicides applied in a minimum tillage cropping system.
plant species may be achieved when to them remain there. If the transport of Translocated foliar herbicides (such as
some plant species are able to detoxify sugar is restricted, as when plants are glyphosate) kill perennial broadleaf
a particular herbicide, while others are under low light intensity, redistribu- weeds and grasses found in the fallow
unable to do so and are killed. For tion of herbicides will not occur. As a vegetation. Where annual weeds
example, rice plants are 40 times more result, the general recommendation for predominate, paraquat is adequate.
tolerant of propanil than Echinochloa many translocated herbicides is that Where volatile preplanting herbicides
the weeds should be in active growth. are used, they must be incorporated
Glyphosate and MCPA are examples into the soil before planting, to avoid
of herbicides translocated in the damage to the rice crop.
phloem.
Herbicide use 51
Runoff Photodegradation propanil is applied to rice when the
Runoff is one of the main pathways of Photochemical and biochemical temperature is above 38 °C, phytotoxic-
herbicide loss from flooded ricefields. degradation of herbicides govern their ity may occur. At high temperatures,
Herbicides are usually transported as fate in a flooded environment. For simetryn will cause injury, even in
solutes in soil water; their movement example, the high pH of the water japonica rice, as a result of higher
will depend on solubility and adsorp- induces hydrolysis of carboxylic esters. absorption through the roots. In
tive capacity. In runoff water, however, Some herbicides adsorb ultraviolet general, the rate at which herbicide
herbicides are transported both as radiation. The presence of humic acids degrades increases with increasing
solutes and on soil sediment particles. in floodwater may also induce photo- temperature.
An irrigation system needs to be well- chemical degradation of herbicides
regulated to retain herbicides within that do not adsorb ultraviolet radiation Relative humidity
the system. Irrigation drainage and (Yaron et al 1985). At high relative humidity, leaf stomata
overflow should be avoided, to reduce are open, which increases absorption of
herbicide losses and to protect down- Persistence herbicide into the leaf. Evaporation of
stream water from pollution. Herbicide persistence is the length of herbicides from leaf surfaces is slowed
time a herbicide remains active in the at high relative humidity. Slow evapo-
Volatilization soil. Persistence depends on the ration lengthens the time the herbicide
All herbicides are volatile (have a amount of herbicide applied, the rate at can enter the plant.
tendency to change from a solid or which it is broken down, properties of
liquid to a gaseous state). Volatility, the particular herbicide, and leaching. Soil moisture
however, varies among herbicides and Herbicides with long persistence keep Soil moisture affects herbicide effec-
increases with higher temperature. a crop weed-free for a longer time than tiveness by influencing the amount of
Volatile herbicides should be mechani- do herbicides with short persistence. herbicide in the soil solution and the
cally incorporated into the soil to avoid Persistence of a herbicide beyond the depth of herbicide movement in the
excessive chemical loss. rice-growing season, however, is soil profile. When a residual herbicide
Volatilization from the floodwater undesirable because other crops is applied to a dry soil, it relies on soil
into the atmosphere is an important sensitive to that herbicide cannot be moisture (from rain or from irrigation)
route of herbicide loss from lowland grown on the same land for some time. to move it to the root zone. Inadequate
ricefields. The volatilization rate movement is a common cause of the
depends on water evaporation rate, Effect of environment on failure of herbicides in upland fields.
water depth, water solubility, and
vapor pressure of the herbicide. herbicidal activity Wind
Volatilization loss of herbicides from a Several environmental factors affect Wind adversely affects the absorption
shallow, warm water flooded field can the success of weed control by soil or of foliar-applied herbicides by increas-
be highly significant. Volatilization foliar-applied herbicides. Temperature, ing the evaporation of spray droplets
from the soil surface of upland rice relative humidity, soil moisture, and and the volatilization of herbicide
may be much greater than that from wind are important. residue from the leaf surfaces.
the floodwater of lowland rice.
Thiocarbamate herbicides are volatile Temperature
in floodwater. Absorption and translocation of herbi- Properties of herbicides
cides increase as temperature When the probable behavior of a
increases, and selectivity can be herbicide can be predicted from its
changed by differences in absorption properties, that information can be
and translocation. For example, if used to design safer and more effective
application. Some properties of herbi-
cides that affect their biological activity
Herbicide use 53
Surfactants are often added to
herbicide formulations by the manu-
facturer, but additional surfactants
may be needed for some postemer-
gence herbicides or in some environ-
mental conditions (e.g., bentazon
under low temperatures). The herbi-
cide label indicates how much
surfactant to use when an additional
surfactant is required. When surfac-
tants are not recommended, their use
with foliage-applied herbicides can
result in loss of selectivity.
Herbicide labels
The best source of information con-
cerning safe and effective herbicide use
is the product label (Fig. 4.1) printed
on or attached to the herbicide con-
tainer. It gives directions on effective,
safe use of the herbicide. The label
should be read carefully and under-
stood before any herbicide is used. The
herbicide label is a legal document that
in many countries has government
approval. Every herbicide product
label should contain the following
information:
Trade or brand name. The trade or
brand name identifies a herbicide of
one company and differentiates it from
those of other companies. Like com-
mon weed names, trade names of
herbicides may change from country to
country. Thus, the trade name is not
always adequate for identifying a
herbicide or determining correct rates
of application. When an application
rate is recommended with a trade
name, the rate normally refers to the
amount of formulated product, and tional Union of Pure and Applied 4.1 All herbicide labels are required to
provide important information. Users should
not the rate of active ingredient or the Chemistry [IUPAC]). The chemical read and understand the label before opening
acid equivalent. name refers to the active ingredient. In any herbicide container.
Chemical and common names. Herbi- addition to the chemical name, each
cides have complex chemical names. herbicide is given a common name
Rules for selecting suitable chemical which also applies to the active ingre-
names are given by appropriate dient. The common name is not the
chemical associations (e.g., the Interna- same as the trade name of the same
herbicide. Common names for herbi-
cides are approved by appropriate pro-
fessional bodies, such as the Interna-
tional Standardization Organization
Herbicide use 55
Controlled-droplet applicator 4.2 Top, a leverage-operated
A controlled-droplet applicator (CDA) knapsack sprayer; bottom, a
controlled-droplet applicator.
produces uniform-sized droplets of
spray. Weeds may be effectively
controlled with as little as 18 liters
water/ha. Droplet size is controlled
within narrow limits by spinning discs
inside the applicator. Controlled-
droplet applicators are lightweight.
They have a plastic spray head, a small
motor that drives the rotating disc, a
liquid reservoir, a handle, and a power
supply (e.g., one to eight 1.5-volt dry-
cell batteries). Models that use a
manually operated air pump (Fig. 4.3)
(e.g., Birky sprayer) have been intro-
duced. This eliminates the cost of
batteries for the first-generation CDA.
Granular applicator
Herbicides sold as granules are applied
using granular applicators. Whatever
its size, a granular applicator consists
of a hopper to contain the granules, a
metering device to control flow rate,
and a distribution mechanism. Granu-
lar herbicides that are soluble in water
may not require the same uniformity of
distribution as those that are not
soluble in water.
Granules are often applied by hand,
especially in the tropics. Because
broadcasting granules by hand is not
precise, the quantity of herbicide used
will be higher than with other
application techniques. More uniform
application can be achieved on small
areas by shaking the granules from a
corrosion-resistant material, such as Operation of the
stainless steel, polyvinyl material, or
container with a perforated lid. fiber glass. The tank has a special lid knapsack sprayer
through which it is filled. The lid Herbicide and water are poured into
Hydraulic applicator (pressurized) contains a strainer to remove debris the sprayer tank. The lever is moved
Hydraulic applicators commonly that might clog the nozzle. The pump up and down, causing the pump to
consist of a tank, pump, and nozzle. drives the solution through the pres- draw spray liquid from the tank into a
Other components include an agitator, sure chamber to the nozzle. The most special chamber inside the tank called
pressure gauge, pressure regulator, widely used small hydraulic applicator the pressure chamber. Air trapped in
hose, and boom. The tank that holds is the lever-operated knapsack sprayer
the spray solution is made of a (Fig. 4.4). For a detailed discussion, see
Pesticide application methods (Matthews
1979).
the pressure chamber is compressed as The faster the lever is moved, the Selecting the nozzle
the liquid is forced in. On one side of higher the pressure becomes. High The flow rate through a nozzle de-
the pressure chamber is the hose, pressure makes the herbicide droplets pends on the size of the nozzle opening
which is connected to the trigger smaller and increases the speed with and the spray pressure. Increasing the
handle, and the on-and-off switch which the solution comes out of the nozzle size (diameter of opening) and
known as the cutoff valve. Compressed nozzle (flow rate). For herbicide increasing the pressure will increase
air forces the liquid from the pressure spraying, where coarse spray droplets the flow rate. A pressure of 70-275 kPa
chamber through the hose to the are desired, 5-6 lever strokes per (see Appendix A) is recommended for
nozzle. The nozzle turns the liquid into minute are adequate. herbicide application. The best way to
droplets, which aid in uniform cover- regulate the flow rate is to change
age of the weed or soil. nozzle tips and to increase or decrease
walking speed.
Herbicide use 57
It is important to select the proper There are several ways to find out
type of nozzle for each activity how much spray is applied to a given
(Fig. 4.5). For residual herbicides area for a given time. A well-tested
applied to the soil, impact nozzles (e.g., technique for calibrating a sprayer
Polijet) should be used because they (Fraser and Burrill 1979) is as follows:
cause less drift problems. For systemic, Step 1. Make sure the sprayer is in
translocated herbicides, where thor- good working condition (no leaks, no
ough wetting of stems and leaves is blocked nozzles, etc.). Calibration
not required, an impact nozzle or fan should be done on a surface similar to
nozzle is recommended. For contact the field to be sprayed. Measure and
herbicides, use a hollow cone nozzle, mark out an area of 100 m2 (10 m x
an impact nozzle, or a fan nozzle 10 m). Trace with a stick the rows on
operated at 275 kPa. which rice normally would be sown in
the field. The row width should be the
Sprayer calibration one you will use in planting rice.
Sprayer calibration determines the Step 2. Place the sprayer on level
volume of water that will be applied on ground and put in 10 liters of clean
a given area by a given applicator water. Mark the outline of the sprayer
under given conditions. The volume of on the ground so the same spot can be
water applied by a sprayer depends on found later. Put the sprayer on your
walking speed, sprayer pressure, and back. Position the nozzle above the first
nozzle size. seed row mark. Pump the sprayer to
Walking speed. An increase in walk- develop pressure. Begin spraying the
ing speed results in less spray mixture plot you have marked, adjusting the
applied to a given area. Conversely, a height of the nozzle to cover whatever
decrease in walking speed results in swath width you desire. Maintain a
application of a greater volume applied constant nozzle height. Walk at a
per unit area. comfortable pace, which you must
Sprayer pressure. Increasing sprayer maintain throughout the calibration,
pressure results in a greater volume of and later in actually spraying the field.
spray mixture applied to a given area. Spray the 100-m 2 plot once. When you
Conversely, a lower spray pressure have completed spraying the plot,
results in less spray mixture applied. place the sprayer back on the ground in
The sprayer should be operated to give its outlined position and measure the
as steady a pressure as possible. A water level.
pressure gauge may be fitted to the Step 3. Determine the application
sprayer. rate by subtracting the volume of water
Nozzle size. The use of a large nozzle remaining in the sprayer from the
opening increases the volume of spray amount you started with. For example,
mixture applied to a given area. if the amount of water in the tank
Smaller openings deliver a smaller before spraying was 10 liters and the
spray volume.
