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Journal of Memory and Language 51 (2004) 231246

Journal of Memory and Language


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Can CANISO activate CASINO? Transposed-letter similarity eects with nonadjacent letter positionsq
Manuel Pereaa,* and Stephen J. Lupkerb
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Departament de Metodologia, Universitat de Valncia, Facultat de Psicologia, Av. Blasco Ib~ez, 21, 46010 Valncia, Spain e an e b Department of Psychology, University of Western Ontario, London, Ont., Canada N6A 5C2 Received 6 February 2004; revision received 14 May 2004 Available online 17 June 2004

Abstract Nonwords created by transposing two adjacent letters (i.e., transposed-letter (TL) nonwords like jugde) are very eective at activating the lexical representation of their base words. This fact poses problems for most computational models of word recognition (e.g., the interactive-activation model and its extensions), which assume that exact letter positions are rapidly coded during the word recognition process. To examine the scope of TL similarity eects further, we asked whether TL similarity eects occur for nonwords created by exchanging two nonadjacent letters (e.g., canisoCASINO) in three masked form priming experiments using the lexical decision task. The two nonadjacent transposed letters were consonants in Experiment 1 (e.g., caniso-CASINO), vowels in Experiment 2 (anamil-ANIMAL) and both consonants and vowels in Experiment 3. Results showed that nonadjacent TL primes produce priming eects (in comparison to orthographic controls, e.g., caviro-CASINO), however, only when the transposed letters are consonants. In a nal experiment we examined latencies for nonwords created by nonadjacent transpositions of consonants versus vowels in a lexical decision task. Both types of nonwords produced longer latencies than matched controls, with consonant TL nonwords being more dicult than vowel TL nonwords. The implications of these ndings for models having position-specic coding schemes as well as for models proposing alternative coding schemes are discussed. 2004 Elsevier Inc. All rights reserved.
Keywords: Coding schemes; Lexical decision; Orthographic similarity; Transposed letters; Vowel/consonant processing

Introduction One key issue for models of visual word recognition is how the ordering of letters in a word is encoded
This research was supported by Natural Sciences and Engineering Research Council (NSERC) of Canada Grant A6333 to Stephen J. Lupker and a grant from the Spanish Ministry of Science and Technology (BSO2002-03286) to Manuel Perea. We thank Ken Forster, Carol Whitney, Iris Berent, and three anonymous reviewers for helpful criticism on an earlier draft. We also thank Edurne Goikoetxea for selecting the stimuli and conducting Experiments 3 and 4. * Corresponding author. Fax: +1-34-96-3864697. E-mail addresses: mperea@uv.es (M. Perea), lupker@uwo.ca (S.J. Lupker).
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within that words orthographic representation. Most current computational models of word recognition simply assume that the positions of the letters are established very early in processing, well before the identities of the letters are known (position-specic coding schemes; e.g., the interactive-activation (IA) model, Rumelhart & McClelland, 1982, and the models deriving from its architecture, the dual-route cascaded model, Coltheart, Rastle, Perry, Ziegler, & Langdon, 2001; and the multiple read-out model, Grainger & Jacobs, 1996). Thus, in these models, a nonword created by transposing two adjacent letters (e.g., JUGDE) would be no more similar to its base word (JUDGE) than a nonword created by simply replacing those letters (JUNPE).

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This type of coding scheme does not, however, appear to t with the available data. For example, a number of experiments have shown that people actually have more diculty accurately perceiving letter order information than letter identity information when a random sequence of letters is briey presented (e.g., Mewhort, Campbell, Marchetti, & Campbell, 1981; Ratcli, 1981). Results have also shown that adjacent transposed-letter (TL) nonwords (e.g., JUGDE) have a strong tendency to be misperceived as words in a lexical decision task, a tendency that is even stronger than that for replacementletter nonwords (JUNGE) (see Chambers, 1979; OConnor & Forster, 1981). Further, in masked priming experiments (Forster & Davis, 1984), TL nonword primes not only produce form-priming eects relative to an orthographic control (e.g., jugde-JUDGE vs. jupte-JUDGE; Perea & Lupker, 2003b; see also Andrews, 1996; Forster, Davis, Schoknecht, & Carter, 1987; Peressotti & Grainger, 1999; Schoonbaert & Grainger, 2004), but also associative-priming eects (e.g., jugde-COURT vs. ocaenCOURT; Perea & Lupker, 2003a). The presence of these TL similarity eects poses a challenge for word recognition models that use a position-specic coding scheme, that is, those models in which letters are assumed to be immediately assigned to their correct positions in the letter string. At the very least, addressing this challenge would require the models to drop this immediate assignment assumption and, instead, assume that letter positions often take more time to encode than letter identities. In addition, one would also need to assume that a letter in position N produces some activation of the representation of that same letter at positions N 1 and N 1 (Rumelhart & McClelland, 1982; see also Andrews, 1996; Peressotti & Grainger, 1995). Indeed, Rumelhart and McClelland (1982) acknowledged that there might be a problem with the coding scheme in their model and suggested that:
information presented in one location might activate detectors in a range of locations rather than simply in one xed position. Perhaps there is a region of uncertainty associated with each feature and with each letter. If so a given feature in a given input position would tend to activate units for that feature in positions surrounding the actual appropriate position. As a result, partial activation of letters from nearby positions would arise in a particular position along with the activation for the letter actually presented. (p. 89)

entry corresponding to its base word (JUDGE) somewhat more than the two-dierent letter nonword JUNPE, allowing the models to account for these types of eects. Simulation work would, of course, be necessary in order to verify that integrating these ideas into the letter coding schemes in the current models would allow those models to: (a) successfully capture TL similarity eects, and (b) maintain their ability to account for the other eects that they currently are able to account for (i.e., consider the discussion below concerning Daviss, 1999, analysis). In principle, however, the presence of TL similarity eects based on adjacent letters may not present an insurmountable challenge for a modied IA model. In the present paper, we wished to examine the scope of TL similarity eects. Specically, using the masked priming technique (Experiments 13), we asked whether TL similarity eects occur for TL nonwords created by transposing two nonadjacent letters (e.g., caniso-CASINO). In all cases, these eects were evaluated relative to the appropriate orthographic controls (i.e., replacement-letter nonwords as primes, as in caviro-CASINO). In an eort to obtain additional evidence on this issue, we also asked whether the TL nonwords created by transposing two nonadjacent letters are more competitive (in terms of the number of false positives and longer latencies) than their corresponding orthographic controls in a lexical decision task (Experiment 4). The presence of TL priming eects when the transposed letters are not adjacent would pose a substantially greater problem for word recognition models using position-specic coding schemes. That is, these eects would require the assumption that a letter in position N activates its representation across letter positions N 2 to N 2. As described by Davis (1999), incorporating this assumption would seriously harm these models ability to recognize highly familiar inputs. The reason is that, in the IA architecture, the mechanism responsible for resolving competition between candidate letters in a given letter position is bottom-up inhibition between the feature and letter levels. If a given letter position were receiving activation from features from up to ve different letters (i.e., letters in positions N 2 to N 2) it would be almost impossible for the inhibitory process to function eectively. Thus, the existence of nonadjacent TL priming eects would strongly suggest that IA-based models would be best served by incorporating a dierent type of coding scheme. The search for a new coding scheme The existence of TL similarity eects, even when the transposed letters are not adjacent, is, in fact, a natural consequence of the letter coding schemes in two recently proposed computational models of the letter coding process: the SOLAR model (Davis, 1999) and the

Similarly, Peressotti and Grainger (1995) indicated that there could be some form of cross-talk between neighboring letter positions (p. 886), and Andrews (1996) indicated that a letter in position n yields some activation of the same letter in positions n 1 and n 1 (p. 797). With this reformulated coding scheme, the TL nonword JUGDE would be expected to activate the lexical

