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MORPHOLOGY OF
FLOWERING PLANTS
Concepts
PLANT MORPHOLOGY
MODIFICATION OF ROOTS
Modified Tap Roots
Floating root : Spongy aerial roots filled with air help the
plant in floating e.g., Jussiaea.
THE ROOT
Root thorn : Roots near the soil become hard like thorns.
These thorns serve as protective organ e.g., coconut.
Modified Adventitious Root
Modified for Physiological Functions
Storage root
Parasitic root : Roots arise from the stem and enter the host
plant developing contact with xylem and phloem of the host
stem e.g., Cuscuta.
Tuberous root : Arise from nodes of stem and one root out
of bunch become tuberous and fleshy for storage of food
e.g., sweet potato.
SHOOT SYSTEM
Aerial or Metamorphosed
LEAF
The leaf is a specialised organ of photosynthesis, transpiration
and gaseous exchange.
Parts of Leaf or Phyllopodium
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Phyllotaxy
The arrangement of mature leaves on the stem or its branches is called phyllotaxy.
It is of three types :
Modifications of Leaves
Leaf tendrils
Leaves or their
portions may
become modified
into tendrils in
various plants.
e.g. Pisum
sativum,
Gloriosa
Leaf spines
Leaves or their parts
are modified into
spines either to
escape transpiration
or for protection
e.g. Opuntia,
Barberry
Scale leaves
Store food and
water and are,
therefore, thick
and fleshy
e.g. Asparagus
Pitcher
Phyllode
The leaf-blade is
The leaf or
modified into a
lamina sheds
and the petiole pitcher-like structure.
modifies into The pitcher is provided
leaf like
with a lid which itself is
structure
an outgrowth of the
e.g. Australian
leaf-apex.
e.g. Nepenthes
acacia
Bladder
The leaf segments
get modified into
tiny bladders. The
bladder like
structure is meant
for catching the
aquatic insects
e.g. Utricularia
Phyllotaxy
Alternate :
Only one leaf is borne
at each node in alternate
manner e.g. Mustard,
Sunflower
Opposite :
two leaves occur
at each node.
INFLORESCENCE
I. Racemose (Indefinite)
(Main axis grows indefinitely bearing
flowers in Acropetal order)
RACEMOSE
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CYMOSE
Special cymes
Typical cymes
Hypanthodium : Receptacle is fleshy and cupshaped. Flowers are sessile and unisexual. Female
flower are found on the lower side and the male
flowers near the aperture e.g. Ficus spp.
Gamophyllous:
In such flowers tepals are fused.
(c) Polygamous : possess more than two types of flowers
e.g. Mango, cashewnut, trees possess three types of Calyx
flowersbisexual, staminate and neuter.
The outermost non-essential but protective whorl of the flower is
the calyx. It is represented by the sepals.
Arrangement of Floral Parts
Polysepalous : Sepals are free.
Hemicyclic: Floral organs are partly in spiral and partly in Corolla Aestivation
whorls, e.g., Anona.
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Androecium
Chambers of Ovary
Placentation
Gynoecium
CARPELS
Simple
e.g. : Leguminoseae,
poaceae
Apocarpous
e.g. : Rosaceae,
buttercup
Compound
Syncarpous
e.g. China rose,
potato, crucifers,
sunflower
Wall
FRUIT= RIPENED OVARY
Simple
Ovules
Aggregate
Etaerio of achenes : Each fruitlet is an achene. The
thalamus is fleshy and becomes red on maturation
and is the edible part in Strawberry, e.g. Clematis,
Ranunculus, Lotus
Etaerio of follicles : Many follicles (fruitlets) make
an etaerio which are arranged on the enlarged
thalamus in a bunch, e.g. Calotropis
Pericarp
Epicarp
Mesocarp
Endocarp
Seeds
Multiple or Composite
Syconus : It develops
from hypanthodium
infloresence.
Receptacle becomes
fleshy and many
achenes develop from
the pistillate flowers.
Develops from Hypanthodium, e.g. Banyan
Hesperidium : Develop
from multicarpellary,
syncarpous, superior
ovary having axile
placentation. Mesocarp is
in the form of white, fibrous
part fused with epicarp, e.g.
lemon, orange
Indehiscent
Achene : One seeded
developed from superior
ovary, fruit wall and seed
coat free, e.g., Hogweed
(Boerhaavia), Mirabilis
jalapa
Caryopsis : A superior
acheme, fruit wall and
seed coat fused e.g., rice,
wheat, maize
Cypsela : An inferior
acheme, e.g., sunflower,
marigold
Nut : A large woody
pericarp acheme, e.g.,
water chestnut, oak,
cashew-nut, litchi
Dehiscent
Follicle : Dehiscing by
ventral suture, e.g.,
Caloptropis.
Legume or Pod :
Dehiscing by both the
sutures. e.g., pea.
Siliqua : Dehiscing from
base to apex, e.g. mustard
to expose replum with
attached
seeds
(Brassica)
Capsule : Dehiscing in
many ways e.g., lady
fiinger.