Herbicide use 59
area less than 1 ha or for a tankful. The To summarize, the steps required to The product labels of most commer-
amount of water needed to spray a arrive at the amount of herbicide and cial herbicide formulations containing
given area can be calculated as water to be sprayed on a given area are salts and esters specify the amount of
1. Determine sprayer output per acid equivalent present in the formula-
hectare (calibrate the sprayer). tion. The acid equivalent is equal to the
2. Determine the quantity of formu- difference in weight, expressed as per-
lated product required per hectare cent, between the parent acid molecule
(from label or other recommendations). (minus a value of 1, representing the
3. Use the size of the area to be loss of the H+) and that of the salt or
If the area to be sprayed is 1,000 m2 and sprayed or the sprayer capacity ester molecule. It is always less than
the sprayer output is 200 liters/ha (whichever is smaller) to determine the l00%, and is calculated by the
(from the calibration described earlier), amount of water needed. following formula:
then the amount of water required to 4. Calculate the formulated product Molecular weight (mol/wt) of
spray this area is : needed for the quantity of water. acid
Appendix B gives calculated dosage
rates per hectare for different formula-
tions, so that you can cross check your
When the area to be sprayed is de- own calculations.
termined on the basis of sprayer
capacity, area can be calculated as Calculation of dosage in acid
equivalents of salts and esters Example: The molecular weight of
Herbicide dosage recommendations 2,4-D is 221. The molecular weight of
computed in kilograms of active its isopropyl ester is 263. The acid
ingredient per hectare refer to the equivalent of the isopropyl ester of
unaltered chemical molecule. But in 2,4-D is determined as
If a knapsack sprayer has a capacity herbicide molecules that are acids, the
of 15 liters and the sprayer output is acidic portion is normally transformed
200 liters/ha, then the area covered by to a salt or ester, to improve such
one full tank (15 liters) is characteristics as solubility in water or
oil and foliar penetration. In general,
the parent acid portion of the herbicide Field techniques for
molecule remains as the herbicidally
The total amount of formulated active portion, while the ester or salt using herbicides
product to be used in the examples attached to the parent acid satisfies the Proper field techniques are important
above can be calculated as functions for increased solubility or to get good weed control from herbi-
increased penetration. The acid equiva- cides. Read all labels before using a
lent of a salt or ester form of a herbi- herbicide, and follow the directions. To
cide, therefore, is that portion of the kill the weeds and not the rice, using
molecule representing the original acid the correct dosage is essential. Contact
form of the molecule. The dosage rec- herbicides work best when they are
ommendation for such herbicides is applied in a high volume of water.
given as kilograms of acid equivalent
of the active ingredient per hectare.
The recommended dosage of herbi-
cides applied as salts or esters is often
based on the herbicidal portion of the
salt or ester molecule, and excludes the
herbicidally inactive portion
(Anderson 1983).
Herbicide use 61
Granular herbicide application To calculate the application rate, use Indiscriminate herbicide use in
Most farmers apply granular herbicides the following formula: irrigated rice can adversely affect wild-
by broadcasting. Whatever the method life, humans, and the environment.
Granules applied
used, safety and uniformity of applica- in g per 100 m Improper herbicide use will contami-
Application rate
tion are important. Unlike liquid formu- in kg/ha = Swath width × 10 nate local water bodies. Inappropriate
lations, where the amount of water used in cm herbicide use or use of herbicides
is not critical as long as the sprayer has highly toxic to fish will kill or contami-
been calibrated to deliver a certain rate, Example: If the weight of granules nate the fish in that water, which will
granular formulations are not further used over a 100-m length with a swath affect the fish consumers.
diluted with a carrier. Thus the formu- diameter of 40 cm is 120 g, then the
lated product is constant and the application rate is Safe handling of herbicides
amount applied is determined by the 120 × 10 Handling an undiluted herbicide is
recommended application rate per 40 = 30 kg/ha more dangerous than handling the
hectare. It is important to follow this diluted product. Herbicides can enter
calibration method: Granular herbicides, although about the human body through the skin,
1. Determine the amount of formulated twice as costly as nongranular ones, mouth, nose, and eyes. Absorption
product per hectare using the formula can be applied once by hand, in asso- through skin is common and can occur
on page 59. ciation with fertilizer. They can also be from chemical spilling and splashing,
Example: To apply 2.0 kg ai/ha of applied using any one of a number of and from drifting of spray. Absorption
Machete with 5% ai, multiply 2 kg by mechanical spreaders. In small plots, through the nose and mouth can occur
100, then divide by 5 = 40 kg/ha. granules can be applied by shaking from inhaling spray droplets, vapors,
2. Measure and mark out a 5-m × 5-m them from a bottle with perforated lid or chemical dust. Safety tips for herbi-
area and determine the amount of (IRRI, unpublished circular). cide use are as follows:
granules required for the area. The Mix herbicides or other pesticides in
amount of butachlor required for the open air, never in enclosed
25 m2 is
Safe use of herbicides places with inadequate ventilation.
25 Improper herbicide use can harm Read the label on the herbicide
× 40 kg = 0.1 kg = 100 g crops, humans, wildlife, domestic
10,000 container and make sure any special
animals, and the environment. There instructions are understood.
3. Walk at a comfortable pace and are two types of herbicide toxicity: Leaks from a badly maintained
practice applying the herbicide several acute (a single oral dose) and chronic sprayer may increase contact
times, until you achieve a 100 g/25 m2 (a sublethal dose repeated over time). between the herbicide and the skin,
application rate. permitting the herbicide to enter the
When the granules are to be mixed in LD50 and LC50 body. Tighten leaking sprayer parts
a carrier such as sand, use a different LD50 is the expression for a single dose and check seals and washers to
calibration method, as follows: that, when taken orally, kills 50% of a avoid leakage.
1. Measure and mark out a length of 100 group of test animal. It is usually Wear protective clothing. Special
m. expressed in milligrams of herbicide protective clothing is heavy and
2. Weigh enough herbicide to cover this per kilogram of body weight of test expensive, and uncomfortable when
area. Walk at a comfortable pace and animal. The higher the herbicide toxi- used in the tropics. For small-scale
practice applying herbicide uniformly city, the lower the LD50. For example, farmers who use herbicides occa-
over this area. the LD50 of paraquat is 150 mg/kg; that sionally, a complete set of protective
3. Reweigh the herbicide remaining. of butachlor is 2,000 mg/kg. clothing may not be necessary. All
The difference will give the amount of We emphasize that the LD50 is NOT
herbicide applied. the safe dose level—50% of test ani-
4. Measure the swath width. mals die at that dose.
LC50 is the concentration required to
kill 50% of the test organisms in an
environment (usually water). The LC50
[96 h] for carp is 0.32 mg/liter.
Herbicide use 63
Chapter 5
Herbicides play an important role in Herbicides that may be combined effects on the environment. Herbicides
integrated weed management in rice. often are sold as formulated products. may be classified, for convenience, by
Early-season weed competition When these are not available, two or method and timing of application. Or
significantly reduces rice grain yield, more herbicides may be mixed in the they may be classified by chemical
and preemergence herbicide treat- spray tank at the time of application- group, which also gives an indication
ments are widely used. But most weed a tank-mix combination. Combina- about how the herbicide may be used.
seeds germinate over a long time, and tions must be selected carefully and
preemergence herbicides, with their comply with manufacturers’ recom- Anilides
relatively short residual life, may not mendations to avoid product Anilides are used to control germinat-
control weeds long enough to opti- incompatibility. ing annual weeds, especially grasses.
mize rice yields. Then, postemergence They often are most active as surface
herbicides may be needed along with Herbicide classification preemergence treatments. The primary
other control measures. Moreover, any mechanism of action is through inter-
one herbicide may not control all the and uses ference with nucleic acid and protein
weeds present in a ricefield. Herbicide The herbicides commonly used for synthesis. Butachlor, pretilachlor, and
mixtures are used to obtain a wider weed control in rice are described propanil are examples of this group.
range of weed control. here. Table 5.1 lists the weeds con- Butuchlor. Butachlor is absorbed
trolled by the herbicides. Details on primarily through germinating shoots
their use are discussed in Chapters 6-9. and secondarily through roots. Its
Herbicide mixtures, Every effort has been made to ensure mode of action is inhibition of protein
rotations, and that the information presented is synthesis. It is used at 2-3 kg ai/ha for
correct (Roberts 1982, Swarbrick 1984, preemergence control of most annual
sequences Attwood 1985, Chemical and Pharma- grasses at the 1- to 2-leaf stages and of
Mixing herbicides and spraying them ceutical Press 1986, Thomson 1986, certain broadleaf weeds, and can be
simultaneously increases the range of Worthing 1987). But because herbicide applied postemergence. For trans-
weed control. Using herbicide mix- activity varies from locality to locality, planted rice, it is applied 3-7 d after
tures can also save time and reduce only general recommendations are transplanting. For direct seeded rice, it
application costs. A broadleaf herbi- given. Specific recommendations is applied 10-12 d after emergence.
cide and a grass herbicide are often should be obtained from weed special-
mixed together (e.g., bensulfuron + ists in the reader’s locality.
butachlor). Residual and foliar contact Most herbicides are organic com-
herbicides may be combined (e.g., pounds. Herbicides are considered
thiobencarb + 2,4-D). ideal if they are toxicologically safe,
selective to rice, cost-effective, effective
on weeds, and have no lasting adverse
Principal herbicides 65
Table 5.1. Susceptibilitles a of important rice weeds to common rice herbicides (treatment at recommended doses and application times).
Herbicide
Weed
Acanthospennum hispidum S R S R R S S S R R MS
Aeschynomene viginica S S R S S MS S S R MS R
Ageratum conyzoides S S S R S S S S R R S
Altemanthera sessilis S MR S R S S S S R S
Amaranthus spinosus S S S S R S S S S S S S R S R S
Ammannia coccinea S S S R S S S R S MR
Bidens pilosa S S R S S MR S S R S
Brachiaria mutica R R R S R S R R R MR R R R
Commelina benghalensis S S R MR R MR S MR S MR R R MR R
Cynodon dactylon R R R R R R R MS R S R R R R R R R R R
Cyperus difformis S S S S S S S S R S S MS MS S S S S S S S R
Cyperus esculentus S MS MS S MS R S MR S S R R R MR R R
Cyperus iria S S S S S s S R S MS MS S S R S S S S S MR S R
Cyperus rotundus R MS R R R R MS R R S MS R R R R R R R R MR R R
Dactyloctenium aegyptium R R S S S R S S S S S R R
Digitaria sanguinalis R S R S S S R S S S S S MR S R
Echinochloa colona R MR S S S S R S MR S R S S S S S S S S S S R
Echinochloa crus-galli S R MR S S S S R S MR S R S S S S S S S S S S R
Echinochloa glabrescens S R MR S S S S R S MR S R S S S S S S S S S S R
Eclipta alba S MR S R MR S S MR S S R MR MR S
Eichhornia crassipes S R S MS R R
Eleusine indica R MS S S R S S S R S S S S S R MS S R
Eleocharis acicularis S S S R S S S S S S R
Euphorbia hirta S MS R S MS S S MR R
Fimbristylis miliacea S S S S S S R S S S MS S S S R S S MR S
lmperata cylindrica R R R R R R MR R S R R R R R R R R R R
Ipomoea aquatica R S R R MR MS R
Ischaemum rugosum R S S R S S R R S S S S MS R
Leersia hexandra R R R R S S R R R MR R
Leptochloa chinensis R MR S R S S S R S S S S S MR R S R
Ludwigia octovalvis S S MS S R S S S S S S R S S S
Marsilea minuta S S S MR R S MR S S R MS
Monochoria vaginalis S S S S S S S R S S S S S S S S S MS MS
Nymphaea stellata S R S S
Butachlor in soil is broken down by
microbial activity. In soil or water, it is
rapidly converted to water-soluble
derivatives. It may persist for 6-10 wk.