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SERIOL model (Whitney, 2001).1 The SOLAR model uses a spatial coding scheme in which letter codes are position-independent, so that the nonadjacent TL nonword CANISO and its base word, CASINO, share the same set of letter nodes. The order of the letters is coded by the relative activity of the set of letter nodes. Thus, CANISO and CASINO would be coded dierently because they would produce dierent activation patterns across the letter nodes they share (e.g., in the word CASINO, the letter node corresponding to C is the one associated with the highest activation value, the letter node corresponding to the letter A is associated with a slightly smaller activation value, etc.). The SERIOL model (Whitney, 2001) uses a letter-tagging coding scheme, in which each letter is marked for the ordinal position in which it occurs within a letter string. For instance, the word CASINO would be represented by C1, A-2, S-3, I-4, N-5, and O-6 with the relevant letter nodes then receiving dierential levels of activation as a function of position. This letter-tagging scheme is accompanied by the activation of bigram nodesordered pairs of lettersso that CASINO would be represented by the following bigram nodes: CA, AS, SI, IN, NO, CS, CI, CN, CO, AS, AI, AN, AO, SI, SN, SO, IN, and IO. The nonadjacent TL nonword CANISO would then share 13 bigram nodes with CASINO (CA, AS, NO, CS, CI, CN, CO, AS, AI, AN, AO, SO, and IO), whereas the two-letter dierent nonword CAVIRO would share only six bigram nodes with CASINO (CA, CI, CO, AI, AO, and IO). For our purposes, the crucial point here is that, according to both the SOLAR and SERIOL models, nonwords created by transposing nonadjacent letters are highly similar to their base words. Further, although the precise similarity of CASINO and its nonadjacent TL
1 Other coding schemes had been proposed in the literature, the more cited of which has been Mozers (1987) BLIRNET model. Mozer (1987) used letter-cluster units that respond to local arrangements of letters in which the only location information retained consisted of the relative positions of letters within a cluster. The letter-cluster units respond to letter triples: either a sequence of three adjacent letters (e.g., CAS in the Spanish word CASINO), or two adjacent letters and one nearby letter, such as CA_I or S_NO, where the line indicates that any letter may appear in the corresponding position. For instance, presentation of CASINO should result in the activation of the following letter-cluster units: **C, **_A, *CA, *_AS, *C_S, CAS, C_SI, CA_I, ASI, A_IN, AS_N, SIN, S_NO, SI_O, INO, I_O*, IN_*, NO*, N_*, and O**. (The asterisk signies a blank space and double-asterisks are used simply to keep all units in the form xxx, xx_x, or x_xx.) The word CASINO and the nonadjacent TL pseudoword CANISO only share 5 (out of 20) letter-cluster units. The word CASINO and the orthographic control CAVIRO also share ve letter-cluster units. Thus, BLIRNET would predict that nonadjacent TL nonwords are no more similar to their base words than orthographic controls.

nonword CANISO depends on a variety of factors, the current parameter sets in the SOLAR and SERIOL models predict that a nonadjacent TL nonword like CANISO is more similar to CASINO than a two-letter dierent nonword like CAVIRO. More specically, in terms of calculated similarity, the similarity match between CASINO and CASINO would be 1.00 in both models. For the SOLAR and SERIOL models, respectively, the similarity match to CASINO would be reduced to .83 or .88 for the adjacent TL neighbor CAISNO, to .75 or .83 for a one-letter dierent nonword like CASIRO, to .62 or .71 for the nonadjacent TL nonword CANISO, and to .54 or .49 for a two-letter dierent nonword like CAVIRO (an unrelated nonword like NOMERA results in a match value of .13 in the SOLAR model and a match value of .20 in the SERIOL model).2 Thus, although the presence of TL similarity eects based on nonadjacent letter positions would pose considerable problems for position-specic coding schemes, their presence would actually support the predictions of these two recent models of letter coding. In Experiments 13 we asked whether TL nonword primes created by transposing two nonadjacent letters do produce reliable form-priming eects relative to twoletter dierent nonwords (e.g., caniso-CASINO vs. caviro-CASINO). Because TL similarity eects are greater for nonwords created by transposing internal rather than external letters (Chambers, 1979; Perea & Lupker, 2003a, 2003b), the transposed letters were always the third and the fth in six-letter words (or nonwords) in Experiments 1 and 2. (They were the third and fth, or the fourth and the sixth in Experiment 3, in which the items were 710 letters long; e.g., tradegia-TRAGEDIA.) For comparison purposes, we also included a one-letter dierent condition (casiro-CASINO), as well as either an identity condition (casino-CASINO; Experiment 1a), or an unrelated condition (nomero-CASINO, Experiments 1b and 2). The two nonadjacent transposed letters were consonants in Experiments 1a and 1b (e.g., caniso-CASINO vs. caviro-CASINO) and vowels in Experiment 2 (anamil-ANIMAL vs. anomel-ANIMAL). Experiment 3 was an attempt to replicate the results of Experiments 1 and 2 using a new set of items. To obtain converging evidence of TL similarity eects using another experimental technique, Experiment 4 employed a single-presentation lexical decision task in which the nonword targets were the masked primes used in Experiment 3. The dierential processing of vowels and consonants The second issue being addressed in the present research concerns potential dierences in the processing of vowels versus consonants. Recent research strongly
2 We thank Colin Davis and Carol Whitney for providing us with the match scores.

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suggests that vowels and consonants are processed differently (visual-word perception: Berent & Perfetti, 1995; Berent, Bouissa, & Tuller, 2001; Lee, Rayner, & Pollatsek, 2001, 2002; speech perception: Boatman, Hall, Goldstein, Lesser, & Gordon, 1997; neuropsychology: Caramazza, Chialant, Capasso, & Miceli, 2000; Caramazza & Miceli, 1990; Cubelli, 1991; Ferreres, Lpez, Petracci, & China, 2000; cognitive modeling, o Monaghan & Shillcock, 2003; language acquisition, Nespor, Pe~a, & Mehler, 2003; linguistics, Gafos, 1998; n Goldsmith, 1990). The experimental tasks used in this research varied widely, from perceptually based tasks to production tasks (see Berent et al., 2001; Monaghan & Shillcock, 2003). The clear implication, as a number of these researchers have suggested, is that the consonant/ vowel distinction is an essential one in that orthographic representations convey information concerning not only letter identity but also consonant/vowel status (e.g., Berent et al., 2001; Caramazza & Miceli, 1990; Tainturier & Caramazza, 1996). Note, as well, that the dimension of consonant/vowel status also emerges as important in patterns of brain damage. That is, there are documented cases of brain damaged patients with a selective decit with vowels (e.g., Caramazza et al., 2000; Cubelli, 1991) or with consonants (e.g., Caramazza et al., 2000; Kay & Hanley, 1994). Thus, it is even possible that consonants and vowels are processed by dierent neural mechanisms (Caramazza et al., 2000). Finally, it is worth noting two additional facts particularly relevant to the present research. First, in normal speech in Spanish (the language used in the present experiments), individuals make substantially more pronunciation errors by transposing two consonants (14.2% of errors; e.g., the nonword escanLaDosa instead of the Spanish word escanDaLosa) than by transposing two vowels (1.9%; see Prez, Palma, & Santiago, 2001). e Second, a post hoc analysis of Experiment 3 of Perea and Lupker (2003b) revealed that transposition of two adjacent internal consonants (e.g., mohter-MOTHER) led to as much priming as that from identity primes, whereas the transposition of two adjacent internal vowels (freind-FRIEND) produced very little priming. Unfortunately, this comparison is not only post hoc, it also fails to control for any dierences between withinand between-syllable transpositions. In the present experiments, due to the syllable structure of Spanish, all transpositions were between-syllable transpositions. How the consonant/vowel distinction actually manifests itself during processing has been a matter of considerable theorizing in recent years. For instance, Berent et al. (2001) have proposed that printed words are represented in the internal lexicon in terms of a consonant/ vowel skeletal structure, in which there are dierent slots for consonants and vowels. Caramazza et al. (2000) proposed that letters are classied as consonants or

vowels and this categorical distinction then plays a key role in processes such as the construction of syllables in speech production. Nespor et al. (2003) suggested that the rapid classication of letters as consonants or vowels allows a division of labor between the their processing, with vowels being used to help the reader interpret grammar, whereas the role of consonants is to aid in accessing the internal lexicon. The important point here is that there is now considerable opinion that consonants and vowels are processed dierently. Thus, although none of these theories would appear to make any specic predictions as to whether TL similarity eects would vary as a function of whether the transposed letters are vowels or consonants, any theories proposing consonant/vowel dierences would be informed by the results of such a comparison. For these reasons, it seemed important to examine whether TL similarity eects do dier as a function of whether the transposed letters are vowels or consonants. What should be explicitly noted is that because the orthographic representations in the SERIOL and SOLAR models do not convey consonant/vowel status, TL similarity eects with nonadjacent letters should be the same for consonant and vowel transpositions. Thus, if consonant/vowel dierences are observed in the present experiments, at the very least, these models will need an additional mechanism in order to explain those dierences.

