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8.

9.

// . : . -1998. - . 113-118.

// - .: - .. , 1978. 87.

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- : - . - 392.

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, [1-4].
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: .
[5] , , , [4]

dX
L
=
X + k4SX + k2G k1G m ,
dt
KL + L

d
dt

m
Z Z + k4SZ + k1G k2G ,
KL +L
dL
1

( + )L ,
=
dt
Y KL + L

68

(1)

(2)

(3)

.., .., ..

dS
= ( k3 k4S )( + ) ,
dt

(4)

, /

t,

, /

X , Z , S , L - , , , , ; - ; K L - , k1 - , k2 - , ; G - ; - ; k3 k4 - , ,
m - , Y - .

(1)-(4)

2
-1
2
= 1,75 10 , K L = 7,5 10 , Y = 0,85 , = 1,55 103 -1
[1] , ,
. 1. , , X ,

a = 0,008 0,02 / L = 0,15 (. 1).

L
)

t,

. 1 L X : ) 0,008 /; )
6 /

, - a = 6 .

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69

. 10, 2009

, 20 (.1 ), .
[6], . (1)(4),
,
2 . ,

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. ,
. , , q = 25,2 3 ,
a = 6 ,
Li = 0,015 . , Zi 0 . q = 45,13 3 L = 0,15 .
1, 2 3
4
-1
3
-1
G = G1 2 = 1,6 10 G = G3 = 1,6 10 [1].

m = 2 , k1 = 5 1010 , k2 = 2,4 105 , [1],


. 2.
. 2 , ( ) ,
.
= 3,6 ,
= 0 , L 0 , X
.
(.2 ), ,
= 0,24 . (. 2 ) : = 0,08 , = 0,25 , L 0 .
a = X + Z = 0,33
70

.., .., ..

a = 3 4 .

, , 25%.
L 0,015 .
.

Z
L

X+Z
, /

, /

X+Z

L
X
)

, /

, /

X+Z

X+Z

L
)

. 2 : ) ; ), )
: 1- 4- , )

X L X Z (1), (2) (3), G1,2


G3 , [1] -

. , (4)

71

. 10, 2009

a L
. ,
, , , . ,
, [7]

dX
= ( ax + bx L ) + k2G k1G m + k4SX
dt

(5)

d
= ( az + bz L )Z k2G + k1G m X + k4SZ
dt

(6)

dL
= ( g x + g z )L
dt

(7)

dS
= ( k3 k4S )( + ) ,
dt

(8)

aX , aZ - ;
bx , bz - , , g x , g z -
.
(5) (6) [7] , . (5)
, , .
(6) ,
[1]. (7) , . (8) (4).

72

.., .., ..

(5)-(8) S
, (1)-(4), , . 3.
X+Z

, /

X+Z

, /

X+Z

Z
X

X+Z

, /

, /

Z
L

L
)

. 3 : ) ; ), )
: 1- 4- , )

:
aX = 1,2 103 -1 ,
bX = 7 102
( ) ,
aZ = 3 103 -1 ,

4
bZ = 3,7 10 1 ( ) , g X = g Z = 2,5 103 ( ) , G1 2 = 1,6 10

-1

3
-1
G3 = 2 10 , k1 = 5 1010 , k2 = 2,4 105 . -

, ax , az , bx , g x , g z
X , [1].
= 2,85 ,
0 , L 0,015 . bZ 5
bx , 2,6 ,

73

. 10, 2009

g x g z . , [1, 8].
, . 2 3,
,
,
. , (. 3 -)
(5)-(8) , , . (1) (4)
.
(5) (8)
(. 3 ), .
(1) (4) (5) (8) , , ,
, . , .

, /

L()

X()

L()

X ()
,

. 4 : ) (1) (4); ) (5) (8)

, (5) (8) aX = 1,55 10 3 -1 , bX = 9,9 10 2 ( ) ,


aZ = 1,2 10 3 -1 ,
bZ = 1,1 102 ( ) ,
g X = g Z = 1,08 10 1
( ) . ,
74

.., .., ..

(1) (4) (5) (8) X L


k1 = k2 = 0 ( ) . 4.
, (5)-(8)
, . , (5)-(8) ,
. 3,
(1)-(4) : = 3,5 101 -1 , K L = 4,77 ,
Y = 27 , = 1,14 103 -1 . (1)-(4)
, , . 5.

Z
L
)

X+Z
Z

, /

, /

)
X+Z

X+Z
, /

, /

X+Z

Z
X
L

L
)

. 5 : ) (); ), ) : 1- 4- ; )

,

. 3 , . 5 , .

75

. 10, 2009

, .
(1)-(4) . ,
. 2 ,
.
= 0,6 , = 0,6 ,
L = 0,08 . : = 3 4 , 0 , L 0,015 .
. X L ,
(1) (4) (5) (8)
( ),
. , , . , ,

(1)-(4), . , (5)-(8)
.
,
.
, .

1.
2.

76

.., ..
.
.: , 1979. 119 .
Monod J. La technigue de culture continue theorie and applications. Ann. Institut Pasteur, 1950. 79 c.
.., .., ..

3.
4.

5.

6.

7.
8.

Herbert D. A theoretical analysis of continuous culture systems.


In: Symp. Of continuous of microorganisms, 1960. L., 1961. Sc. I,
monogr. N 12, p. 44-52.
.., .. --
... ., . .
: - . 4, 2005. . 46-53.
.., .. , . . . . . . . .7.- :
, 2008. .51-57.
.., .. , , . . - . . :
, .81. : , 2008. . 133-139.
. . .: , 1976. - 288 .
.. . . ,
2003. 512 .

77