Formulations available include
granules and emulsifiable concentrate.
When applied postemergence, it can
be tank-mixed with propanil.
Water solubility is 20 mg/liter at
20 °C. LD50 for rats is 2,000 mg/kg.
Contact causes irritation to skin and
eyes. It has high fish toxicity-LC 50
(96 h) for carp is 0.32 mg/liter.
Pretilachlor. Pretilachlor is a selec-
tive chloroacetamide rice herbicide
that can be applied before transplant-
ing or any time between transplanting
and weed emergence. Pretilachlor at
0.6 kg ai/ha is well-tolerated by trans-
planted rice, but it cannot be used for
direct seeded rice without an antidote.
Rice can be protected against injury by
applying fenclorim one day before, at
the same time as, or up to 3 d after,
pretilachlor application (Christ 1985).
Fenclorim is also effective in protect-
ing water seeded rice, but is not
suitable for use on upland rice.
Water solubility is 50 mg/liter at
20 °C. LD50 is 6,099 mg/kg. It has high
fish toxicity-LC 50 (96 h) for rainbow
trout is 0.9 mg/liter.
Propanil. Propanil is a contact
herbicide that can be applied post-
emergence. It is effective against
several grassy and broadleaf weeds at
the 2- to 3-leaf stages, and has no
residual effects. Its effectiveness
decreases on grasses to negligible at
the tillering stage. Rice is extremely
tolerant of propanil because rice plants
have high levels of the hydrolyzing
enzyme aryl acylamidase, which
detoxifies propanil.
Principal herbicides 67
Propanil can be used at 3-4 kg ai/ha Paraquat. Paraquat is nonselective, propanil and the residual activity of
in irrigated and rainfed rice. It should with fast contact action when applied pendimethalin. Because soil and
be applied when most weeds have postemergence. It kills most annual weeds must be completely exposed to
emerged. The water level in a flooded weeds and grasses, including rice. It spray coverage, no floodwater should
ricefield should be lowered to expose can be used in zero tillage or mini- be on the field at the time of applica-
weeds about 24 h before propanil mum tillage systems and in stale tion. The residual activity of
application. Raise the water level again seedbed land preparation, at applica- pendimethalin is activated by mois-
1-3 d after treatment, before any new tion rates of 140 to 840 g ai/ha. It is ture. It is most effective when ade-
weeds emerge. A slight leaf burn may rapidly inactivated on contact with soil quate rainfall or irrigation is received
occur on rice after application, but the by strong adsorption to clay. within 7 d after application.
rice plant normally recovers quickly. Water solubility is 70 g/liter at Water solubility is 0.3 mg/liter at
Propanil should not be applied if 20 °C. LD50 for rats is 150 mg 20 ºC. LD50 for albino rats is 1,050-1,250
rain threatens to fall within 5-6 h after paraquat/kg. It has medium fish mg/kg. It has high fish toxicity-
application. It should not be applied to toxicity, depending on the formulation LC50 (96 h) for channel catfish is
rice at the late tillering stage, about used-LC 50 (96 h) for rainbow trout is 0.42 mg/liter.
60 d after planting. Propanil-treated 32 mg/liter.
rice crops should not be treated with Diphenyl ethers
organophosphorus or carbamate in- Dinitroanilines This group includes bifenox,
secticides within 14 d before or after Butralin and pendimethalin are fluorodifen, oxyfluorfen, and
treatment because those insecticides examples of dinitroanilines. Members chlomethoxynil. Diphenyl ether
inhibit the detoxification of propanil of this family are active when applied herbicides are classified as contact
by rice. Commercial mixtures include to the soil and must be applied before herbicides. When applied preemer-
propanil + molinate, propanil + weed seed germination. In general, gence, they inhibit seed germination
bentazon, and propanil + bifenox. dinitroaniline herbicides do not and early seedling growth. They are
Water solubility is 0.2 g/liter at control established weeds. Their mode relatively insoluble in water, and do
25 °C. LD50 is 1,400 mg/kg. It has high of action is inhibition of both root and not readily leach. They are used
fish toxicity-LC 50 (96 h) is shoot development (mitotic poison). principally preemergence or early pos-
13 mg/liter, indicating that streams Butralin. Butralin is a preemergence temergence to control broadleaf weeds
and lakes should be protected from herbicide. It is selective to upland rice and grassy weed seedlings. In general,
contamination. when applied pre-emergence and to these herbicides more effectively
pregerminated rice at the 1- to 4-leaf control broadleaf seedlings than
Bipyridyliums stages (4-6 d after seeding). It is active grassy seedlings.
Herbicides from this group are against many broadleaf weeds. Bifenox. Bifenox is primarily a
primarily postemergence, foliar-acting Water solubility is 1 mg/liter at broadleaf herbicide. It can be applied
compounds with no soil activity. 24 °C. LD50 for albino rats is 12,600 at 2 kg ai/ha as preemergence or
Paraquat and diquat are generally mg/kg. It has high fish toxicity-LC 50 postemergence up to the 2-leaf stage of
nonselective. On treated plants, (48 h) for rainbow trout is 3.4 mg/liter. rice. Bifenox has been found to be
toxicity symptoms are a characteristic Pendimethalin. Pendimethalin used highly toxic to direct seeded flooded
rapid scorch and desiccation. Light, as a preemergence herbicide inhibits rice, resulting in low grain yields (IRRI
oxygen, and chlorophyll are required germination and seedling develop- 1974). Mixtures include bifenox +
for maximum manifestation of ment of susceptible weeds. Where 2,4-D and bifenox + propanil.
phytotoxicity. grasses are expected to be a problem, it
should be applied after planting and
before emergence of rice and weeds. It
can also be applied as a postemer-
gence treatment with propanil, which
combines the direct contact action of
Principal herbicides 69
Thiocarbamates Thiobencarb. Thiobencarb is more Simetryn. Simetryn is used as a
Thiocarbamates are active when effective on grasses and sedges than mixture with thiobencarb to control
applied to the soil. Some are highly on broadleaf weeds. Thiobencarb broadleaf weeds in rice. LD50 for rats is
volatile, which makes soil incorpora- interferes with protein synthesis and 1,830 mg/kg.
tion necessary. Thiocarbamate herbi- inhibits photosynthesis. Protein Dimethametryn. Dimethametryn is a
cides control germinating annual synthesis and amylase biosynthesis are triazine compound used as a selective,
grassy and broadleaf weeds. The inhibited more in susceptible grass preemergence, and postemergence
process of lipid biosynthesis appears species than in rice. herbicide. It controls annual broadleaf
to be the most sensitive to thiocar- In transplanted rice, thiobencarb weeds and grasses.
bamates. Examples of this group are shows maximum herbicidal selectivity
thiobencarb and molinate. Most by taking advantage of differences in Sulfonylureas
thiocarbamate herbicides have a growth between rice seedlings at the Sulfonylureas have very high biolgical
relatively short soil persistence. Recent 3- to 6-leaf stages and germinating activity, and rates as low as
evidence (Roeth 1986) has shown weeds. Thiobencarb (2-3 kg ai/ha) is 0.002 kg ai/ha have been used.
increased breakdown of some thiocar- distributed in the top soil after applica- Although weed seed germination is
bamate herbicides in soils with a tion and is readily leached from the not usually affected, subsequent root
history of thiocarbamates application. soil. Rice plant injury symptoms and shoot growth are severely inhib-
Molinate. Molinate is a herbicide for include dwarf malformation and deep ited in sensitive seedlings. Weed
selective weed control in rice at greening. In some cases, leaf scorching growth inhibition is rapid, with visual
2-4 kg ai/ha. It is particularly effective appears. symptoms within 1-2 d in rapidly
for controlling Echinochloa weed Thiobencarb should be applied growing plants. The site of action of
species. It can be applied as a granular between preemergence and the 2-leaf the sulfonylureas is the enzyme
formulation to the water in flooded stage of Echinochloa spp. because acetolactate synthase. Inhibition of this
rice. Emulsifiable concentrates herbicidal activity decreases after this enzyme, which is needed for the
applied to the soil surface are stage. For post-emergence application, production of the essential amino acid
extremely volatile and must be thiobencarb should be applied after building blocks valine and isoleucine,
incorporated immediately. Molinate the 1.5-leaf stage of rice to the 2- to results in rapid cessation of growth
can be applied before sowing rice, at 3-leaf stage of Echinochloa spp. to and eventual plant death. All plants
postemergence preflood, at flooding, obtain the best weed kill without contain this target enzyme, but the
or postflood. Moisture is required to damaging the rice. Cool temperatures ability of some plants to rapidly
activate molinate. In pre-flood treat- may delay onset of herbicidal activity. convert the herbicide to an inactive
ment, the area should be flooded as Commercial mixtures available product is the basis for selectivity
soon as possible. Once applied, a include thiobencarb + propanil and (Beyer et al 1988).
continuous water cover must be main- thiobencarb + simetryn. Bensulfuron. Bensulfuron (0.05 kg
tained. Rice is extremely tolerant of Water solubility is 30 mg/liter at ai/ha) is a sulfonylurea herbicide used
this herbicide. Cool temperatures will 20 °C. LD50 for rats is 1,300 mg/kg. in direct seeded and transplanted rice.
cause less than optimal weed control. LC50 (48 h) is 3.6 mg/liter for carp. It has good crop safety on indica rice
Commercial mixtures include varieties, but less crop safety on
molinate + propanil. Triazines japonica types. Indica rices metabolize
Water solubility is 880 mg/liter at The triazines are a large group of the herbicide more rapidly than
20 ºC. LD50 for male rats is 369 mg/kg. herbicides. They are applied preemer- japonica rices. Thiocarbamates
LC50 (96 h) for goldfish is 30 mg/liter. gence and postemergence to control herbicides, except molinate, show an
At recommended rates, molinate had seedling grasses and broadleaf weeds. antidote effect on bensulfuron through
no detectable effects on fish in ditches They control broadleaf weeds better the acceleration of detoxification by
draining water from treated ricefields than grasses and do not control estab- rice.
in California. lished annual or perennial weeds.
Their mode of action is through
inhibition of photosynthesis in plants.
Simetryn and dimethametryn are
triazines.
Principal herbicides 71
Quinclorac. Quinclorac is a chino- Susceptibility of rice cultivars to
line-carboxylic acid compound used as grass herbicides has been observed
a selective preplant, preemergence, with propanil, butachlor, thiobencarb,
and postemergence herbicide. It has pendimethalin, molinate, and
been experimentally tested in rice at piperophos-dimethametryn. For
125-300 g ai/ha for grass weed control. example, IR5, IR28, and IR46 are
Organic matter in the soil will decrease susceptible to thiobencarb (Shin et a1
herbicidal activity. Early postemer- 1989). Indica varieties are usually more
gence application when grass weeds susceptible to simetryn than are japon-
are at the 1- to 3-leaf stage will give the ica varieties. On the other hand, ben-
best results. Although results are best sulfuron has good crop safety on
when quinclorac is applied onto indica rice varieties but less crop safety
saturated soil, it can also be applied on japonica types. Cultivar tolerance
onto dry soil or into standing water may be due to differences in growth
not deeper than 5 cm. Quinclorac can rate, growth stage, morphology, physi-
be safely used in dry seeded and water ology, and biochemistry.
seeded rice when application is made Herbicide selectivity is relative and
postemergence, from the 1- to 2-leaf can be overcome by increasing dosage
stage onward. It may be tank-mixed and by changes in environmental
with other rice herbicides. conditions. Herbicide label informa-
Oral LD50 , is 2,610 mg/kg. tion should be followed at all times to
prevent severe damage to the rice
crop.