Experiment 1 The prime types in Experiments 1a were: (1) identity (casino-CASINO), (2) one-letter replacement nonword (casiro-CASINO), (3) nonadjacent TL nonword (canisoCASINO) and (4) two-letter replacement nonword (caviro-CASINO). As noted, both SERIOL and SOLAR predict that the latencies for these four conditions should increase monotonically between the rst and fourth conditions. The only dierence between Experiments 1a and 1b was that the identity condition in Experiment 1a was replaced by an unrelated nonword condition in Experiment 1b. Thus, Experiment 1b allowed us to better gauge the sizes of the form priming eects in the other three conditions (by comparing their latencies to that in the unrelated condition). It also allowed us the opportunity to replicate the main nding of Experiment 1a, the advantage of the nonadjacent TL prime condition over the two-letter replacement letter prime condition. Method Participants Fifty-six students from the University of Valncia e received course credit for participating in the experiment

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(28 in Experiment 1a and 28 in Experiment 1b). All of them either had normal or corrected-to-normal vision and were native speakers of Spanish. Materials The targets were 128 Spanish words of six letters (mean word frequency per one million words in the Alameda & Cuetos, 1995, count: 42, range: 2418; mean Colthearts N: 2.3, range: 011). The targets in Experiment 1a were presented in uppercase and were preceded by primes in lowercase that were: (1) the same as the target (identity condition), e.g., casino-CASINO, (2) the same except for the substitution of one internal letter (always the fth letter; one-letter dierent condition), casiro-CASINO, (3) the same except for a transposition of the third and the fth letters (nonadjacent TL condition), caniso-CASINO, and (4) the same except for the substitution of two internal letters (the third and the fth letters; two-letter dierent condition), caviro-CASINO. Except in the identity condition, the primes were always nonwords. An additional set of 128 nonwords of six letters was included for the purposes of the lexical decision task (mean Colthearts N: 1.0, range: 05). The manipulation of the nonword trials was the same as that for the word trials. All items had a CV.CV.CV syllabic structure. Four lists of materials were constructed so that each target appeared once in each list, but each time in a dierent priming condition. Dierent groups of participants were used for each list. The materials of Experiment 1b were the same as in Experiment 1a, except that the identity primes were replaced by unrelated nonword primes (e.g., nomero-CASINO). The related pairs are given in the Appendix. Procedure Participants were tested in groups of four to eight in a quiet room. Presentation of the stimuli and recording of response times were controlled by Apple Macintosh Classic II microcomputers. The routines for controlling stimulus presentation and reaction time collection were obtained from Lane and Ashby (1987) and from Westall, Perkey, and Chute (1986), respectively. Reaction times were measured from target onset until the partic-

ipants response. On each trial, a forward mask consisting of a row of six hash marks (######) was presented for 500 ms in the center of the screen. Next, a centered lowercase prime was presented for 50 ms. Primes were immediately replaced by an uppercase target item, which remained on the screen until the response. Participants were instructed to press one of two buttons on the keyboard to indicate whether the uppercase letter string was a legitimate Spanish word or not (c for yes and z for no). Participants were in structed to make this decision as quickly and as accurately as possible. Participants were not informed of the presence of lowercase items. Each participant received a dierent order of trials. Each participant received a total of 20 practice trials (with the same manipulation as in the experimental trials) prior to the 256 experimental trials. The whole session lasted approximately 15 min. Results and discussion Incorrect responses (6.7% of the data for word targets) and reaction times less than 250 ms or greater than 1500 ms (0.6% of the data for word targets) were excluded from the latency analysis. Although the critical contrast was the comparison between the nonadjacent TL condition and its control condition (i.e., the twoletter dierent priming condition), we also conducted F tests, based on both the subject (F 1) and item (F 2) means, for the following contrasts: one-letter dierent primes vs. two-letter dierent primes, one-letter dierent primes vs. identity primes (for Experiment 1a), and twoletter dierent primes vs. unrelated primes (for Experiment 1b). To extract the variance due to the error associated with the lists, List was included as a dummy variable in all comparisons. All signicant eects had p values less than the .05 level. The mean response times and error percentages from the subject analysis are presented in Table 1. Experiment 1a Word data. Targets preceded by a nonadjacent TL prime were responded to 21 ms faster than the targets preceded by a two-letter dierent prime, F 11; 24

Table 1 Mean lexical decision times (in ms) and percentage of errors (in parentheses) for word and nonword targets in Experiment 1 Type of prime Identity Experiment 1a Word trials Nonword trials Experiment 1b Word trials Nonword trials 598 (4.5) 695 (6.3) One-letter dierent 615 (6.9) 691 (3.9) 665 (6.1) 822 (5.4) NonAdj. TL 628 (7.8) 694 (5.0) 679 (6.4) 814 (7.1) Two-letter dierent 649 (8.6) 696 (3.3) 696 (6.4) 813 (6.4) 703 (6.7) 832 (6.0) Unrelated

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13:00; F 21; 124 10:02. In addition, targets preceded by a one-letter dierent prime were responded to 34 ms faster than the targets preceded by a two-letter dierent prime, F 11; 24 42:18; F 21; 124 35:58, and targets preceded by an identity prime were responded to 13 ms faster than the targets preceded by a one-letter dierent prime, F 11; 24 14:14; F 21; 124 18:76. None of the contrasts on the error data were statistically signicant (all ps > :10). Nonword data. In the latency analyses, there were virtually no dierences across the dierent priming conditions. With respect to the error data, participants committed 1.7% more errors to nonwords preceded by a nonadjacent TL prime than to nonwords preceded by a two-letter dierent prime (5.0 vs. 3.3%, respectively), F 11; 24 7:42; F 21; 124 3:10, p :08. The other two contrasts did not approach signicance (all ps > :10). Experiment 1b Word data. As in Experiment 1a, targets preceded by a nonadjacent TL prime were responded to 17 ms faster than the targets preceded by a two-letter dierent prime, F 11; 24 7:47; F 21; 124 5:21, and targets preceded by a one-letter dierent prime were responded to 31 ms faster than the targets preceded by a two-letter dierent prime, F 11; 24 42:18; F 21; 124 20:11. Finally, the 7-ms dierence between targets preceded by a two-letter dierent prime and the targets preceded by an unrelated prime was not signicant, both ps > :10. The error analyses did not reveal any signicant eects (all ps > :10). Nonword data. There were virtually no dierences across the dierent priming conditions in either the latency or error data. The results were straightforward. There was a sizable priming eect (1721 ms) from nonadjacent TL nonword primes relative to the appropriate orthographic

control condition (i.e., the two-letter dierent condition) in both Experiments 1a and 1b (also see Fig. 1). Thus, it appears that nonadjacent TL nonwords do activate, to a greater degree than two-letter dierent nonwords, the lexical representation of their base words. Note also that the pattern of priming eects across the various conditions is consistent with the predictions of the SERIOL and SOLAR models using their default parameter settings (see Introduction). The only possible exception to this is the comparison between the unrelated word primes and the two-letter dierent primes. Although two-letter dierent primes are much more similar to their base words than unrelated primes according to both models, there was no signicant latency dierence between the two conditions.

Experiment 2 The primetarget conditions in Experiment 2 were the same as in Experiment 1b, except that the transposed letters were vowels (e.g., anamil-ANIMAL vs. anemolANIMAL) instead of consonants. Method Participants Twenty-four students from the same population as in Experiment 1 participated in this experiment. Materials The word targets were 128 Spanish words of six letters (mean word frequency per one million words in the Alameda & Cuetos, 1995, count: 28, range: 1379; mean Colthearts N: 1.4, range: 08) and 128 nonwords of six letters (mean Colthearts N: 0.6, range: 05). All word and nonword targets had vowels in positions three and

Fig. 1. Response times for word targets in the dierent primetarget relationships in Experiments 1 and 2.

