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a.AccordingtoDarwinstheoryofnaturalselection,competitionforlimitedresourcesresultsin
differentialsurvival.Individualswithmorefavorablephenotypesaremorelikelytosurviveand
producemoreoffspring,thuspassingtraitstosubsequentgenerations.
b.Evolutionaryfitnessismeasuredbyreproductivesuccess.
c.Geneticvariationandmutationplayrolesinnaturalselection.Adiversegenepoolis
importantforthesurvivalofaspeciesinachangingenvironment.
d.Environmentscanbemoreorlessstableorfluctuating,andthisaffectsevolutionaryrateand
directiondifferentgeneticvariationscanbeselectedineachgeneration.
e.Anadaptationisageneticvariationthatisfavoredbyselectionandismanifestedasatrait
thatprovidesanadvantagetoanorganisminaparticularenvironment.
f.Inadditiontonaturalselection,chanceandrandomeventscaninfluencetheevolutionary
process,especiallyforsmallpopulations.
g.ConditionsforapopulationoranalleletobeinHardyWeinbergequilibriumare:1.alarge
populationsize,2.absenceofmigration,3.nonetmutations,4.randommating,and5.absence
ofselection.Theseconditionsareseldommet.
h.Mathematicalapproachesareusedtocalculatechangesinallelefrequency,providing
evidencefortheoccurrenceofevolutioninapopulation.
1.A.2:Naturalselectionactsonphenotypicvariationsinpopulations.
a.Environmentschangeandactasselectivemechanismonpopulations.
b.Phenotypicvariationsarenotdirectedbytheenvironmentbutoccurthroughrandomchanges
intheDNAandthroughnewgenecombinations.
c.Somephenotypicvariationssignificantlyincreaseordecreasefitnessoftheorganismandthe
population.
d.Humansimpactvariationinotherspecies.
1.A.3:Evolutionarychangeisalsodrivenbyrandomprocess.
a.Geneticdriftisanonselectiveprocessoccurringinsmallpopulations.
b.Reductionofgeneticvariationwithinagivenpopulationcanincreasethedifferencesbetween
populationsofthesamespecies.
1.A.4:Biologicalevolutionissupportedbyscientificevidencefrommanydisciplines,including
mathematics.
a.Scientificevidenceofbiologicalevolutionusesinformationfromgeographical,geological,
physical,chemicalandmathematicalapplications.
b.Molecular,morphologicalandgeneticinformationofexistingandextinctorganismsaddtoour
understandingofevolution.
1.B.2:Phylogenetictreesandcladogramsaregraphicalrepresentations(models)of
evolutionaryhistorythatcanbetested.
a.Phylogenetictreesandcladogramscanrepresenttraitsthatareeitherderivedorlostdueto
evolution.
b.Phylogenetictreesandcladogramsillustratespeciationthathasoccurred,inthatrelatedness
ofanytwogroupsonthetreeisshownbyhowrecentlytwogroupshadacommonancestor.
c.Phylogenetictreesandcladogramscanbeconstructedfrommorphologicalsimilaritiesof
livingorfossilspecies,andfromDNAandproteinsequencesimilarities,byemployingcomputer
programsthathavesophisticatedwaysofmeasuringandrepresentingrelatednessamong
organisms.
d.Phylogenetictreesandcladogramsaredynamic(i.e.,phylogenetictreesandcladogramsare
constantlybeingrevised),basedonthebiologicaldataused,newmathematicaland
computationalideas,andcurrentandemergingknowledge.
1.C.1:SpeciationandextinctionhaveoccurredthroughouttheEarthshistory.
a.Speciationratescanvary,especiallywhenadaptiveradiationoccurswhennewhabitats
becomeavailable.
b.Speciesextinctionratesarerapidattimesofecologicalstress.
1.C.2:Speciationmayoccurwhentwopopulationsbecomereproductivelyisolatedfromeach
other.
a.Speciationresultsindiversityoflifeforms.Speciescanbephysicallyseparatedbya
geographicbarriersuchasanoceanoramountainrange,orvariouspreandpostzygotic
mechanismscanmaintainreproductiveisolationandpreventgeneflow.
b.Newspeciesarisefromreproductiveisolationovertime,whichcaninvolvescalesof
hundredsofthousandsorevenmillionsofyears,orspeciationcanoccurrapidlythrough
mechanismssuchaspolyploidyinplants.
1.C.3:Populationsoforganismscontinuetoevolve.
a.Scientificevidencesupportstheideathatevolutionhasoccurredinallspecies.
b.Scientificevidencesupportstheideathatevolutioncontinuestooccur.