Differences in herbicide
tolerance among rice
cultivars
Rice cultivars may vary in their
tolerance for or susceptibility to
herbicides. Broadleaf herbicides are
expected to have little effect on rice.
The differences in sensitivity to 2,4-D
and MCPA (phenoxy acetic acid
herbicides) that has been observed is
due to differences in the growth stage
of the cultivars at the time of herbicide
application.
About 77 million ha of rice (53% of the Weed problems Because nursery areas are small
world’s rice area) are partially or fully The weeds common in transplanted (about 21 × 21 m will provide enough
irrigated throughout the growing rice (Monochoria vaginalis, Echinochloa seedlings for 1 ha rice) and seedling
season (IRRI 1988b). In South and crus-galli, Cyperus difformis, Cyperus establishment takes only 20-25 d,
Southeast Asia, irrigated rice com- iria, and Scirpus maritimus) are in controlling weeds is easy. Propanil,
prises 33% of the rice-growing area; in general highly competitive. They have thiobencarb, butachlor, quinclorac,
temperate Asia, most ricelands are discontinuous germination and rapid bensulfuron, pretilachlor + fenclorim,
irrigated. In Europe, Australia, Egypt, growth and are adapted to aquatic and pendimethalin give good weed
Pakistan, and USA, ricelands are conditions. Weeds grow and infest an control in rice seedling nurseries.
entirely irrigated. irrigated field if optimum water depth Doubling the seed rate, hand
Irrigated rice is classified into four is not maintained. In poorly flooded weeding, or removing large weed
culture groups according to the crop ricefields, most semiaquatic lowland seedlings from rice seedling bundles
establishment technique used. rice weeds can germinate and survive. resulted in less than 50% control of
Transplanted in puddled soil. weeds. Careful examination of each
Direct seeded on puddled soil Stand establishment method plant to ensure that most weeds are
(broadcast or drill seeded using Twenty- to 30-d-old rice seedlings are removed from the seedling bundles is
pregerminated seed). normally transplanted into a puddled laborious, time-consuming, and more
Direct seeded on dry soil (broadcast soil. In the irrigated rice-growing areas expensive than herbicide treatment
or drill seeded using nongermi- of Asia, seedlings are raised in wet (Moody et al 1988).
nated seed. bed, dapog, or dry bed nurseries. In
Water seeded. the wet bed method, pregerminated Land preparation
seeds are broadcast uniformly on a Mechanical land preparation should
Transplanted in raised bed of puddled soil. Seedlings provide a weed-free field to allow
are ready for transplanting 20-25 d optimal early rice growth. The initial
puddled soil after sowing. In the dry bed method, plowing buries weeds and crop
Among irrigated rice cultures, trans- seedlings are grown similarly, but the stubble from the previous crop.
planted rice has the lowest potential soil is not puddled and drainage is Puddling uproots weeds that grow
loss to weeds, because of the head start provided. after plowing and buries them in the
rice seedlings have over weeds and Weeds in rice seedling nurseries can layers of mud. The field is leveled after
because of the weed control effects of cause the complete failure of the puddling to eliminate inadequately
floodwater. Despite these advantages, nursery. Nurseries used to raise rice flooded areas that are ideal for the
uncontrolled weeds can reduce rice seedlings should be kept weed free to growth and development of difficult-
yields by an average 48%, through prevent transplanting grassy look- to-kill semiaquatic weeds.
competition for light and nutrients. alike weeds along with the rice seed-
lings. Transplanted weeds are highly
competitive and extremely difficult to
control by hand weeding or by
selective herbicides.
Irrigated rice 73
Planting method
Transplanting into well-puddled soil
helps rice seedlings to establish
quickly. Healthy 20- to 30-d-old rice
seedlings transplanted in rows in a
well-prepared weed-free field will
have a head start over weeds. Small
rice seedlings are not competitive
against weeds.
Plant population
Most modern early-maturing rices,
which have a short vegetative period
that limits tillering, do best when
transplanted at close spacing. No
single spacing recommendation,
however, is best for all rice cultivars.
In the absence of lodging and weeds,
yields of most varieties do not change
much with planting distances between
25 and 10 cm in rows or hills. Dense 6.1 Single and 2-row cono weeders.
Drawings, design information, and limited
planting increases the competitiveness technical supported are provided free to 6.2 Rice growth stages
of rice against weeds by reducing later manufacturers who want to produce IRRI when herbicides can be
germinating weed seedlings through designs on a commercial basis. IRRI retains applied in transplanted
worldwide distribution and patent rights for irrigated rice. Bars (-)show
shading. Rice should be transplanted all designs developed by the Institute, and periods during which a
in straight rows, to allow mechanical does not grant exclusive manufacturing particular herbicide is
weeders to be used for weeding. rights or licenses in any country or region. applied. *Timing of herbicide
application is based on weed
emergence and growth stage
Water management within the rice growth stage.
Good water management will elimi-
nate all normal upland weeds in trans-
planted irrigated rice. The anaerobic
conditions prevailing in soil under
5 cm of water inhibit most weed
growth. Reductions in water level
expose the soil surface, which leads to
aerobic conditions that allow weed
seed germination. A field should be
flooded 2-3 d after transplanting and a
5-cm water depth should be main-
tained throughout the growing season.
Fertilizer
Rice response to fertilizer nitrogen is
markedly increased by good weed
control, with maximum yields when
weeds are controlled before fertilizer is
applied.
Hand weeding is the most common Herbicide Rate Comments and source of information
weed control method in irrigated (kg ai/ha)
transplanted rice. The first 6 wk after
Bensulfuron 0.05 Apply 3-5 d after transplanting (DT)
transplanting is the critical time of (IRRI 1986).
weed competition. Two or three timely Bentazon 1.0-2.0 Apply postemergence to control
weedings will provide adequate weed broadleaf weeds and sedges
(IRRI 1979). Drain before application,
control. if necessary to expose weeds.
Weeding by machine is possible Bifenox + 2,4-D 2.0 + 0.5 Apply preemergence to weeds 4 DT
when irrigated rice is transplanted in (IRRI 1981).
Butachlor 1.0-2.0 Apply 3-6 DT. Water depth of 5-10 cm
rows. Hand weeding requires about at application and for 3-5 d after
120 labor-hours per ha. That is (IRRI 1974).
reduced to 30-90 labor-hours when 2,4-D or MCPA 0.8-1.0 Apply 3-4 wk after weeds have emerged
to control sedges, broadleaf weeds,
mechanical weeders are used. The and aquatic weeds. Drain before
conventional single-row rotary application to expose weeds. Reflood
weeders require 80-90 labor-hours, but within 2-3 d after application. Can also
be applied at 4-5 DT, before weeds
they are difficult to push and must be emerge. Granular herbicides can be
moved back and forth for proper broadcast directly into floodwater
operation. (IRRI 1973, 1981).
Molinate 2.0-4.0 For preflood appllcation, flood as
The IRRI cono weeder (Fig. 6.1) soon as possible. For postflood
uproots and buries weeds with conical application, deepen water at
shaped rotors. Forward movement of application to cover weed foliage,
then lower water after 4-6 d. No need to
the weeder creates a horizontal back- incorporate granular formulation
and-forth soil movement in the top (COPR 1976).
3-cm layer, and the cono weeder Oxadiazon 0.5-0.75 Apply 2-8 DT to control annual grasses
and broadleaf weeds. Field should be
weeds satisfactorily in a single pass. flooded 3-5 cm and maintained at that
Power requirements are low because level for 2-5 d after treatment
only a small quantity of soil is moved. (COPR 1976).
Oxyfluorfen 0.15-0.25 Apply 4 DT (IRRI 1979, 1983).
IRRI’s two-row cono weeder can weed Pendimethalin 0.75 Apply 4 DT to control annual grasses
three to four times faster than conven- (IRRI 1979, 1983).
tional single-row rotary weeders. Piperophos + 0.5 Apply 2-5 DT to control annual
dimethametryn grasses, sedges, and broadleaf weeds
Weeds within or close to rice hills (IRRI 1977).
must be hand-pulled. Piperophos + 2,4-D 0.3 + 0.2 Apply 2-8 DT to control annual grasses,
sedges, and broadleaf weeds
(IRRI 1986).
Herbicides Propanil 3.0-4.0 Apply as postemergence spray to
In fields where heavy weed infesta- control several annual grasses and
tions are expected, weed competition broadleaf weeds at the 2- to 3-leaf
stages. Draln flooded fields 24 h
can be prevented by a wide range of before application and reflood 3-5 d
herbicides. Rice herbicides show after treatment. Do not spray organo-
maximum selectivity in transplanted phosphorus and carbamate
insecticides within 14 d after
rice because of differences in growth appllcation (COPR 1976).
between rice seedlings transplanted at Quinclorac 0.3 Apply 3-5 DT (IRRI 1986).
the 3- to 6-leaf stage and the germinat- Thiobencarb 1.5-4.0 Apply 4-8 DT at the 1- to 2-leaf stages of
weeds. Water should not be drained
ing weeds. Several herbicides and or overflowed for 3-5 d after
herbicide combinations can be used in appltcation (IRRI 1971).
transplanted rice. Applying a pre- Thiobencarb + 2,4-D 1.0 + 0.5 Apply 4-5 DT (IRRI 1971).
emergence herbicide together with
effective water management will
provide season-long weed control.
Some important rice herbicides and
their times of application in irrigated
rice are given in Table 6.1 and
Figure 6.2.
Irrigated rice 75
Direct seeded on Land preparation Water management
Thorough land preparation is essential Good water management is an impor-
puddled soil in direct seeded flooded rice. Land tant factor in weed control in direct
In direct seeded irrigated rice culture, preparation is similar to that for trans- seeded flooded rice. Seeds are broad-
the field is leveled after puddling and planted flooded rice. However, the cast onto puddled soil with little or no
pregerminated seeds are broadcast or final leveling of the field is even more standing water. The water level is
machine-drilled onto the puddled soil. critical than for transplanted rice increased gradually as the rice grows.
Direct seeding, also known as wet because the water level in direct Because the field cannot be flooded
seeding, is practiced in parts of India, seeded fields is kept shallow. An until seedlings are established, some
Bangladesh, and Sri Lanka and has uneven land surface results in areas weeds will grow along with the rice.
become an increasingly important rice where the soil surface is exposed to air. After rice establishment, the water
crop establishment method in South- That creates an ideal condition for level should be raised as rapidly as
east Asia. Broadcast seeded flooded weed germination and growth. Rice possible without damaging the young
rice is also practiced in several rainfed stands in areas that have deeper rice seedlings, then kept uniform and
areas in South and Southeast Asia flooding will be reduced. continuous. Weed emergence and the
(De Datta and Flinn 1986). type of weeds that emerge are closely
Direct seeding has become an Planting method related to floodwater depth. Shallow
acceptable alternative to transplanting Rice seeds are pregerminated (soaked (less than 2.5 cm), continuous flooding
as labor costs have increased, less in water for 24 h, then incubated for facilitates weed growth.
expensive herbicides have become 48 h) before they are sown in the field. For preemergence herbicide appli-
available, and irrigated area has This assures a quick and even stand. cation in direct seeded flooded rice,
increased. However, root anchorage is Pregerminated rice seeds may be the following water management is
poor, and lodging can be more serious broadcast or machine drilled. Mechan- suggested.
in direct seeded than in transplanted ical weeding is possible when seeds Keep the field saturated from
rice. are drilled in rows. sowing to herbicide application. If
the soil dries within this period, add
Weed problems Cultivar enough water to resaturate the
Uncontrolled weeds in direct seeded The cultivar used should have excel- plots.
flooded rice can reduce yields about lent seedling vigor and good tillering Flood the field to 2-3 cm deep and
53%. The culture requires shallow capacity. IRRI and national programs apply herbicide directly into the
flooding, which results in more have released several such rices. water.
exposed soil areas and aerobic condi- Raise water depth to 5 cm 1 wk
tions. Because rice and weeds germi- Plant population after herbicide application and
nate and emerge together, competition Close spacing is essential to reduce maintain that depth until 1 wk
is more intense than in transplanted weed infestation and for high grain before harvest.
rice. The range of herbicides that can yields. In wet seeded rice, less weed
be used safely is also limited, because competition has been observed with Fertilizer
rice and the weeds are at the same seeding rates of 100 kg/ha and higher. High fertilizer application to increase
development stages. Where weeds are not a problem, no yields of modern improved rice
Lowland weeds such as E. crus-gall, rice grain yield advantage has been cultivars enhances weed growth.