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ve; e.g., the word ANIMAL, or the nonword ECUDAR. The primetarget conditions were the same as in Experiment 1b: (1) the one-letter dierent condition, e.g., animol-ANIMAL, (2) the nonadjacent TL condition, anamil-ANIMAL, (3) the two-letter dierent condition, anomel-ANIMAL, and (4) the unrelated condition, agapn-ANIMAL. Primes were always nono words. The manipulation for the nonword trials was the same as that for the word trials. Four lists of materials were constructed so that each target appeared once in each list, but each time in a dierent priming condition. Dierent groups of participants were used for each list. The related pairs are given in the Appendix. Procedure The procedure was the same as in Experiment 1. Results and discussion Incorrect responses (5.9% of the data for word targets) and reaction times less than 250 ms or greater than 1500 ms (1.9% of the data for word targets) were excluded from the latency analysis. As in Experiment 1, the critical contrast was the comparison between the nonadjacent TL condition and its orthographic control condition (i.e., the two-letter dierent condition). The mean response times and error percentages from the subject analysis are presented in Table 2. Word data. Unlike in Experiment 1, targets preceded by a nonadjacent TL prime were responded to 9 ms slower (rather than faster) than the targets preceded by a two-letter dierent prime. This dierence was not signicant (both ps > :20). In addition, targets preceded by a one-letter dierent prime were responded to 15 ms faster than the targets preceded by a two-letter dierent prime, F 11; 20 6:02; F 21; 124 6:85, and targets preceded by a two-letter dierent prime were responded to 20 ms faster than the targets preceded by an unrelated prime, F 11; 20 9:54; F 21; 124 4:50. Finally, the 11-ms dierence between the nonadjacent TL priming condition and the unrelated priming condition approached signicance in the subject analysis, F 11; 20 4:09, p :057; F 21; 124 1:20. The error analyses did not reveal any signicant eects. Nonword data. There were virtually no dierences across the dierent priming conditions in either the latency or error data.

Although the results of Experiment 2 parallel those of Experiment 1 in some ways, the central nding is that priming eects from nonadjacent TL nonwords do not seem to occur when the TL nonwords are created by transposing two vowels (see Fig. 1).

Experiment 3 The main goal of Experiment 3 was to establish the generality of the priming eects obtained in Experiments 1 and 2, with a new set of (710 letter long) words. For reasons of design eciency, we focused on the transposed-letter condition and its corresponding orthographic control condition, both for consonants (e.g., tradegia-TRAGEDIA vs. trabepia-TRAGEDIA) and for vowels (absuloto-ABSOLUTO vs. abselito-ABSOLUTO). The SOA was the same as in Experiments 1 and 2 (50 ms). Method Participants Sixty-two students from the Universidad de Deusto took part in the experiment. All of them either had normal or corrected-to-normal vision and were native speakers of Spanish. Materials The targets were 80 Spanish words that were 710 letters long. Forty of these words (mean word frequency per one million words in the Alameda & Cuetos, 1995, count: 69, range: 28210; mean Colthearts N: 0.47, range: 02) were presented in uppercase and were preceded by primes in lowercase that were: (1) the same except for a transposition of two internal consonants (either letter positions 35 or 4-6; nonadjacent TL-consonant condition), tradegia-TRAGEDIA, or (2) the same except for the substitution of the corresponding internal consonants, trabepia-TRAGEDIA. The remaining forty of these words (mean word frequency per one million words in the Alameda & Cuetos, 1995; count: 68, range: 31143; mean Colthearts N: 0.37, range: 02) were also presented in uppercase and were preceded by primes in lowercase that were: (1) the same except for a transposition of two internal vowels (either letter positions 35 or 46; nonadjacent TL-vowel condition), absuloto-ABSOLUTO, or

Table 2 Mean lexical decision times (in ms) and percentage of errors (in parentheses) for word and nonword targets in Experiment 2 Type of prime One-letter dierent Word trials Nonword trials 691 (4.9) 819 (4.3) Nonadj TL 714 (6.2) 818 (5.5) Two-letter dierent 707 (6.0) 814 (6.3) Unrelated 725 (6.5) 820 (4.2)

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M. Perea, S.J. Lupker / Journal of Memory and Language 51 (2004) 231246 Table 3 Mean lexical decision times (in ms) and percentage of errors (in parentheses) for word and nonword targets in Experiment 3 Type of prime Nonadj TL Word trials Consonants Vowels Nonword trials Consonants Vowels Two-letter dierent 591 (5.3) 587 (2.9) 700 (7.3) 705 (8.6) Priming

(2) the same except for the substitution of the corresponding internal vowels, abselito-ABSOLUTO. Primes were always nonwords. An additional set of 80 target nonwords that were 710 letters long was included for the purposes of the lexical decision task. The manipulation of the nonword trials was the same as that for the word trials. Two lists of materials were constructed so that each target appeared once in each list, but each time in a different priming condition. Dierent groups of participants were used for each list. The pairs are given in the Appendix. Procedure Participants were tested individually in a quiet room. Presentation and timing of stimuli were controlled by DMDX software (Forster & Forster, 2003) on a PC computer. Reaction times were measured from target onset until the participants response. On each trial, a forward mask consisting of a row of ten hash marks (##########) was presented for 500 ms in the center of the screen. Next, a centered lowercase prime was presented for 50 ms. Primes were immediately replaced by an uppercase target item, which remained on the screen until the response. Participants were instructed to press one of two buttons on the keyboard to indicate whether the uppercase letter string was a legitimate Spanish word or not (m for yes and z for no). Participants were instructed to make this decision as quickly and as accurately as possible. Participants were not informed of the presence of lowercase items. Each participant received a dierent order of trials. Each participant received a total of 24 practice trials (with the same manipulation as in the experimental trials) prior to the 160 experimental trials. The whole session lasted approximately 12 min. Results and discussion Incorrect responses (3.8% of the data for word targets) and reaction times less than 250 ms or greater than 1500 ms (0.2% of the data for word targets) were excluded from the latency analysis. The mean response times and error percentages from the subject analysis are presented in Table 3. ANOVAs based on the subject and item mean correct response latencies and error rates were conducted based on a 2 (Relatedness: transposition, control) 2 (Type of transposition/replacement: consonants, vowels) 2 (List: list 1, list 2) design. Word data. Targets preceded by a nonadjacent TL prime were responded to 12 ms faster than the targets preceded by a two-letter dierent prime, F 11; 60 21:42; F 21; 76 18:55. Response times to targets that involved the transposition/replacement of two consonants did not dier from the response times to targets that involved the transposition/replacement of two vowels, both F s < 1. More important, the interaction of

573 (3.6) 581 (3.5) 697 (8.5) 708 (7.5)

18 (1.7) 6 ()0.6) 3 ()1.2) )3 (1.1)

the two factors was signicant, F 11; 60 4:37; F 21; 76 3:98: there was a signicant 18-ms relatedness eect for the transposed consonants, F 11; 60 32:29; F 21; 38 29:35, whereas there was a small (6 ms) nonsignicant relatedness eect for the transposed vowels, F 11; 60 2:19, p > :10; F 21; 38 2:02, p > :10. The ANOVA on the error data showed a similar pattern: The interaction of the two factors was also signicant, F 11; 60 4:94; F 21; 76 4:88: there was a 1.7% relatedness eect for the transposed consonants, F 11; 60 6:52; F 21; 38 4:71, whereas there was a )0.6% nonsignicant relatedness eect for the transposed vowels, both F s < 1. Nonword data. The ANOVAs on the nonword data did not reveal any signicant eects (all ps > :10). The results were again clear-cut. When the transposed letters were consonants (tradegia-TRAGEDIA), there was a sizable priming eect (18 ms) from nonadjacent TL nonword primes relative to the orthographic control condition (i.e., the two-letter dierent condition; trabepia-TRAGEDIA). When the transposed letters were vowels, there was only a nonsignicant 6 ms eect.

