Professional Documents
Culture Documents
Learning
Objec9ves
A"er
studying
this
chapter
you
should
be
able
to
1. Explain
the
dierences
between
gap-phase
disturbances
and
stand-replacing
ones.
2. Describe
how
disturbance
is
followed
by
a
successional
recovery,
but
not
necessarily
back
to
the
original
kind
of
ecosystem.
3. Explain
the
broad
paIerns
and
mechanisms
of
successional
recoveries
aJer
disturbance.
4. Discuss
the
key
dierences
between
primary
and
secondary
succession.
Learning
Objec9ves
5. Explain
how
the
paIerns
and
processes
of
natural
disturbances
can
be
emulated
to
soJen
the
environmental
impacts
of
anthropogenic
harves9ng
and
management
systems.
Disturbance
Destruc9on
of
part
of
a
community.
Community-level
recovery:
succession.
Wide
range
of
spa5al
scales.
Wide
range
of
temporal
scales.
Smaller-scale
Disturbances
Gap-phase
disturbances
occur
at
a
small
scale
in
an
otherwise
intact
community.
Gaps
caused
by:
Death
of
a
single
tree
in
a
forest.
Small
landslides.
Diggings
by
animals
such
as
prairie
dogs.
Microsuccession
restores
the
community
in
the
gap
created
by
a
fallen
tree.
(Shown
here:
Hardwood
forest
in
southern
Ontario.)
Large-scale
Disturbances
Stand-replacing
disturbances
aect
the
en9re
community.
Disturbance
caused
by:
Wildre.
Windstorms.
Pests
and
diseases.
Glacia9on.
In
the
prairies
and
boreal
forest,
res
belong
to
natural
ecosystem
dynamics.
So, beneath the Aspen canopy there is young Spruce and Fir (not Aspen).
Primary
Succession
Succession
in
an
ini5ally
abio5c
habitat
Volcanic
erup9ons.
Glacial
retreat.
Landslides.
Sand
dune
succession.
Primary
succession
on
a
scree
slope
in
the
Swiss
Alps
with
Campanula
cenisia.
Secondary
Succession
Some
life
survives
the
disturbance,
such
as
seed
bank,
below- ground
organisms,
soil
microbes.
Examples:
Fire.
Abandoned
elds.
Storm
damage.
PaIerns
of
Succession
No
intervening
disturbance.
A
sequence
of
community
types
(seral
stages).
The
seral
stages
together
form
a
sere.
A
stable
community
develops,
depending
on
enduring
site
features,
species
present
and
stochas9c
factors.
Sedge
meadow
is
a
seral
stage
in
succession
following
a
collapse
of
a
beaver
dam
in
Killarney
Provincial
Park,
Ontario.
Temperate Forest
Ini9al
seral
stages
dominated
by
ruderal
plants
and
survivors.
Pioneer
trees
are
fast-growing
and
shade-intolerant,
with
good
dispersal
ability.
Shade-tolerant
trees
increasingly
dominate
over
9me.
Fig.
10.2.,
p.
300
Hydrosere
Begins
with
a
young
lake
or
pond;
e.g.,
aJer
glacial
mel9ng.
Ini9ally
oligotrophic,
sediment
accumula9on
slow.
It
may
become
more
nutrient-rich
over
9me.
Eventually,
the
en9re
lake/pond
may
turn
to
a
wetland.
Fig.
10.3.,
p.
301
Facilita9on
Model
Earlier
seral
stages
are
required
for
later
stages
to
occur.
E.g.,
lichens
and
mosses
create
a
substrate
for
grasses
and
herbs,
which
create
substrate
for
trees.
Leave
nutrients
and
shade.
Tolerance
Model
Species
vary
in
their
ability
to
u9lize
resources
and
tolerate
certain
condi9ons.
E.g.,
colonizing
species
are
displaced
at
later
seral
stages
as
they
cannot
tolerate
the
shady
condi9ons.
Inhibi9on
Model
Early
successional
species
prevent
or
delay
the
establishment
of
other
species.
E.g.:
Later
species
only
can
establish
when
some
of
the
earlier
ones
die
(herbivore,
disease,
microdisturbance),
crea9ng
an
opportunity.
Successional
Mechanisms
in
Rocky
Inter9dal
Zone
Sousas
study
of
inter9dal
zone
succession
tested
the
inhibi9on
model:
If
the
inhibi9on
model
is
occurring,
early
successional
species
should
be
inhibi9ng
the
establishment
of
later
successional
species.
Facilita9on
Example
Turner
(1983)
Algae
and
surfgrass
coloniza9on
on
rocks
Surfgrass
aIach
seed
pods
to
algae
Tested
if
this
was
facilita9on
Frequency
of
Disturbance
Return
frequency
of
a
disturbance
can
be
regular
or
unpredictable.
Tides,
cold
or
dry
seasons,
and
spring
oods
are
regular
and
predictable.
Wildres
can
be
fairly
regular
but
not
exactly
Winter
is
predictable.
Organisms
can
adapt
to
it predictable.
e.g.,
plants
seed
dormancy
Volcanic
erup9ons
are
irregular.
Connell (1975)
Mean
number
of
plants
per
plot
in
sand-dunes
is
the
highest
at
intermediate
levels
of
rabbit
grazing.
Fig.
9.24,
p.
268
richness. richness.
Species richness
Disturbance
and
Species
Richness
in
the
Inter9dal
Zone
Sousa
(1979)
studied
eects
of
disturbance
on
species
richness
of
algae
and
invertebrates
growing
on
boulders
in
the
inter9dal
zone.
Predicted
level
of
disturbance
depends
on
boulder
size.
Large
boulders
require
more
force
to
move.
1 4 2
Timber
Harves9ng
3
years
aJer
clearcut
Logging
Prac9ces
Prevalent
even
in
highly
fragmented
landscapes.
Edge
crea9on
and
loss
of
interior
habitat.
Many
small
woodlots
are
at
risk
of
degrada9on
due
to
loss
of
interior
habitat.
Private
Public
Silvicultural
Systems
Even-aged
Systems
Clearcut
Systems
Uneven-aged
Systems
Selec9on
Systems
Natural
disturbance
Non-uniform
size
of
patches
Less
frequent
disturbance
Leaves
woody
debris
Nutrients
retained
Clear-cupng
Uniform
size
of
patches
More
frequent
harvest
Leaves
liIle
woody
debris
Nutrients
removed