Ischaemum rugosum, Leptochloa observed at these seeding rates. Incorporating N into the seedbed at
chinensis, Cyperus difformis, Fimbristylis 5-10 cm reduces N losses and at the
miliacea, and Scirpus maritimus are same time reduces the availability of N
adapted to the wet conditions of direct to weed seedlings that germinate near
seeded flooded rice. the soil surface.
Irrigated rice 77
Table 6.2. Herbicides suitable for use in irrigated rice direct seeded on puddled soil.
Rate
Direct seeded on
Herbicide
(kg ai/ha)
Comments and source of information
dry soil
Bensulfuron 0.05 Apply 6-8 d after seeding (DAS) Dry seeded irrigated rice culture is
(IRRI 1985).
Bentazon 2.0 Apply postemergence to control practiced in Africa, Australia, Europe,
broadleaf weeds and sedges, and the USA. Nongerminated seeds
including S. maritimus. Water level are broadcast or drill seeded in dry or
must be lowered for good coverage.
Annual weeds must be small—2- to moist soil. Broadcast seeds are covered
7-leaf stages (IRRI 1984). by harrowing. More seeds are required
Bifenox + 2,4-D 2.0 + 0.6 Apply at early postemergence of for broadcast than for drill seeding,
weeds, about 4-6 DAS (IRRI 1981).
Butachlor 0.75 Apply 6 DAS to control annual grasses and stand establishment is poorer with
and sedges. Soil should be saturated broadcast seeding than with drill
at application and remain nonflooded seeding.
for 3 d after application (IRRI 1983).
Granular butachlor applied 3 DAS
gives better weed control, less stand Weed problems
reduction, and higher yields than when After broadcast or drill seeding rice
applied 6 DAS (IRRI 1986).
Butralin 2.0 Apply 2-3 DAS to control annual into dry soil, the field is irrigated just
grasses (COPR 1976). enough to provide the soil moisture
2,4-D or MCPA 0.5-1.0 Apply 3-4 wk after seeding to control that allows the seeds to germinate.
annual broadleaf weeds and sedges.
Lower water level to expose weeds Flooding the soil would prevent rice
before spraying and reflood within a seedling emergence. Thus, aerobic
few days (COPR 1976). conditions remain ideal for the germi-
Molinate 3.0 Apply 6-7 DAS. Raise water level after
application (IRRI 1977). nation of upland and aquatic weeds,
Oxadiazon 0.75-1.0 Apply preemergence 4-6 DAS. Soil and weed problems are much worse in
must remain moist after application to dry seeded irrigated than in wet
maintain herbicide activity (IRRI 1985).
Oxyfluorfen 0.15-0.25 Apply 3-6 DAS (IRRI 1978, 1982).
seeded rice. Because the water level is
Pendimethalin 0.75-2.0 Apply up to 6 DAS (IRRI 1975, 1982). increased gradually, it is 2-6 wk before
Piperophos + dimethametryn 0.4 + 0.1 Apply 4-6 DAS (IRRI 1977). a continuous flood at 5 cm depth can
Piperophos + 2,4-D 0.3 + 0.2 Apply 6-8 DAS (IRRI 1987).
Pretilachlor + antidote 0.3-0.4 Apply 3 DAS (IRRI 1987). be achieved. Many well-established
Propanil 3.0-4.0 Apply postemergence to control grass upland weeds will survive, making
and broadleaf weeds at the 2- to 5-leaf weed competition more intense in
stages (about 10 DAS). Water level
should be lowered before application
this rice culture than in the cultures
and the field reflooded as soon as described earlier. The fact that rice and
possible (IRRI 1980). weeds germinate together restricts the
Quinclorac 0.3 Apply 6-8 DAS (IRRI 1986).
Thiobencarb 1.5-2.0 Apply about 6 DAS, when grasses have
number of herbicides that can be
1-2 leaves but before the 3-leaf stage of used safely.
grasses and sedges. Keep water low
enough to avoid submerging the rice
plants (IRRI 1972).
Land preparation
Thiobencarb + 2,4-D 1.0 + 0.5 Apply 6-8 DAS (IRRI 1986). Land preparation should provide
weed-free conditions at planting and
favorable conditions for rice growth
Herbicides can be soil-incorporated and development. Land preparation
before sowing rice, applied preemer- and leveling should be thorough
gence to water a few days after sow- because large soil clods will reduce
ing, or applied postemergence before germination of rice seedlings and
weeds reach the 3- to 4-leaf stage. cause irregularity in herbicidal
Table 6.2 and Figure 6.3 outline efficacy. As clods melt down, the
various herbicides and their applica- inner, unexposed soil will allow weeds
tion times for this rice culture. to germinate.
Water management
Good water management is important Hand weeding Herbicides
in controlling weeds in broadcast or Interrow mechanical weeding is not The effects of herbicides are similar for
drill seeded flooded rice. After dry possible in broadcast seeded rice. Even broadcast seeded or drilled dry-sown
seeding, the soil may be intermittently when seeds are drilled, the interrow rice and wet-sown flooded rice.
flooded and drained to allow for rice spacing is so narrow that only hand or Because of water management prob-
emergence. The water level is then hoe weeding is possible. In this rice lems and difficulties in hand weeding,
increased gradually for a few weeks culture, two to three-timely hand herbicides are particularly important
until a continuous flood of 5-cm depth weedings are sufficient to ensure in this rice culture. Covering the seeds
is maintained. optimum yields. The soil disturbance with soil after drill or broadcast
involved, however, can cause as much seeding increases the tolerance of rice
damage to rice as to weeds. The first for herbicides but decreases a rice
weeding may be done between 14 and seedling’s flooding tolerance.
21 d, depending on weed growth, Butachlor, molinate, oxadiazon,
followed by subsequent weedings propanil, and thiobencarb are used as
when necessary.
Irrigated rice 79
6.4 Rice growth stages when
herbicides can be applied in dry-seeded Water seeded Weed problems
irrigated rice. Bars (–) show periods Many weeds and rice will germinate
during which a particular herbicide is Water seeding of rice is practiced in through either soil or water, but not
applied. *Timing of herbicide
application is based on weed
several parts of Asia, including India, through both. Water seeding takes
emergence and growth stage within Sri Lanka, Malaysia, and Thailand. advantage of that by establishing an
the rice growth stage. It is widely practiced in the USA, early water covering to suppress
southern Europe, USSR, and Australia. weeds. Continuous flooding in water
preplant, preemergence, postemer- Pregerminated rice is broadcast seeded rice culture, however, encour-
gence, or postflood treatment. directly onto the flooded field. The ages aquatic weeds. Where continuous
Propanil is widely used as a rice, which is seeded into water 7-10 flooding is not maintained, many
postemergence herbicide, either in cm deep, sinks to the soil, germinates, semiaquatic weeds typical of
addition to or in place of a preemer- and emerges from the water. The field discontinuously flooded, dry seeded
gence treatment. Preplanting herbi- remains flooded at a depth of 7-10 cm fields can be found in water seeded
cidal treatments, such as with until a few weeks before maturity. In ricefields. Under such conditions.
glyphosate, may also control perennial the USA, seeding into continuously E. crus-galli, Lepfochloa sp.,
weeds. Table 6.3 and Figure 6.4 outline flooded fields began in the 1930s as a Aeschynomene virginica, and Sesbania
the common herbicides and their times cultural method to control E. crus-galli exaltata are weed problems.
of application for this culture. and later as part of a program to
control red rice.
Irrigated rice 81
Fertilizer Table 6.4. Herbicides suitable for use in water seeded irrigated rice.
Herbicides
Use of herbicides is essential to
increase the efficiency of other crop
management practices to control
weeds. Because water seeded rice is
grown under continuous flooding, the
ideal times of herbicide application are
preflood, preplanting; postplanting
into the water; and postplanting, post-
emergence above the water.
For foliar-applied herbicides such
as MCPA, bentazon, and propanil, the
weed foliage must be exposed to the
herbicide. The water level may have to
be decreased for 24-48 h to avoid
washing the herbicide off the leaf, to
allow sufficient uptake of the herbi-
cide. The water level should then be
reestablished. Common herbicides of
importance in water seeded rice are
listed in Table 6.4. Time of application
is indicated in Figure 6.5.
Weed control in
rainfed lowland rice
Rainfed lowland rice is grown on between field capacity and satura- Weed problems
about 23% of the world’s rice area tion. When there is no rainfall, Because the amount and distribution
(IRRI 1988b). It accounts for about however, moisture content may drop of rainfall for growing rainfed low-
45% of the rice area in South and below field capacity. With excessive land rice are uncertain, fields may not
Southeast Asia, 22% in Africa, and rainfall, deepwater conditions may remain flooded from planting to
6% in Latin America (De Datta 1981). develop. maturity. In Asia, most rainfed low-
In South and Southeast Asia, rainfed Rainfed lowland rice is classified land ricefields change from upland to
lowland rice dominates the area, into three culture groups according to submerged conditions during the
although its importance differs the crop establishment technique monsoon season. Lack of water
among countries. For example, 76% used. control reduces the effectiveness of
of the rice area in Bhutan is rainfed Transplanted in puddled soil. using water as a tool in weed man-
lowland, 69% in Thailand, 56% in Direct seeded on puddled soil agement. Upland, semiaquatic, and
Bangladesh, 56% in Myanmar, 43% in (broadcast or drill seeded using aquatic weeds all present problems.
Philippines, 37% in India, and 17% in pregerminated seed). Conditions favorable for weed germi-
Indonesia (IRRI 1988b). Direct seeded on dry soil (broad- nation and growth, such as exposure
Most of the rainfed lowland rice cast or drill seeded using nonger- of the soil surface (aerobic conditions)
area of Southeast Asia is in major rice minated seed). and high soil moisture, occur for
deltas, such as the Mekong in Viet- extended periods. Once weeds
nam, the Chao Phraya in Thailand, Transplanted in become established, deeper flooding
the Irrawaddy in Myanmar, and the is needed to reduce weed growth
Ganges-Brahmaputra in India and puddled soil substantially (Moody et al 1986).