Experiment 4 The results of the rst three experiments have clearly shown that nonadjacent TL-consonant nonwords activate, to a greater degree than two-letter dierent nonwords, the lexical representation of their base words. Further, they demonstrate that this eect seems to occur only for consonants (Experiments 1a, 1b, and 3) and not for vowels (Experiments 2 and 3). The goal of Experiment 4 was to obtain converging evidence on the role of nonadjacent TL consonants vs. vowels using another experimental technique: a single-presentation lexical decision task. We used the masked primes of Experiment 3 as the nonword targets. A set of word targets was selected for the purposes of the lexical decision task. It is a well established nding that wordlike nonwords (e.g., one-letter dierent nonwords) produce

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slower no responses and more errors than nonwordlike nonwords (e.g., those with no orthographic neighbors) in lexical decision tasks (e.g., Coltheart, Davelaar, Jonasson, & Besner, 1977; Forster & Shen, 1996; Perea & Rosa, 2000; Sears, Hino, & Lupker, 1995; Siakaluk, Sears, & Lupker, 2002). The explanation is that the former partially activate the lexical representations of their word neighbors. As a result, additional time is needed for the activation levels to settle and for the participant to realize that no word unit is being activated over threshold. Similarly, if nonadjacent TL nonwords (e.g., TRADEGIA) activate the lexical representation of their corresponding base words (TRAGEDIA) to a higher degree than orthographic controls (TRABEPIA), one would expect a higher rate of word responses and longer latencies for the nonadjacent TL nonwords than for the orthographic controls in the present experiment. Finally, based on the results of the previous experiments, we would expect this eect to exist for consonant transpositions but not necessarily for vowel transpositions. One nal point should be noted. There is good evidence for syllable-frequency eects in single-word presentation lexical decision tasks in Spanish (e.g., Carreiras & Perea, 2004; Perea & Carreiras, 1998). These eects, however, are restricted to the frequency of the initial syllable. Therefore, in order to avoid any uncontrolled eects of initial syllable frequency, all the experimental nonwords in Experiment 4 maintained the initial syllable of their base words. That is, the transposed/replacement letters all came from later syllables, meaning that the experimental and control nonwords had the same initial syllables. Method Participants Twenty-six students from the Universidad de Deusto took part in the experiment. All of them either had normal or corrected-to-normal vision and were native speakers of Spanish. Materials The 80 word targets from Experiment 3 were used as the base words for the four nonword conditions (nonadjacent TL-consonant nonword and its corresponding controle.g., TRADEGIA and TRABEPIA, nonadjacent TL-vowel nonword and its corresponding control e.g., ABSULOTO and ABSELITO). That is, the nonword targets in the present experiment had been the nonword primes in the previous experiment. The TLconsonant nonwords and their orthographic controls both had, on average, 0.075 word neighbors (range 01) (note that all these neighbors were always very-lowfrequency words, with a frequency no higher than 3 per million). The mean token trigram frequencies for the

critical trigrams in the transposed and the replacementletter conditions (e.g., DEG and BEP in TRAGEDIA and TRAPEDIA) were 255 and 226 per million (t < 1), respectively. All these trigrams occurred in Spanish words. The mean token bigram frequencies for the critical bigrams in the two conditions (e.g., AD, DE, EG, GI in TRADEGIA vs. AP, PE, ED, DI in TRAPEDIA) were comparable (22,780 vs. 20,560 per million, for the transposed and replacement-letter conditions, respectively; p > :15). The TL-vowel nonwords and their orthographic controls had no orthographic neighbors. The mean token trigram frequencies for the critical trigrams in the transposed- and replacement-letter conditions (e.g., ULO and ELI in ABSULOTO and ABSELITO) were 632 and 583 per million (t < 1), respectively. All these trigrams occurred in Spanish words. (Note that the trigrams were more frequent for VCV than for CVC transpositions/replacements because of the obvious fact that there are only ve vowel letters.) The mean token bigram frequencies of the critical bigrams in the two conditions (e.g., SU, UL, LO, OT in ABSULOTO vs. SE, EL, LI, IT in ABSELITO) were comparable (24,996 vs. 26,230 per million in the transposed- and replacement-letter conditions, respectively; p > :15). As noted, in all cases, the rst syllable of the base word remained unchanged. Two lists of materials were constructed so that if the TL-consonant nonword TRADEGIA appeared in one list, its orthographic control (TRABEPIA) would appear in the other list. Dierent groups of participants were used for each list. An additional set of 80 words that were 710 letters long (mean frequency per million words: 38, range: 1206) was included for the purposes of the lexical decision task. Procedure Participants were tested individually in a quiet room. Presentation and timing of stimuli were controlled by DMDX software (Forster & Forster, 2003) on a PC computer. On each trial, a centered uppercase target item remained on the screen until response. Participants were instructed to press one of two buttons on the keyboard to indicate whether the letter string was a legitimate Spanish word or not (m for yes and z for no). Participants were instructed to make this decision as quickly and as accurately as possible. Each participant received a dierent order of trials. Each participant received a total of 24 practice trials (with the same manipulation as in the experimental trials) prior to the 160 experimental trials. The whole session lasted approximately 12 min. Results and discussion Incorrect responses (19.5% of the data for nonword targets, 5.3% for the word targets) and reaction times

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Table 4 Mean lexical decision times (in ms) and percentage of errors (in parentheses) for nonword targets in Experiment 4 Type of nonword Nonadj TL Consonants Vowels 943 (43.5) 915 (24.4) Two-letter dierent 869 (4.6) 853 (5.4) Dierence 74 (38.9) 62 (19.0)

Note. The mean correct RT for word trials was 752 ms and the error rate was 5.3%.

less than 250 ms or greater than 1500 ms (6.2% of the data for nonword targets) were excluded from the latency analysis. The mean response times and error percentages from the subject analysis are presented in Table 4. ANOVAs, based on both subject and item mean response latencies and error rates to nonword targets, were conducted based on a 2 (Type of nonword: transposition, control) 2 (Type of transposition/replacement: consonants, vowels) 2 (List: list 1, list 2) design. Nonword targets created by transposing two nonadjacent letters were responded to 68 ms slower than nonwords created by replacing those two letters, F 11; 24 34:86; F 21; 76 44:48. Nonwords created by transposing/replacing two consonants had slower latencies than nonwords created by transposing/replacing two vowels, F 11; 24 4:60; F 21; 76 3:97. The dierence between the transposed-letter nonwords and the replacement-letter nonwords was only slightly greater for the transpositions involving consonants than for the transpositions involving vowels (74 vs. 62 ms). Hence, the interaction between the two factors was not signicant, both ps > :10.3 The ANOVA on the error data showed that there were signicantly fewer errors to replacement-letter nonwords than to transposed-letter nonwords, F 11; 24 116:21; F 21; 76 207:63. There were also more errors to nonwords created by transposing/replacing two consonants than to nonwords created by
3 The lack of an interaction between Type of nonwords and Type of transposition/replacement was not due to the use of a 1500 ms cuto. With a more conservative cuto, the critical interaction was also not signicant (e.g., with a 2000 ms cuto, the TL eect was 94 vs. 79 ms, for the consonant TL nonwords and for the vowel TL nonwords, respectively). Nonetheless, there were some signs of a greater TL eect for consonant nonwords than for vowel nonwords when examining the RT distributions (using all the correct responses): As in the mean RT analyses, the TL eect was only slightly greater for consonant nonwords than for vowel nonwords in the bulk of the RT distributions (.5 quantile: 101 vs. 89 ms for the consonant and the vowel TL nonwords, respectively); however, the TL eect was substantially greater for consonant nonwords than for vowel nonwords with the slowest responses (.9 quantile; 213 vs. 86 ms, respectively; interaction: F 1; 24 6:18, p:025).