Schizocarpic
Dehiscing into one seeded
parts :
Lomentum : A legnum
breaking into one seeded
parts e.g., Acacia,
Mimosa.
Cremocarp : Capsule
breaking into two
mericarp e.g. Coriander,
Fennel.
Regma : Single seeded
fruitlets dehisce to release
out its seeds called cocci
e.g. Castor, Geranium
Double samra : Breaking
into two winged fruitlets
e.g. Acer (Maple)
The endosperm has one to three layered peripheral proteins Floral Characters
layer called aleurone layers which separates the embryo Inflorescence : Racemose
with endosperm. The cotyledon is also called scutellum in Flower : Bisexual, zygomorphic
cereals. It is laterally attached to the embryonal axis. The
Calyx : Sepals five, gamosepalous; imbricate aestivation.
lower end of the axis is called the radicle which has a
protective sheath termed coleorhiza. The upper end of the Corolla : Petals five, polypetalous, papilionaceous, consisting of
a posterior standard, two lateral wings, two anterior ones forming
axis is called the plumule which is covered by coleoptile.
There are two types of seeds i.e., albuminous and exalbuminous. a keel (enclosing stamens and pistil), vexillary aestivation.
Albuminous (endospermic) seeds : In these seeds food is Androecium : Ten, diadelphous, anther dithecous.
stored in the endosperm, e.g., corn, wheat, caster, onions, Gynoecium : Ovary superior, mono carpellary, unilocular with
many ovules, style single.
etc.
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Fruit : Legume; seed: one to many, non-endospermic
G
Floral formula : % K(5) C 1 + 2 + (2) A
(9) +1 1
Floral Diagram of Pisum sativum :
Economic Importance
SOLANACEAE
LILIACEAE
It is commonly called as Lily family.
HabitUsually perennial herb as Polygonatum, sometimes small
shrubs as Ruscus.
RootAdventitious, fibrous.
StemErect or climbing as Smilax, branched or unbranched,
herbaceous.
Leaf Exstipulate, alternate or whorled, parallel venation.
InflorescenceRaceme, umbel or sometimes solitary.
FlowerPedicellate, bracteate, hermaphrodite, actinomorphic,
trimerous, hypogynous.
Perianth Tepal six (3 + 3) perianth in two whorls, imbricate
aestivation,. Androecium six (3 + 3) stamens in two whorls.
GynoeciumTricarpellary, syncarpous, superior ovary, axile
placentation, trilocular, two or more ovules in each locule.
FruitCapsule or rarely berry.
Floral formula Br.
P3 + 3 A 3 + 3 G(3)
Economic Importance
Allium cepa, A. sativum and Asparagus racemosus are used as
vegetables.
Leaves of Allium cepa (onion) are used in dental and ear troubles.
The leaves of Allium sativum (lahasun) are used in cough and
fever.
Colchicine, an alkaloid obtained from Colchicum autumnale is
used to induce polyploidy in plants.
Exercise
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(a) A - Hilum, B - Micropyle, C - Radicle, D - Cotyledon,
E - Plumule
(b) A - Hilum, B - Micropyle, C - Plumule, D - Cotyledon,
E - Radicle
(c) A - Micropyle, B - Hilum, C - Plumule, D - Cotyledon,
E - Radicle
(d) A - Hilum, B - Micropyle, C - Plumule, D - Radicle,
E - Cotyledon
56.
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(a)
(b)
94.
(c)
(d) Br % K(5) C(5) A(10) G1
Which one of the following is a true fruit?
(a) Apple
(b) Coconut
(c) Plum
(d) Mango
Trilocular ovary with axile placentation is found in :
(a) Liliaceae
(b) Gramineae
(c) Compositae
(d) Cruciferae
Cross pollination is a rule in :
(a) Cleistogamy
(b) Chasmogamy
(c) Dicliny
(d) Bisexuality
95.
(b) Pear
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(c) When the male and female reproductive organs develop 56. (b) The edible part of cauliflower represents a fleshy
on different individual plants the condition is called
inflorescence. Here the upper part of the peduncle is
dioecious. In such plant species, fertilization can not
suppressed, so that all the flowers appear to lie in one
occur in a single isolated plant.
plane. The lower and older flowers are with long
(a)
pedicels, and the upper and younger flowers are with
(a) A flower with all the petals of the same size produces a
shorter pedicels. This is a type of corymb inflorescence.
circular out line, thereby rendering an actinomorphic 57. (c) In cyathium inflorescence, extreme case of reduction is
symmetry. Corolla with one petal larger or smaller than
noticed. Here the peduncle is reduced to a small disc
the rests provides zygomorphic symmetry and corolla
which bears single female flower and many male
with petals of different sizes forms asymmetric condition.
flowers. The flowers are achlamydous , i.e., without
(a)
any accessory whorls. The whole inflorescence is
(d) In an androecium, when the stamens are united only
covered by a large bract called involucre, thus
by anthers the condition is called syngenesious (a case
producing a single flower-like appearance.
of cohesion). When the stamens (filaments) are united 58. (b) Raceme is a typical racemose inflorescence and is
to petals, it is called epipetalous (a case of adhesion),
characterised by indetermiante growth of peduncle and
Syngenesious-epipetalous stamens are characteristic
acropetal succession of flowers, i.e., older flower at the
of Compositae (Asteraceae).
base and younger at th apex.