Bangladesh. Transplanting is the major crop estab- Transplanting gives rice a head start
Rainfed lowland rice is not lishment method for rainfed lowland over weeds, but uncontrolled weeds
irrigated, but the soil is flooded to a rice in most of tropical Asia. can still reduce rice yields as much as
maximum depth of less than 50 cm Primarily grown as a monsoonal 50%.
during a portion of the crop cycle. crop, rainfed lowland rice is known Weed species of importance in this
Water is supplied by frequent rains as kharif in India and as aman in rice culture include Echinochloa spp.,
during the growing season. Soil northeastern India and Bangladesh. lschaemum rugosum, Monochoria
moisture is usually maintained Seedlings raised by a wet bed, dapog, vaginalis, Sphenoclea zeylanica, Cyperus
or dry bed technique are transplanted difformis, Cyperus iria, Fimbristylis
into a puddled soil. miliacea, and Scirpus maritimus.
Weed problems
Direct seeded rainfed rice is more
susceptible to weed competition than
is transplanted rainfed rice. Although
soil puddling reduces the weed
problem, uncontrolled weeds still
reduce rice yields about 60%. In some
cases, a puddled lowland field may
be saturated but without any stand-
ing water, because of lack of water
control. The moist, warm, aerobic soil
condition created promotes germina-
tion and rapid growth of many
upland, semiaquatic, and aquatic
weeds which are little affected by
7.1 Rice growth stages when herbicides Herbicides later flooding. Deeper-than-normal
can be applied on rainfed lowland rice flooding is often required to signifi-
The efficiency of herbicides on trans-
transplanted on puddled soil. Bars (—)
show periods when a particular herbicide is planted rainfed lowland rice depends cantly reduce weed growth. Because
applied. *Timing of herbicide application is on water management. If the water in this culture, rice and weeds germi-
based on weed emergence and growth nate at the same time, competition by
stage within the rice growth stages.
depth in the field exceeds 10 cm
during the first week after herbicide weeds is more intense than it is in
application, herbicidal efficacy will be transplanted rice.
Hand weeding reduced due to dilution and leaching. Weeds of importance in this
Hand weeding is the most common Despite water management limita- culture include E. crus-galli and other
weed control method in rainfed tions, however, weeds in rainfed Echinochloa spp., I. rugosum,
lowland transplanted rice. The field lowland rice can be adequately M. vaginalis, S. zeylanica, C. difformis,
should be weed free for 30 d after controlled by herbicides. C. iria, F. miliacea, and Scirpus spp.
transplanting (DT) to prevent yield Herbicides that can be used as an
losses caused by weeds. Two to three alternative or supplement to manual
properly timed hand weedings, the or mechanical weeding include 2,4-D,
first about 21 DT, combined with MCPA, butachlor, thiobencarb,
good water management will ensure propanil, oxadiazon, pendimethalin,
optimum rice yields. Mechanical piperophos, and oxyfluorfen. Table
weeders can be used in rice trans- 7.1 and Figure 7.1 provide informa-
planted in rows, but weeds within tion on available herbicides and
the rows still have to be removed by timing of their application.
hand.
Weed problems
Among rainfed lowland rice cultures,
weed problems in direct seeded rice
are more intense and wider in range
than those in transplanted or broad-
cast rice where puddling reduces
7.2 Rice growth stages when
herbicides can be applied on wet-seeded dependent on soil moisture condi- weed problems. Rice yield losses
rainfed lowland rice. Bars (—) show tions and is markedly reduced by dry from uncontrolled weeds can be as
periods when a particular herbicide is
or deep flooding conditions immedi- high as 74%. Rice-weed competition
applied. *Timing of herbicide application
is based on weed emergence and growth ately after herbicide application. In for moisture is heavy during the early
stage within the rice growth stage. the tropics, butachlor, thiobencarb, growth stage, when there is no
butralin, and propanil have been standing water. Dry rice seeds germ-
used in direct seeded rice. Table 7.2 inate 3-5 d later than pregerminated
Hand weeding and Figure 7.2 provide herbicide rice seeds. Weeds germinate and
Hand pulling of weeds can be done information for this rice culture. establish faster than rice.
in broadcast or drill seeded rice; The prevailing moist, aerobic
mechanical weeding is feasible only Direct seeded condition for direct seeded rice
when rice is planted in rows. Two to encourages the growth of upland,
three hand weedings at 2-3 wk after on dry soil semiaquatic, and aquatic weeds.
sowing are usually sufficient to Direct seeding on dry soil provides Dominance of any of these weed
ensure optimum yields. an opportunity to increase cropping communities depends on the availa-
intensity in rainfed lowland rice. bility and depth of standing water.
Herbicides Eliminating puddling shortens land Many well-established upland weeds
The use of herbicides for weed use time. In this culture, dry seed is continue to survive under flooding
control has proved effective and sown directly into moist, nonpuddled later in the crop cycle. Lowland
economical in direct seeded rainfed soil at the beginning of the rainy weeds that are important include
lowland rice. Because weeds and rice season. The field may be bunded to E. crus-galli, I. rugosum, L. chinensis,
germinate together, the number of accumulate water as the rainy season C. difformis, F. miliacea, and
herbicides that can be used safely is progresses and the crop may end its S. maritimus.
limited. The efficacy of herbicides is cycle under flooding.
Upland rice, also known as dryland or Upland rice, like all upland crops, is upland weeds include Cyperus rotun-
pluvial rice, is grown on rainfed, planted in moist soil, that, in general, dus, Echinochloa colona, Eleusine indica,
naturally well-drained soils. Strictly does not retain moisture beyond field Rottboellia
defined, upland ricefields are not capacity. Water is supplied by rains cochinchinensis, Cynodon dactylon,
bunded and no surface water accumu- during the growing season. Optimum Digitaria sanguinalis, Imperata cylindrica,
lates. temperature, sufficient aeration, and Amaranthus spinosus, Commelina
About 13% (18.8 million ha) of the ideal moisture for weed germination benghalensis, Trianthema portulacastrum,
world's rice area is upland (IRRI and growth exist at planting time. This Ageratum conyzoides, Portulaca oleracea,
1988b). About 11.9 million ha is in enables weeds to germinate earlier and and Euphorbia hirta.
Asia, 4.5 million ha in Latin America, grow more vigorously than the rice
and 2.2 million ha in Africa. It is the crop. Land preparation
dominant rice culture in Latin America Weed competition is more intense Land preparation for upland rice
and West Africa. in upland rice than in irrigated and varies greatly among regions. In West
Upland rice is grown under a wide rainfed lowland rice because upland Africa, where shifting cultivation is
range of management practices that fields do not have standing water to common, slash-and-burn is practiced
vary from shifting cultivation-as suppress weed growth. Some weeds in with hand tools. In South Asia, upland
practiced in Malaysia, Philippines, upland rice can withstand drought fields are plowed by bullocks and in
Peru, and West Africa-tothe mecha- better than rice because their roots Southeast Asia, by water buffalo.
nized cultivation practiced in Brazil. penetrate deeper into the soil to tap Deep plowing (25 cm or deeper)
Most of the world’s upland rice is moisture. Poor rice germination due to moist soil at the end of the rainy
grown on poor soils in areas with drought results in excessive weed season is recommended for upland
uncertain rainfall by small farmers growth, especially if semidwarf rice grown in the West African savan-
using traditional, low-input techno- varieties are grown. nahs (FAO 1976). Deep plowing and
logy. Because weeds in upland rice subsoiling conserve soil moisture in
germinate throughout the season, the rainy season and enhance root
Weed problems dense weed growth may reoccur after growth and extraction of soil moisture
Weeds rank second to drought stress hand weeding or after the residual from deeper soil layers. It also will
in reducing upland rice grain yields effects of herbicides have worn off. bury weed seeds deep enough to pre-
and quality (Sankaran and De Datta A mixture of annuals and peren- vent them from emerging. Subsequent
1985). Yield losses caused by uncon- nials, and grasses and broadleaf tillage operations must be shallow.
trolled weeds in upland rice are about weeds, intensifies the competitive
96%. effects of weeds in upland rice. C4
weeds, which have higher water-use
efficiency than rice, prevail (Ampong-
Nyarko and De Datta 1989). Common
Upland rice 91
An upland field should be Latin America. Dibbling, or hilling, is Crop rotation is practiced to pre-
harrowed to break up soil clods, but practiced in Africa and Asia by farm- vent the buildup of weeds adapted to
field leveling is not critical. Rice ers using slash-and-burn systems. A upland ricefields, but easy to control in
should be planted as soon as possible pointed stick is used to make holes in other crops. Herbicides that control
following the last harrowing to pro- the soils; 4-8 unsprouted seeds are problem weeds but are toxic to rice
vide rice an even start with the weeds. dropped in and covered with soil. In also can be used on a tolerant crop in
The stale seedbed technique can Latin America, mechanized drilling is the rotation. That will reduce the rice
reduce weed problems in upland rice. becoming increasingly popular. weed problem.
After land preparation, weeds that Row drilling or dibbling in rows
grow are killed at the 2- to 5-leaf stage makes weeding and other manage- Fertilizer
by herbicides or using mechanical ment practices easier. Timely sowing Nitrogen response is high for modern
methods. Germination of most of the and rapid canopy closure minimize short- to medium-statured upland rice
viable weed seeds is essential to the weed growth and ensure good stand cultivars that are resistant to lodging.
success of the stale seedbed technique. establishment. In most soils, split application of N
Under unfavorable conditions for Weed problems in broadcast seeded gives higher grain yield than does
weed seed germination (e.g., dry fields are higher because mechanical single basal application.
soils), no weed control advantage may hand weeding cannot be done, and Applying fertilizer to upland rice,
be achieved. weeding must be delayed because it is however, will reduce grain yield if the
Use of the stale seedbed technique difficult to tell grassy weeds from rice field is not weeded. The first applica-
should not delay rice seeding beyond at early growth stages. tion should be delayed until after
the optimum time. Planting at the weeding. Subsequent topdressings of
optimum time assures more rainfall Cultivar N after weeding will maximize fertil-
from seeding through spikelet filling; Upland rice cultivars with drought izer-use efficiency. The N application
late-planted rice is likely to suffer avoidance (through deep root sys- rate should be reduced in drought-
drought and reduced solar radiation tems) and drought recovery abilities prone areas.
during the reproductive stage, which are preferred. Intermediate-statured Phosphorus deficiency is common
can substantially reduce yields. cultivars with moderate tillering, big in upland rice, especially in Oxisols
Zero tillage can be used to establish panicles, blast resistance, and tolerance and Ultisols in Brazil, West Africa, and
an upland rice crop where no difficult- for iron deficiency and aluminum some parts of South and Southeast
to-control perennial weeds are found toxicity are also desirable. Asia (Gupta and O'Toole 1986). In the
in the fallow vegetation or when Philippines, 18 kg P/ha is recom-
appropriate herbicides are available. Plant population mended for upland rice (PCARR
But zero tillage may not be successful Seeding rate and spacing for upland 1977). Some coarse-textured soils in
under all soil conditions; restricted rice vary with planting method and high-rainfall areas are affected by
root development and reduced grain the rice cultivar used. A high plant potassium deficiency. Applying
yield have been observed (Stone et a1 population is important for upland 33 kg K/ha has been adequate in some
1980). rice to quickly develop a canopy that African countries (IRCN 1982). Zinc,
will suppress weed growth. Seeding iron, and sulfur deficiencies also occur
Planting methods rates vary from 80 to 150 kg/ha, in upland rice. Deficiency of any of
Broadcasting, dibbling, and drilling depending on the seeding method. these nutrients will reduce the vigor of
are the common seeding practices for Row spacing also varies between 20 rice and, hence, its competitiveness
upland rice. Broadcasting is common and 30 cm. Broadcast seeding requires against weeds.
in many countries in Asia, Africa, and more seeds than drilling or dibbling.
Tall leafy cultivars should be planted
at wider spacing than semidwarf culti-
vars. Growing tall cultivars at narrow
spacing increases lodging.