transposing/replacing two vowels, F 11; 24 23:06; F 21; 76 15:24 The interaction of the two factors was also signicant, F 11; 24 54:22; F 21; 76 24:31: the transposition-letter eect was substantially larger for the nonwords created by transposing two consonants (38.9%) than for the nonwords created by transposing two vowels (19.9%). These two simple main eects were both signicant: for consonants, F 11; 24 168:15; F 21; 38 167:23, for vowels, F 11; 24 40:00; F 21; 38 50:95. Consistent with the previous experiments, transposed-letter nonwords created by transposing two nonadjacent consonants seem to activate their base word to a considerable degree. The eect was quite dramatic in terms of error rates (43.5 vs. 4.6%) for the transposedletter condition and its orthographic control, respectively. One dierence between the present experiment and the previous experiment is that this time, there were signicant eects for transposing two nonadjacent vowels. (With these same materials, using the masked priming technique, there was only a nonsignicant 6-ms priming eect in Experiment 3.) In fact, only in the error data was there clear evidence that the eect of transposing two vowels was smaller than the eect of transposing two consonants.4 It appears, therefore, that vowel transposition nonwords are perceptually similar to their base words, they are simply less similar than consonant transposition nonwords. Finally, it is important to note that the very high error rates for the TL nonwords do not reect a very lenient decision criterion for word responses, but rather they reect the high degree of similarity between the TL nonwords and their corresponding base words. Experiment 4 has been replicated with a dierent set of items in another lab in Spain, and the error rates for the TL-vowel nonwords and the TL-consonant nonwords were virtually the same (41 vs. 22%, respectively) as those reported here (43 vs. 24 %). Indeed, for any skilled reader of Spanish, the TL nonword DEYASUNO seems to activate its base word (DESAYUNO) to a large degree, and it is rather dicult to process/pronounce correctly a TL nonword such as DEYASUNO under time pressure.
This experiment was replicated with a shorter stimulus duration (200 ms, followed by a pattern mask composed of a series of # signs). The pattern of results was essentially the same, except that error rates were slightly higher. For the transposed consonant conditions, error rates were 46.3 vs. 9.0% for the transposed-letter nonwords and their orthographic controls (the mean RTs were 865 vs. 779 ms, respectively). For the transposed vowel conditions, error rates were 30.5 vs. 9.0% for the transposed-letter nonwords and their orthographic controls (the mean RTs were 846 vs. 773 ms, respectively). That is, the transposed-letter similarity eects were noticeably greater for the transposition of consonants than for the transposition of vowels, especially in the error data (37.3 vs. 21.5%; 86 vs. 73 ms).
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General discussion The present experiments allow the following conclusions: (1) nonword primes created by transposing two nonadjacent letters produce masked priming eects relative to the appropriate orthographic controls (canisoCASINO vs. caviro-CASINO), (2) these priming eects seem to be restricted to the case in which the transposed letters are consonants (i.e., anamil-ANIMAL is no faster than the orthographic control anomel-ANIMAL), and (3) in a single-presentation lexical decision task, both TL-consonant nonwords (TRADEGIA) and TL-vowel nonwords (ABSULOTO) produce longer latencies and more errors than control nonwords (TRABEPIA, ABSELITO), however, TL-consonant nonwords are somewhat more problematic (as indicated by the false positive error rates) than TL-vowel nonwords. The presence of nonadjacent TL similarity eects poses a clear problem for models that assume a positionspecic coding scheme (e.g., the interactive-activation model and its extensions). As stated in the Introduction, the only real way to try to explain an eect of this sort is to incorporate the notion of noise in the coding process, so that the representation of one letter is not immediately tied to a single letter position but, instead, extends activation into nearby letter positions. That is, the letter S in the nonadjacent TL nonword CANISO would provide considerable activation to the representation of the letter S in letter position 5, somewhat less activation to the representations of the letter S in adjacent positions (4 and 6), as well as at least some activation to the representation of the letter S in letter position 3. Ratclis (1981) letter coding model is a model based on these ideas. In Ratclis model, each letter in a letter string creates a distribution of activation over positions. As a result, the representation of a letter in a given position would be activated by the appearance of that letter in any nearby letter position. Therefore, the overall overlap between CASINO and the TL nonwords CANISO would be substantially higher than that between CASINO and the orthographic control CAVIRO (see Gmez, Perea, & Ratcli, 2003). o Even if Ratclis scheme were integrated into the interactive-activation architecture, however, new problems may emerge. That is, as Davis (1999) has noted, noise of this sort in the visual input code would harm the ability of IA-based models to recognize highly familiar inputs. Thus, it would be extremely unlikely that IA models with this modied coding scheme would still be able to successfully simulate many of the eects they now can. Further, there would, of course, be an additional problem with respect to the present data. An account of this sort has no way to distinguish between vowel and consonant processing. An alternative way of explaining nonadjacent TL priming would be to assume either a spatial coding

scheme (as in the SOLAR model) or a letter-tagging coding scheme (as in the SERIOL model). Both of these coding schemes can readily capture the overall pattern of TL eects in the present experiments. That is, in both cases, the similarity between the nonadjacent TL nonwords and their corresponding base words is higher than the similarity between the orthographic controls (the two-letter dierent nonwords) and their corresponding base words (see Introduction), leading to the prediction of nonadjacent TL priming eects. (Similar predictions can be made by very recently proposed letter-coding models: the open-bigram model, Grainger & van Heuven, 2003; and the overlap model, Gmez et al., 2003.) o The problem, however, is still that, like a modied position-specic model, the current versions of all these models do not make any distinctions between the processing of vowels and consonants. Thus, some of the model assumptions would need to be altered to accommodate the presence of priming eects from nonadjacent TL nonwords based on consonants but not on vowels (Experiments 13) and the presence of a substantially larger number of false positive errors in a single-presentation lexical decision task for TL-consonant nonwords than for TL-vowel nonwords (Experiment 4). To allow these models to explain the present data, one would need to start by assuming that very early in processing, the consonant/vowel status of a given letter is obtained and that this information allows the two types of letters to be segregated from each other. For example, some formulations (e.g., Berent et al., 2001; Caramazza & Miceli, 1990; Tainturier & Caramazza, 1996) assume that very early in processing, the word CASINO activates a C.V.C.V.C.V structure with the identied letters then lling those slots. This segregation of consonants from vowels, however it is accomplished, would then allow subsequent processing of the consonant letters c s n to dier from the processing of the vowel letters a i o (e.g., see Caramazza et al. (2000) and Nespor et al. (2003) for discussions of how consonants and vowels might play dierent roles in lexical processing). One would next need to assume that the dierent processing that consonants and vowels underwent made it somewhat more likely that one would observe writing/reading errors involving transpositions of two consonants than transpositions of two vowels. Note that one implication of these assumptions would be that it should be very unusual to nd errors involving transpositions of one consonant and one vowel. Indeed, the usual pattern obtained with brain damaged patients is the transposition/replacement of vowels by vowels, and consonants by consonants (see Caramazza et al., 2000). One way in which a segregation of consonants and vowels could occur would be for the orthographic coding of vowels to be more distinct than that of consonants. For example, in Ratclis (1981; Gmez et al., o

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2003) framework, the activation gradients corresponding to vowels and consonants could be assumed to dier (see Drewnowski, 1980, for a similar suggestion). More specically, Ratclis model could explain the present ndings by assuming that the activation of consonant graphemes extends further (i.e., into nearby letter slots) than the activation of vowel graphemes.5 Within the framework of the SERIOL or SOLAR models, the key aspect of the models would be the set of activation levels of the sublexical units. As noted, the SOLAR model uses activation levels to code order information (i.e., the rst letter is coded by the highest activation value, the second letter is coded with a slightly smaller activation value, etc.). According to the model, there is a constant ratio (e.g., 2:1) between the activation levels of successive letters (the so-called the invariance principle, see Davis, 1999), without any distinctions between consonants and vowels. The SERIOL model works in a fairly similar way. In that model (Whitney, 2001; Whitney & Berndt, 1999), once letter positions have been tagged, the activation of a given letter node is set to 0.7position-1 . Thus, for the word CASINO, the activation levels for its component letters would be C-1, A0.7, S-0.49, I-0.34, N-0.24, and O-0.17, with, again, no distinction between consonants and vowels. If, as suggested above, consonant/vowel status is ascertained very early in processing, these models could propose that the letters achieve dierent activation levels depending on whether they are consonants or vowels. Probably, the most reasonable way to do this would be either to propose a heightened activation level if the letter were a vowel (making it more distinct) or a lowered activation level if a letter were a consonant (making it less distinct). For example, in SERIOLs framework, the consonant S in CASINO could have a reduced activation level of 0.40, which would make its level more similar to that of the N (although the activation level of the N would, presumably, also be lowered). Thus, the S and the N would be more likely to show TL eects than the A and the I (i.e., CANISO would be more similar to CASINO than CISANO would). A similar x for SOLAR would lead to similar predictions. There are, of course, limits to the degree to which activation levels can be changed without altering the basic structure of the models. For example, the activation level for the consonant S in position 3 in CASINO cannot be reduced below the activation level for the vowel I in position 4 or else the activation levels would
A possible reason why the activation gradient for consonants might be broader than the activation gradient for vowels could be simply because consonants are less frequent than vowels (rather than being a function of basic structural dierences). Although this is an unlikely explanation (e.g., see Caramazza et al., 2000; for discussion), it is, potentially, a testable hypothesis.
5