(b) Both bract and bracteole are leafy structure associated 59. (d)
with flowers. Bract develops one peduncle 60. (d) Spadix is a type of spike, covered by large bract/bracts.
(inflorescence axis) and bracteole develops on pedicle
The female inflorescence of Maize is covered by many
or at the base of the thalamus.
bracts, hence represents a spadix.
(a) The mustard (Brassica campestris) belongs to 61. (d)
62. (c)
63. (b)
64. (b)
Cruciferae (Brassicaeae). The stamens of this family is 65. (b) An aggregate fruit is an association of simple fruits,
characteristically tetradynamous (two small and four
called etaerio. An aggregate fruit can develop only from
large free stamens).
apocarpous pistil (free multiple carpels).
(d) Glumes are characteristically present in gramineae. They 66. (b) A simple fruit develops from single ovary, irrespective
are scaly (leafy) structures and develop on peduncle.
of the number of carpel.
(b)
67. (a) In dehiscent fruits, there are one or more longitudinal
(c) In protogyny, the gynoecium matures before
lines, along which the mature fruit opens, called suture.
androecium in a bisexual flower. Such condition is found
The number of suture is one of the important features
in Magnolia, Michelia (Magnoliaceae), Anona,
in identifying the type of dehiscent fruits. Follicle differs
Polyalthia (Anonaceae), Ficus (Moraceae) etc.
from all other dehiscent fruits in having a single suture.
(b)
68. (b) In samara, the pericarp is modified to wing. In Shorea,
(b) When the stamens (by their filaments) are united to
the calyx is persistent and modified to wing.
petals the stamens are called epipetalous. Such 69. (a) Cabbage is a condensed shoot i.e., a vegetative bud
condition is found in Solanaceae, Malvaceae,
(terminal).
Compositae, etc.
70. (c) The mesocarp of coconut fruit is represented by fibres,
(d) In floral formula, sexuality, symmetry, number of floral
which provides the raw material for coir of commerce.
parts and their cohesion and adhesion, and the position 71. (c)
of ovary can be depicted. The characters like 72. (b) Lomentum is a schizocarpic fruit.
placentation, aestivation and nature of thalamus can 73. (a) Fleshy fruits are always indehiscent. The seeds are
not be given.
exposed only after the decay of the fleshy part (usually
(c) Sometimes zygomorphy may result from oblique
pericarp).
position of ovary as in the case of Solanaceae.
74. (c) Geitonogamy is found in monoecious plants like maize.
(c) Didynamous stamen is found in Ocimum (Labiatae).
This type of pollination may also occur in
(a) In papilionaceous corolla (sweet pea), the two lower
hermaphrodite plant species.
petals (keel remain united and cover the stigma surface. 75. (b) Papilionaceous corolla is characteristic to papilionaceae
As a result cross pollination can not takes place.
(Fabaceae). Bean is an example of this family.
(c)
76. (a) In geitonogamy, pollination occur in the same plant,
(c) In capitulum inflorescence, the flowers (florets) are
resulting no new recombinations.
sessile and arranged in centripetal succession 77. (a) In Mimosaceae and Typhaceae, the pollen grains are
(originally acropetal). It is , therefore, a modification of
not set free after meiosis but are held together in the
spike.
form of tetrad or polyad.
(a) In onion (Allium cepa) and garlic (A. sativum) the 78. (b) The pollinia are connected with each other by slender
inflorescence appears like umbel but the arrangmeent
stalk-like process forming translator apparatus.
of flowers is more or less centrifugal i.e., cymose like. 79. (b) Pollen grains or microspores are halpoid structures and
This is a special type of inflorescence called
are produced from pollen or microspore mother cell by
scapigerous cyme or unbellate cyme.
meiosis. The process is called microsporogenesis.
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15
80. (d) Pollen grains of certain plants cause allergic reaction
to human beings. The allergic response to pollen is
called hayfever or pollinosis. Protein content of the
pollen (mainly g -globulin) affects the mucous
membrane of the upper respiratory tract.
81. (a) In majority (70%) of angiosperms, the embryo sac or
female gametophyte is 8-nucleate (called Polygonum
type), of which the two nuclei are united to form as
diploid (2n) secondary nucleus. So the mature female
gametophyte is 7-celled (1 egg cell, 2 synergids, 1
secondary nucleus and 3 antipodal cells).
82. (c) The pollen grains is haploid and represents the first cell
of male gametophyte.
83. (d)
84. (d)
85. (d) The male gametophyte of angiosperms initiatesfrom
the formation of pollen grain and ends with the
production of micro-sporangium of anther (i.e., on male
sporophyte) and the later part of development takes
place at stigmatic surface and style (i.e., on the female
sporophyte).
86. (b)
87. (c) In certain seeds, like castor, an outgrowth develops
from the integument (testa) and covers the micropyle,
called caruncle.