Upland rice 93
Herbicides Table 8.1. Herbicides suitable for use in upland rice.
Chemical weed control in upland rice
is economical and effective under Herbicide
Rate
Comments and source of information
(kg ai/ha)
certain conditions. It may be the only
weed control method feasible for
Bentazon 1.0-2.0 Selective postemergence control of
large-scale rice farms. Herbicides can certain broadleaf weeds and sedges
complement other weed control (IRRI 1982). Does not control grasses.
Apply postemergence when weeds are
methods or can be used in combina-
at the 4- to 10-leaf stages. Delayed
tion with hand weeding to give application allows the weeds to exceed
acceptable weed control. maximum size, resulting in inadequate
For optimum effectiveness of pre- control.
Bifenox 2.0 Apply preemergence from seeding to
emergence herbicides, which require early spikelet stage. Apply to moist
moist or wet soil, timely rains after soils for best results. Bifenox +
application are essential. When resid- propanil can be applied post-
emergence when rice has 2-3 leaves
ual herbicides are applied to soils that and weeds are 2.5 cm high (Akobundu
have remained dry for a long time, 1987).
they give variable results. The persis- Butachlor 2.0 Effective against grasses and broad-
leaf weeds; less effective against
tence of many preemergence herbi- perennial sedges (IRRI 1975).
cides is so short, they cannot control Butralin 2.0 Apply preemergence to control annual
successive flushes of weed seedlings. grasses (IRRI 1976).
2,4-D or MCPA 0.5 Apply as postemergence spray after
This makes follow-up hand weeding tiller initiation, about 21-30 d after
or application of a postemergence seeding (DAS) (Dixit and Singh 1981).
Dinitramine 1.5 Apply preemergence (IRRI 1978).
herbicide necessary.
Fluorodifen 2.5-4.0 Apply 0-3 DAS for residual pre-
Among the preemergence emergence control of annual grasses
herbicides, butachlor, oxadiazon, (De Datta 1972).
dinitramine, pendimethalin, Oxadiazon 0.75-1.5 Apply preemergence 0-2 DAS. Rice
growth depression has been observed
thiobencarb, fluorodifen, and 15-20 d after emergence (IRRI 1975).
piperophos-dimethametryn have been Oxyfluorfen 0.35 Apply preemergence. May be
widely tested. Postemergence herbi- moderately toxic to rice (IRRI 1983).
Pendimethalin 2.0 Apply preemergence or early post-
cides such as propanil and MCPA are emergence,
also effective. Herbicide combinations combined with propanil (IRRI 1975).
are often more effective than a single Excellent for control of Rottboellia
cochinchinensis.
herbicide. Applying a preemergence Piperophos + 1.0-2.0 Apply 4-6 DAS (Dubey et al 1980).
herbicide such as oxadiazon, dimethametryn
Propanil 3.0-6.0 Apply at 2- to 3-leaf stages of grass.
butachlor, or thiobencarb, followed by
High temperatures increase contact
propanil 15-25 d later will provide burn and may injure rice (IRRI 1980).
better weed control than will any of Thiobencarb 3.0 Apply before 2-leaf stage of weeds
and after 1-leaf stage of rice
these herbicides applied alone. Early
(IRRI 1988a).
postemergence application of a poste-
mergence herbicide (eg, propanil)
combined with a residual herbicide,
done at the time the postemergence
herbicide is most effective, has the
advantages of providing season-long
control and saving time and labor by
combining two spraying operations.
Table 8.1 and Figure 8.1 provide
information on common herbicides
that may be used in upland rice.
Deepwater and floating rice constitute that successively compete with the rice seed or transplant deepwater rice,
about 11% of the world’s rice area crop. Weeds adapted to dryland puddling reduces weed infestation.
(IRRI 1988b). They are grown in the compete at early crop stages, die upon Amphibious weeds, such as the
deltas, estuaries, and river valleys of flooding, and are succeeded by aquatic wild rices, Scirpus sp., Sesbania sp., and
Bangladesh, Cambodia, India, weeds. Leersia hexandra, can compete with rice
Indonesia, Myanmar, Thailand, and Before flooding, deepwater and under dryland or flooded conditions.
Vietnam. In Africa, deepwater and floating rice may be infested by Eichhornia crassipes, Ipomoea aquatica,
floating rice-growing areas are found Echinochloa colona, Eleusine indica, Monochoria vaginalis, and Pistia
in the inland Niger River delta in Mali, Cyperus rotundus, C. iria, and stratiotes are found in deepwater rice
in the Niger River delta, Nigeria, and C. difformis. These weeds also may after flooding has occurred.
in inland swamps. compete with rice during the early Weeds such as the wild rices
stages of flooding. possess flood tolerance and elongation
ability, and grow with rice in rising
Deepwater and floating Deepwater and floating rice may
suffer severe weed infestations during floodwater. Therefore, they are able to
rice cultures the preflood period. Weed problems cause considerable yield losses. Masses
Deepwater rice is grown in areas are often aggravated by poor stand of E. crassipes are usually introduced to
where the maximum depth of stand- establishment due to drought. At the the ricefield by water currents or
ing water exceeds 50 cm for a signifi- preflood stage, weeds reduce growth strong winds. When this occurs, E.
cant period of rice growth. Where and tillering of rice plants and can crassipes can completely smother the
water depth is greater than 100 cm, the reduce grain yield as much as 33% (De rice crop within a few days. In West
culture is referred to as floating rice or Datta and Hoque 1982). Total crop loss Africa, the most troublesome weeds
very deep water rice. Fields are not can occur if flooding is below normal. include E. stagnina and E. pyramidalis,
bunded and flooding usually occurs Minimizing competition from and the wild rices Oryza longistaminata
only during the later part of the grow- weeds at early rice growth stages and O. barthii.
ing season. Rice may grow under largely determines how well deep-
drought conditions for 40-60 d before water and floating rice will tolerate Weed control before
flooding occurs. stresses caused by flooding at later
stages. Aquatic weeds that occur upon flooding
Weed problems flooding are less a problem when The deepwater rice cycle has two
Weeds are a major problem in deep- stand establishment is good and rice distinct phases—before flooding and
water and floating rice because rice tillering is adequate than when early after flooding. Weed control methods
normally is seeded into soils that may weed competition is severe. In small can be described best within these
remain dry for 6-12 wk. Fields may cropping areas where farmers wet phases. Deepwater rice before flooding
also be flash flooded. The dry soil and is treated either as rainfed lowland rice
intermittent flash flooding result in a or as upland rice, using the same weed
broad spectrum of weeds-from control methods.
upland to semiaquatic and aquatic-
Hand weeding
Attempts to weed deepwater and
floating rice from boats-adifficult
task-havebeen made in cases of
severe weed infestation. In Mali,
farmers hand weed after flooding to
minimize competition from
O. longistaminata and O. barthii.
Perennial E. stagnina is also removed
by hand.
Herbicides
Herbicides are effective in deepwater
and floating rice only before flooding.
Management of some
difficult weeds in rice
Difficult weeds 99
Paspalum distichum Echinochloa species Hand weeding
Hand weeding is effective if done
Paspalum distichum is a creeping peren- The genus Echinochloa, which includes repeatedly, to remove succeeding
nial grass found in lowland ricefields about 50 weed species, includes some flushes of weeds and weeds over-
(see page 22). Its underground growth of the most important rice weeds. The looked earlier due to their similarity to
system consists of adventitious roots most common are E. crus-galli, rice.
and rhizomes. It tends to resist con- E.glabrescens, E.oryzoides,
ventional weed control measures, E. pyramidalis, and E. colona. Herbicides
including the use of herbicides, but is E. crus-galli grows widely in both Several herbicides can selectively
sensitive to shading. temperate and tropical regions (see control Echinochloa spp. This include
page 16). Its world distribution ranges butachlor, oxadiazon, oxyfluorfen,
Cultural control from 50° N to 40º S latitude. E. colona pendimethalin, thiobencarb, simetryn,
Thorough land preparation is one way (see page 33) occurs in tropical and molinate, propanil, chlomethoxynil,
of controlling P. distichum. Frequent subtropical regions. E. glabrescens is pretilachlor, and quinclorac. Preemer-
tillage reduces P. distichum problems. found in the Indian subcontinent, gence and early postemergence appli-
Cutting up rhizomes by tillage encour- Southeast Asia, China, and southern cations are most effective because the
ages dormant buds to sprout, deplet- Japan. E. pyramidalis is abundant in the weeds are most susceptible at the
ing the weeds food reserves. For long- floating rice areas of West Africa. seedling stage. Weed resistance to
term control, its rhizome must be E. crus-galli prefers moist conditions herbicides increases with age.
killed. and continues to grow when Butachlor applied preemergence
Because P. distichum is sensitive to submerged; E. colona ceases to grow inhibits the enzyme activity of
shade, closely spaced, vigorous rice when submerged. Echinochloa spp., delays radicle
plants offer better competition against Rice yield losses from season-long emergence, and inhibits emergence of
this weed than do widely spaced competition with Echinochloa spp. can the root and primary leaf. Selectivity
plants. be as high as 90%. Echinochloa spp. are for propanil depends on the different
troublesome in rice because their levels of the hydrolyzing enzyme,
Herbicides ecological requirements are similar: at which is high in rice but low in
P. distichum is resistant to many pre- early growth stages, they resemble Echinochloa spp.
emergence herbicides used on rice. It rice, and they accumulate considerable
is, however, susceptible to glyphosate amounts of nutrients, to the disadvan- Integrated control
applied at 2.0 kg ai/ha and moder- tage of rice. Moreover, these weeds Echinochloa spp. can be managed
ately susceptible to paraquat. A pre- compete with rice for light and effectively through an integrated
plant herbicide, such as glyphosate, moisture. approach. Because the weed is sensi-
used in combination with land prepa- Echinochloa spp. germinate earlier tive to shade, a vigorous rice stand
ration, improves control. Translocation than direct seeded rice. During the (achieved with high planting density
of glyphosate into the rhizomes is best first 3 wk of weed growth, profuse and high fertilizer application),
achieved when the glyphosate is emergence of new leaves and vegeta- combined with hand weeding or with
applied on actively growing tive growth of tillers and adventitious preemergence or early postemergence
P. distichum with maximum leaf area. roots occur. Echinochloa spp. produce herbicide, will lead to early rice
extremely large numbers of seeds, canopy closure. Late-germinating
Integrated control which ensure the weeds dispersal and Echinochloa spp. can be controlled by
Integration of cultural control and reestablishment. herbicides. A 10-20 cm water depth
herbicides will give good control of will generally suppress Echinochloa
P. distichum. Cultural control spp. weeds.
Crop rotation can effectively reduce
Echinochloa spp. populations.
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Anderson W P (1983) Weed science Consortium for International Crop Oregon State University, Corvallis,
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Important crops of the world and their Sons, New York, Chichester, Brisbane, Survey, England.