no longer accurately code letter order. Whether changes to the models within these limits would actually allow the models to predict the vowel/consonant dierences and whether these changes would then harm the models abilities to explain other letter coding results would, of course, be questions for future research. An additional point that should be noted is that SERIOL, unlike SOLAR, incorporates a bigram level. Thus, in theory, SERIOL has the potential to explain the vowel/consonant dierences in terms of activation patterns at that level, rather than at the letter level. For example, vowel bigrams (i.e., the AI pair, the AO pair, and the IO pair in CASINO) could be assumed to have higher activation levels than consonant bigrams (e.g., SN). In this way, CISANO would not be very similar to CASINO because the set of bigram codes activated by CISANO would not include the crucial AI pair. Hence, CISANO would not be expected to produce much priming of CASINO. Alternatively, it would be possible to propose a locus of the consonant/vowel dierences observed here which is entirely outside of the architecture of the models under consideration. For example, one could propose that those dierences arise at the sub-lexical phonological level.6 Subjectively, the transposition of two consonants does appear to preserve more of the sound of the original word than the transposition of two vowels (e.g.,  compare the TL-consonant nonword LIREBACION to  its base word, LIBERACION, in contrast to the TL vowel nonword LIBARECION). Indeed, vowel sounds seem to become phonologically relevant earlier in life than consonant sounds (see Bertoncini, Bijeljac-Babic, Jusczyk, Kennedy, & Mehler, 1988) and (at least in Spanish) young children spell the vowel sounds before consonant sounds (e.g., A_I_O_A for MARIPOSA, the Spanish for buttery; see Ferreiro & Teberosky, 1982). If the masked priming eects we have observed are phonological eects (see Hino, Lupker, Ogawa, & Sears, 2003), the obvious prediction would be that TL-consonant primes would be better primes than TL-vowel primes because TL-vowel primes would be more dissimilar to their targets than TL-consonant primes. In contrast, if negative lexical decision responses are based more on an analysis of the orthographic structure of the letter string, it would make sense that the consonant/ vowel dierence might diminish. Further research and simulation work is, of course, needed to examine this possibility. With respect to the general issue of vowel-consonant dierences and the impact of phonological codes, it is worth noting that Berent and Perfetti (1995) have proposed that because consonants and vowels are distinct linguistic entities (with the argument being that this is a universal structural distinction; see also Caramazza
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We thank Carol Whitney for this suggestion.

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et al., 2000), the time course of phonological assembly for vowels and consonants diers. In particular, Berent and Perfetti have produced evidence that consonants are processed faster than vowels in English (see also Lee et al., 2001). As just discussed, the present ndings do support the claim that there is an essential structural distinction between vowels and consonants. However, an explanation based on Berent and Perfettis specic claim about the time course of phonological assembly for vowels and consonants is not likely to be relevant here. Instead, it seems more likely that the time course of vowel versus consonant processing would actually vary as a function of the characteristics of a given language. Specically, in English, the graphemephoneme relationship for consonants is much more consistent than it is for vowels (Brown & Besner, 1987; Carr & Pollatsek, 1985). Thus, it would follow that English consonants would be coded more rapidly than English vowels. In contrast, in Spanish and in other Romance languages the graphemephoneme relationship for vowels is at least as consistent as that for consonants. Thus, vowel coding should occur at least as rapidly for vowels as for consonants. It is not at all surprising, therefore, that Colombo, Zorzi, Cubelli, and Brivio (2003) were unable to replicate Berent and Perfettis consonant advantage using Italian. Note also that, in spite of the fact that English and Spanish dier dramatically in terms of the consistency of phonological coding of vowels, the present results do have parallels in English. As noted, Perea and Lupker (2003b), using English stimuli, also found substantially larger TL priming eects when the transposition involved consonants rather than vowels. In addition, an experiment parallel to Experiment 4 carried out with English stimuli in the second authors lab yielded results that completely parallel Experiment 4s results (i.e., TL similarity eects involving nonadjacent letter positions are greater with consonants than with vowels when considering the nonwords in a lexical decision task). It seems likely, therefore, that the basis of the consonant/ vowel dierences is something that is common to the two languages. Thus, what we have attempted to do is to try to explain these dierences in terms of a general principle (e.g., establishing the exact position of vowels may be more important for a words identication than establishing the exact position of consonants) that would be relevant in both languages. To summarize, we have provided empirical evidence that TL similarity eects occur even when the transposed letters are not adjacent. These results are as predicted by models with a spatial or letter-tagging coding scheme (SOLAR and SERIOL models). In addition, however, we discovered that this conclusion applies especially to consonants, supporting the claims that there are some basic processing dierences between vowels and consonants (see Caramazza et al., 2000). This latter

fact cannot be readily explained by either the SOLAR or SERIOL models. Whether those models can be amended to allow them to explain the present results, as well as consonant/vowel dierences in general, is a question for future research. Appendix. Related pairs in Experiments 13 Experiment 1
The items are arranged in quadruplets in the following order: one-letter dierent prime, TL-prime, RL-prime, and target word. camafa, cadama, cafasa, CAMADA; gaceda, gateca, gadena, GACETA; sonifo, sodino, sovo, SONIDO; morabo, modaro, mobaso, MORADO; felivo, fenilo, fevito, FELINO; dibupo, dijubo, diputo, DIBUJO; humaro, hunamo, huraco, HUMANO; romaso, ronamo, rosavo, ROMANO; manesa, marena, maseva, MANERA; masape, majase, mapane, MASAJE; basuva, barusa, bavuna, BASURA; malefa, matela, mafeda, MALETA; meduva, mesuda, mecuta, MEDUSA; cadesa, caneda, casela, CADENA; visila, vitisa, vilica, VISITA; genosa, gemona, gesova, GENOMA; tiravo, tinaro, tivaso, TIRANO; fabata, fadaba, fatala, FABADA; gemeto, gelemo, geteco, GEMELO; guca, ruga, cupa, FIGURA; mirata, midara, mitasa, MIRADA; parato, padaro, pataso, PARADO; dorato, dodaro, dotaso, DORADO; tutefa, tuleta, tufeba, TUTELA; lativo, lanito, lavido, LATINO; galoje, gapole, gajode, GALOPE; regafo, relago, refapo, REGALO; navaya, najava, nayasa, NAVAJA; corosa, conora, cosova, CORONA; veciso, venico, vesiso, VECINO; debapo, dejabo, depafo, DEBAJO; modebo, moledo, mobeto, MODELO; monela, modena, molesa, MONEDA; minudo, mituno, miduso, MINUTO; pecafo, pedaco, pefano, PECADO; golono, gosolo, gonoto, GOLOSO; pesato, pedaso, petano, PESADO; gusazo, gunaso, guzavo, GUSANO; rulefa, rutela, rufeda, RULETA; dineso, direno, diseco, DINERO; diviro, dinivo, dirico, DIVINO; camiva, casima, cavica, CAMISA; rodape, rojade, ropate, RODAJE; camivo, canimo, cavico, CAMINO; divima, disiva, dimica, DIVISA; lede, tele, debe, FILETE; relado, retalo, redafo, RELATO; pasino, paviso, panino, PASIVO; butava, bucata, buvala, BUTACA; sumivo, susimo, suvino, SUMISO; cadeva, careda, cavela, CADERA; cubaso, cunabo, cusato, CUBANO; decaso, denaco, desavo, DECANO; tariba, tara, tabima, TARIFA; madeva, mareda, maveta, MADERA; senafo, sedano, sefavo, SENADO; noveta, noleva, noteca, NOVELA; rutiva, runita, ruvifa, RUTINA; ganato, gadano, gataso, GANADO; nacito, nadico, natiso, NACIDO; pasato, padaso, patano, PASADO; comifa, codima, cosa, COMIDA; famono, fasomo, fanovo, FAMOSO; gotena, goreta, goneda, GOTERA; lujomo, lusojo, lumopo, LUJOSO; tomafe, totame, tofave, TOMATE; cocifo, codico, cono, COCIDO; zano, zarifo, zanito, ZAFIRO; docesa, doneca, dosema, DOCENA; tocabo, todaco, tobaso, TOCADO; marifo, madiro, mano, MARIDO; facela, fateca, falena, FACETA; polaso, pocalo, posato, POLACO; racino, ramico, ranino, RACIMO; herifa, hedira, heva, HERIDA; jugomo, jusogo, jumopo, JUGOSO; harisa, hanira, hasima, HARINA; herepe, hejere, hepese, HEREJE; salubo, sadulo, sabufo, SALUDO; canefa, calena, cafesa, CANELA; moliso, monilo, mosifo, MOLINO; futuvo,