1.D.1:ThereareseveralhypothesesaboutthenaturaloriginoflifeonEarth,eachwith
supportingscientificevidence.
a.Scientificevidencesupportsthevariousmodels.
1.PrimitiveEarthprovidedinorganicprecursorsfromwhichorganicmoleculescouldhave
beensynthesizedduetothepresenceofavailablefreeenergyandtheabsenceofasignificant
quantityofoxygen.
2.Inturn,thesemoleculesservedasmonomersorbuildingblocksfortheformationofmore
complexmolecules,includingaminoacidsandnucleotides.
3.Thejoiningofthesemonomersproducedpolymerswiththeabilitytoreplicate,storeand
transferinformation.
4.Thesecomplexreactionsetscouldhaveoccurredinsolution(organicsoupmodel)oras
reactionsonsolidreactivesurfaces.
5.TheRNAWorldhypothesisproposesthatRNAcouldhavebeentheearliestgenetic
material.
1.D.2:Scientificevidencefrommanydifferentdisciplinessupportsmodelsoftheoriginoflife.
a.GeologicalevidenceprovidessupportformodelsoftheoriginoflifeonEarth.
Evidenceofstudentlearningisademonstratedunderstandingofeachofthefollowing:
1.TheEarthformedapproximately4.6billionyearsago(bya),andtheenvironmentwastoo
hostileforlifeuntil3.9bya,whiletheearliestfossilevidenceforlifedatesto3.5bya.Taken
together,thisevidenceprovidesaplausiblerangeofdateswhentheoriginoflifecouldhave
occurred.
2.Chemicalexperimentshaveshownthatitispossibletoformcomplexorganicmolecules
frominorganicmoleculesintheabsenceoflife.
b.Molecularandgeneticevidencefromextantandextinctorganismsindicatesthatall
organismsonEarthshareacommonancestraloriginoflife.
1.Scientificevidenceincludesmolecularbuildingblocksthatarecommontoalllifeforms.
2.Scientificevidenceincludesacommongeneticcode.
2.A.2:Organismscaptureandstorefreeenergyforuseinbiologicalprocesses.
a.Autotrophscapturefreeenergyfromphysicalsourcesintheenvironment.
Evidenceofstudentlearningisademonstratedunderstandingofeachofthefollowing:
1.Photosyntheticorganismscapturefreeenergypresentinsunlight.
2.Chemosyntheticorganismscapturefreeenergyfromsmallinorganicmoleculespresentin
theirenvironment,andthisprocesscanoccurintheabsenceofoxygen.
b.Heterotrophscapturefreeenergypresentincarboncompoundsproducedbyother
organisms.
2.B.3:Eukaryoticcellsmaintaininternalmembranesthatpartitionthecellintospecialized
regions.
c.ArchaeaandBacteriagenerallylackinternalmembranesandorganellesandhaveacellwall.
2.D.2:Homeostaticmechanismsreflectbothcommonancestryanddivergencedueto
adaptationindifferentenvironments.
a.Continuityofhomeostaticmechanismsreflectscommonancestry,whilechangesmayoccur
inresponsetodifferentenvironmentalconditions.
C.Homeostaticcontrolsystemsinspeciesofmicrobes,plantsandanimalssupportcommon
ancestry.
4.C.1:Variationinmolecularunitsprovidescellswithawiderrangeoffunctions.
b.Multiplecopiesofallelesorgenes(geneduplication)mayprovide
newphenotypes.
1.Aheterozygotemaybeamoreadvantageousgenotypethan
ahomozygoteunderparticularconditions,sincewithtwodifferentalleles,theorganismhastwo
formsofproteinsthatmayprovidefunctionalresilienceinresponsetoenvironmentalstresses.
4.C.2:Environmentalfactorsinfluencetheexpressionofthegenotypeinanorganism.
B.Anorganismsadaptationtothelocalenvironmentreflectsaflexibleresponseofitsgenome.
4.C.3:Thelevelofvariationinapopulationaffectspopulationdynamics.
b.Geneticdiversityallowsindividualsinapopulationtoresponddifferentlytothesamechanges
inenvironmentalconditions.
Tofosterstudentunderstandingofthisconcept,instructorscanchooseanillustrativeexample
suchas:
Notallanimalsinapopulationstampede.
Notallindividualsinapopulationinadiseaseoutbreakareequallyaffectedsomemaynot
showsymptoms,somemayhavemildsymptoms,orsomemaybenaturallyimmuneand
resistanttothedisease.
c.AllelicvariationwithinapopulationcanbemodeledbytheHardyWeinbergequation(s).