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kusen Federal Republic of Germany. control in deepwater rice areas. Pages Green D H, Ebner L (1972) A new selective
1465 p. 427-442 in Proceedings of the 1981 herbicide for rice S-(2-methyl-l-
Bernasor P C, De Datta S K (1986) Chemi- international deepwater rice workshop. piperidyl-carbonylmethyl)-0,0-di-N-
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(Scirpus maritimus L.) and annual weeds P.O. Box 933, Manila, Philippines. or in mixtures. Pages 822-829 in
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References 105
Appendices
A. Useful conversions
Weights
1 gram = 0.03527 ounces
1 ounce = 28.4 grams
1 kilogram = 2.2 pounds
1 pound = 454 grams (0.454 kilogram)
1 metric ton = 2,204.6 pounds (1,000 kilograms)
Area
1 acre = 4,840 sq yards = 43,560 sq feet = 0.405 hectare = 4,050 sq meters
1 hectare = 2.41 acres
Volume
1 gallon (Imperial) = 4.546 liters
1 gallon (Imperial) = 1.2 U.S. gallons
1 gallon (U.S.) = 3.785 liters
1 fluid ounce = 28.4 milliliters
1 liter = 0.22 gallons = 1.76 pints
Measurements
1 inch = 2.54 centimeters
1 foot = 0.305 meter
1 yard = 0.914 meter
1 mile = 1.609 kilometers
1 centimeter = 0.394 inch
1 meter = 3.281 feet
1 meter = 1.094 yards
1 kilometer = 0.621 mile
1 kilopascal (kpa) = 0.145 pound per square inch (psi)
Quick conversions
1 liter/hectare = 0.089 gallon/acre
1 kilogram/hectare = 0.892 pound/acre
1 kilogram/liter = 8.33 pounds/gallon
1.0 1.0 1.11 1.25 1.33 1.45 1.66 2.0 2.5 3.33 4.0 5.0
1.5 1.5 1.66 1.87 2.0 2.18 2.49 3.0 3.75 5.0 6.0 7.5
2.0 2.0 2.22 2.5 2.66 2.9 3.32 4.0a 5.0 6.60 8.0 10.0
2.5 2.5 2.78 3.13 3.33 3.63 4.15 5.0 6.25 8.33 10.0 12.5
3.0 3.0 3.33 3.75 4.0 4.35 4.98 6.0 7.5 9.99 12.0 15.0
3.5 3.5 3.89 4.38 4.66 5.08 5.81 7.0 8.75 11.66 14.0 17.5
4.0 4.0 4.44 5.0 5.32 5.80 6.64 8.0 10.0 13.32 16.0 20.0
4.5 4.5 5.0 5.63 6.0 6.53 7.5 9.0 11.25 15.0 18.0 22.5
5.0 5.0 5.55 6.25 6.67 7.14 8.33 10.0 12.5 16.67 20.0 25.0
5.5 5.5 6.11 6.88 7.32 8.0 9.13 11.0 13.75 18.32 22.0 27.5
6.0 6.0 6.66 7.5 7.98 8.7 7.96 12.0 15.0 10.0 24.0 30.0
a For example, to apply 2.0 kg ai/ha using a formulation containing 50% active ingredient, use 4.0 liters of the formulated product.
Appendices 107
B. Conversion table for granular formulations
0.5 2.5 3.33 5.0 6.67 10.0 12.5 16.67 25.0 50.0
1.0 5.0 6.67 10.0 13.3 20.0 25.0 33.3 50.0 100.0
1.5 7.5 10.0 15.0 20.0 30.0 37.5 50.0 75.0 150.0
2.0 10.0 13.3 20.0 26.6 40.0a 50 66.7 100.0 200.0
2.5 12.5 16.6 25.0 33.3 50 62.5 83.3 125.0 250.0
3.0 15.0 20.0 30.0 40.0 60 75.0 100.0 150.0 300.0
3.5 17.5 23.3 35.0 46.6 70 87.5 116.7 175.0 350.0
4.0 20.0 26.6 40.0 53.3 80 100.0 133.3 200.0 400.0
4.5 22.5 30.0 45.0 60.0 90 112.5 150.0 225.0 450.0
5.0 25.0 33.3 50.0 66.6 100 125.0 166.7 250.0 500.0
5.5 27.5 36.3 55.0 73.3 110 137.5 183.3 275.0 550.0
6.0 30.0 40.0 60.0 80 120 150.0 200.0 300.0 600.0
a For example, to apply 2.0 kg ai/ha using granules containing 5% active ingredient, use 40 kg offormulated product.
Year Original
Common name Trade names
introduced manufacturer
Appendices 109
Index
Index 111
in deepwater and floating rice, 96 Oryza rufipogon Griff., 1,101 classification, 83
in dryland, 42 Oryza sativa L., 1,101 distribution, 87
in wetland, 42 Oxadiazon, 66, 67, 71, 75, 77, 78, 79, 80, 84, 85, 86, time of seeding, 42
limited, 42 87, 88, 89, 93, 94, 96, 97, 100 yield loss due to weeds, 1
Latin America, 42, 83, 91, 92, 102 Oxisols, 92 weed control in, 83-89
Leaching of herbicides, 51, 53 Oxyflourfen, 49, 66, 67, 69, 75, 77, 78, 84, 85, 86, 87, Rainbow trout, 67, 68, 69, 71
Leersia hexandra Sw., 2, 20, 66 88, 89, 93, 94, 97, 100 Rats, 68, 69, 70,71
Leptochloa chinensis (L.)Nees., 3, 21, 66, 87 Red rice, 2,80,101
Leptochlou spp., 2, 80 P Residual herbicide, 52
Light, Pacific Islands, 102 nozzle type for, 61,58
for rice growth, 3 Pakistan, 9, 11, 24, 29, 31, 37, 73 Rice
rice-weed competition, 3 Paraquat, 49, 50, 62, 66, 67, 68 diseases, 2
penetration, 43 Partition coefficient, 53 drought tolerance, 4
Lodging, Paspalum paspalodes (Michx)Scribn., 22 growth requirements, 3
Lowland rice weeds, 9-24, 42 Paspalum distichum L., 2, 7, 22, 41, 66, 67, 100 herbicide tolerance, 72
Ludwigia octovalvis (Jacq.) Raven, 15, 66 Pendimethalin, 66, 67, 68, 72, 73, 75, 78, 79, 80, 84, modern varieties, 42
85, 86, 88, 89, 93, 94, 97, 100, 101, 103 mutation, 3
M Perennial weeds, 1 susceptibility to herbicide, 50, 72
Machete, 55, 59, 62 effect of herbicide, 50 similarities with weeds, 4, 5
Madagascar, 35 reproduction, 5 traditional varieties, 42
Malaysia, 9, 10, 12, 14, 15, 16, 19, 20, 22, 23, 24, 27, under limited tillage, 42 wild species of, 2
29, 30, 33, 34, 37, 38, 80, 91, 101 Persistence of herbicide, 52 Rice culture classification, 7
Mali, 95 Peru, 9, 13, 16, 19, 32, 39, 91 effect on weed flora, 1
Manufacturer of herbicides, 55 Pest, Rice dwarf, 2
Marsileaceae, 14 Phenoxy acetic acids, 69,72 Rice grassy stunt virus, 2
Marsilea crenata Presl., 14 Philippines, 9, 10, 11, 12, 13, 14, 15, 16, 18, 19, 20, Rice maja blanca, 2
Marsilea minuta L., 14, 66 21, 22, 23, 24, 25, 26, 27 Rice mimics, 5,44
Mauritius, 28, 31, 35, 37 28, 29, 30, 31, 32, 33, 34, 35, 36, 37, 38, 39, 43, 83, 85, Rice production cost, 1
MCPA, 50, 63, 66, 67, 69, 71, 72, 75, 77, 78, 80, 81, 91,92 Rice stripe, 2
82, 84, 85, 86, 87, 88, 89, 93, 94, 96, 97 Phloem, 50 Rice tungro virus, 2
Mechanical transplanters, Phosphoms, 3, 4, 43-44, 92 Rice-weed competition, 2
Mechanical weeding, 44 Photodegradatio of herbicide, 52 critical periods, 4
Meloidogyne, 2 Photosynthesis, 3 factors, 3-3
Metolachlor, 101 C3 pathway, 3,4 in direct seeded flooded rice, 77
Metribuzine, 101 C4 pathway, 3,4 factors, 3-4
Mexico, 9, 25, 26, 29, 30, 34, 37 Physalis angulata, 66, 67 Rice yellow dwarf, 2
Mimics of rice, 5,44 Phytotoxicity, 52, 68 Rottboellia exaltata L.f., 36
Minimum tillage, Pictograms, 55 Rottboellia cochinchinensis (Lour.)W.D. Clayton,
Molinate, 66, 67, 68, 70, 71, 75, 78, 79, 80, 81, 82, 84, Piperophos, 66, 67, 69, 71, 72, 75, 78, 79, 84, 85, 86, 3,4,36,66,67,91,103
100,101 93, 94, 96, 97, 98 Runoff of herbicide, 52
Monochoria vaginalis (Burm.f.)Presl, 2, 3, 23, 66, Pistia stratiotes, 95
73,83,85,95 Placement of herbicide, 49 S
Monocotyledons, 7 Plant height, advantages of, 4, 42 Safety,
Myanmar, 9, 12, 19, 20, 26, 28, 30, 32, 34, 37, 38, 83, Planting method, 42, 74, 76, 79, 86, 88, 91, 96 Samoa, 28
95 Plant population, 74, 76, 81, 84, 86, 91, 96 Scirpus maritimus L., 13, 66, 67, 71, 73, 83, 87, 99
Poaceae, 16-22, 31-36 Scirpus spp., 85, 95
N Poisoning, 63 Seed, dispersal, 4-5
Nematodes, 2 Polycyclic alkanoic acids, 71 dormancy, 5
Nicaragua, 32 Pontederiaceae, 7, 23, 38 growth, 3
Niger, Portulacaceae, 37 longevity,
Nigeria, 10, 25, 26, 27, 30, 34, 35, 37, 95 Portulaca oleracea L., 37, 66, 67,91 production, 4-5
Nilapurvata lugens, 2 Postemergence herbicides, 51, 65 Seeding, methods for rice, 7
Nitrogen, 3, 4, 74, 76, 84, 86, 88 Potamogetonaceae, 7 rate, 79
recommended timing of application, 43 Potassium, 3, 4, 43, 92 timing, 42
Nontilled rice fields, 92 Preemergence herbicides, 51, 65, 76 Selectivity of herbicide, 49-50, 51, 54
effect on weed flora, 1 Preplanting herbicides, 50 Senegal, 29
Nozzles, 56, 57, 58, 61, 63 Pressure chamber, 57 Sequential application, 65
Nurseries, 84 Pretilachlor, 49, 50, 66, 67, 73, 78, 85, 86, 88, 100 Sesbania aculeata, 84, 98
Nutrient uptake, 3, 4, 43, 44 Prevention of weed introduction, 41 Sesbania exaltata, 80
Nymphae stellata, 66 Propanil, 49, 51, 52, 66, 67, 68, 69, 71, 72, 73, 75, 77, Sesbania sp., 95
Nymphula depunctalis, 2 78, 79, 80, 81, 82, 84, 85, 86, 87, 88, 89, 93, 94, 97, Setaria glauca, 66, 67
96, 99, 100 Setaria viridis, 2
O Propazine, 101 Shading, 3
Onagraceae, 15 Protective clothing, 62-63 effect on rice growth, 4
Organophosphorus compounds, 69 Sierra Leone, 10
insecticides, 50 Q Signal words, 55
Oryza barthii A.Chev., 1, 2, 67, 95, 98, 101 Quality, Simetryn, 52, 70, 100
Oryza glaberrima, 1, 101 Quinclorac, 66, 67, 72, 73, 75, 78, 84, 86, 88, l00c Slash and burn, 91
Oryza longistaminata A.Chev. & Roehr., 1, 2, 41, Soil, effect on C 3 -C4 competition, 3
67,95,97,98,101 R herbicide activity,
Oryza nivara Sharma & Shastry, 1, 101 Radiation from ultraviolet, 52 moisture, 51, 52
Oryza officinalis Wall. ex Watt., 101 Rainfed rice pH,
Oryza punctata Kotschy ex Steud., 101 N application timing, residual herbicide activity,
Index 113