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M. Perea, S.J. Lupker / Journal of Memory and Language 51 (2004) 231246  cidus, codis, acedus, ACIDOS; chicus, chacis, choces, CHI a a CAS; cuider, cuadir, cuoder, CUIDAR; origon, oregin, orugan, ORIGEN; ciegis, cioges, ciagus, CIEGOS; lluvea, llivua, llevoa, LLUVIA; examon, exeman, eximon, EXAMEN; ayudor, aya~ dur, ayedor, AYUDAR; a~ador, a~ idar, a~ edor, ANADIR; n n n reunor, reinur, reanor, REUNIR; clasos, clesas, clisos, CLASES; brutol, bratul, bretil, BRUTAL; imitor, imatir, imetor, IMITAR; clamir, clomar, clumer, CLAMOR; azcor, azac r, u u   azec AZUCAR; crnuo, creno, crin o, CRANEO; ayuner, r, a a u ayanur, ayenor, AYUNAR; deudar, deodur, deadir, DEUDOR; chinus, chanis, chenos, CHINAS; cremus, crames, crimos, CREMAS; orader, orodar, oruder, ORADOR; gui~ al, n ~ guo~il, gua~ el, GUINOL; suizes, suozis, suezas, SUIZOS; n n ahogur, ahagor, ahiger, AHOGAR; grutos, gratus, grotes, GRUTAS; ilegol, ilagel, ilugol, ILEGAL; clavol, cleval, clivol, CLAVEL; oculor, ocalur, ocelir, OCULAR; asesir, asoser, asusir, ASESOR; aner, afanir, afenor, AFINAR; anemoa, animea, anumoa, ANEMIA; acosur, acasor, acuser, ACOSAR; abulea, abilua, abeloa, ABULIA; bribn, brbin, br e o ban,  BRIBON; agitor, agatir, agetor, AGITAR; grupol, grapul, gripel, GRUPAL; apuror, aparur, apirer, APURAR; adaguo, adigao, adugeo, ADAGIO; asoler, asalor, aselir, ASOLAR; alor, afalir, afelor, AFILAR; clonir, clanor, cliner, CLONAR; orel, arol, eril, FLORAL; erizus, erozis, eruzas, ERIZOS; grasus, grosas, gruses, GRASOS; acu~ or, aca~ ur, aci~er, n n n ~ ACUNAR; fritus, frotis, frates, FRITOS; tribel, trabil, trubel, TRIBAL; brujis, brojus, brajes, BRUJOS; gru~ or, gri~ur, n n ~ gre~ ar, GRUNIR; asader, asodar, asidur, ASADOR; aludor, n  alidur, aledar, ALUDIR; crtor, cretr, cr or, CRATER; aca a t ogur, acegor, acigur, ACOGER; aboler, abilor, abelur, ABOLIR; troter, trator, triter, TROTAR; dietos, diates, diutos, DIETAS; acoter, acator, acuter, ACOTAR; peat petan, n, o   pe ton, PEATON; apag ap gan, ap gen, APAGON; olivus, n, o u olovis, olevas, OLIVOS; llanus, llonas, llines, LLANOS; alegor, alager, aligor, ALEGAR; abetus, abotes, abutis, ABETOS; trus, trafus, trifes, TRUFAS; avera, av u rea, avroa, u AVER reinor, reanir, reunor, REINAR; grifes, gros, greIA; fas, GRIFOS; peinor, peanir, peonur, PEINAR; acus, ocas, ices, FLACOS; evadur, evidar, evudor, EVADIR; acusor, acasur, acosir, ACUSAR; anotir, anator, aniter, ANOTAR; chozus, chazos, chizes, CHOZAS; crudis, crodus, crades, CRUDOS; charel, choral, cherul, CHAROL; suecis, suoces, suacis, SUECOS; apelor, apaler, apolir, APELAR; frutol, fratul, fritel, FRUTAL; alojer, alajor, alijer, ALOJAR; orugos, oragus, origes, ORUGAS.

furuto, fuvubo, FUTURO; sonoso, sorono, sosoco, SONORO; ligeso, lirego, liseyo, LIGERO; tesono, teroso, tenovo, TESORO; boleno, borelo, boneto, BOLERO; sucemo, suseco, sumevo, SUCESO; marivo, maniro, mavico, MARINO; veraco, venaro, vecaso, VERANO; rotuna, roruta, ronula, ROTURA; rebapa, rejaba, repata, REBAJA; seguso, serugo, sesuyo, SEGURO; tabavo, tacabo, tavato, TABACO; pedano, pezado, penato, PEDAZO; sonefo, soteno, sofeco, SONETO; gomisa, gonima, gosisa, GOMINA; helafo, hedalo, hefabo, HELADO; salifa, sadila, sata, SALIDA; coyobe, cotoye, coboge, COYOTE; semasa, senama, sesava, SEMANA; casiro, caniso, caviro, CASINO; butaso, bunato, busafo, BUTANO; moromo, mosoro, momovo, MOROSO; malago, mayalo, magato, MALAYO; bonifo, botino, bovo, BONITO; sujedo, sutejo, sudeyo, SUJETO; gorita, golira, gotina, GORILA; peloda, petola, pedofa, PELOTA; corava, cozara, covasa, CORAZA; celoro, cesolo, cerofo, CELOSO; bigole, bitoge, bilope, BIGOTE; vasipa, vajisa, vapira, VASIJA; balata, badala, batafa, BALADA; motino, movito, monido, MOTIVO; lejavo, lenajo, levapo, LEJANO; menuto, meduno, metuco, MENUDO; ranuva, raruna, ravusa, RANURA; tapele, tatepe, talege, TAPETE; casato, cadaso, catano, CASADO; mesefa, metesa, mefeva, MESETA; bikiri, biniki, bisili, BIKINI; payamo, pasayo, pamago, PAYASO; cabena, cazeba, caneta, CABEZA; coliva, conila, coviba, COLINA; rabivo, ranibo, ravito, RABINO; repima, resipa, remiga, REPISA; parape, pajare, papase, PARAJE; cobaga, coyaba, cogala, COBAYA.

Experiment 2
The items are arranged in quadruplets in the following order: one-letter dierent prime, TL-prime, RL-prime, and target word. agujes, agajus, agejos, AGUJAS; alemn, almen, al o a mun,  ALEMAN; animol, anamil, anomel, ANIMAL; ruides, ruodis, ruades, RUIDOS; piedod, piaded, piudod, PIEDAD; apoyur, apayor, apiyer, APOYAR; huesas, huoses, huasis, HUESOS; elegar, eliger, elogar, ELEGIR; chalut, chelat, chilot, CHALET; avidaz, avediz, avoduz, AVIDEZ; glorua, gliroa, glerua,  GLORIA; adem admes, ad s, a mos, ADEMAS; agona, u ag noa, agnua, AGON vuelis, vuoles, vuilas, VUELOS; e IA; prevoa, privea, pruvoa, PREVIA; emisar, emosir, emusar,  EMISOR; frgel, frigl, frogl, FRAGIL; nusoa, nesua, a a e a a  nisoa, NAUSEA; azota, azetoa, azitua, AZOTEA; xitas, a u e  xotis, xutas, EXITOS; planis, plonas, plines, PLANOS; acue e dar, acidur, acedor, ACUDIR; llegor, llager, llogir, LLEGAR; cuevos, cuaves, cuivos, CUEVAS; tregoa, trugea, trigoa,  TREGUA; chfar, chefr, chifar, CHOFER; llenor, llaner, o o  llunor, LLENAR; rabos, rebas, rubos, ARABES; plurel, a a a plarul, plerol, PLURAL; acidaz, acediz, acudoz, ACIDEZ; plumis, plamus, plemos, PLUMAS; nietus, niotes, niatus, NIETOS; globel, glabol, glibel, GLOBAL; criman, cremin, cramon, CRIMEN; amiges, amogis, amuges, AMIGOS; asu mer, asimur, asemor, ASUMIR; intel, initl, inatl, INUTIL; u u e viejis, viojes, viujas, VIEJOS; unided, unadid, unedod, UNI DAD; trav s, trvas, tr o e vos, TRAVES; llorer, llaror, llurer,  LLORAR; adem admen, ad min, ADEMAN; orus, eros, n, a o iras, FLORES; placir, plecar, plicor, PLACER; evitor, evatir, evutor, EVITAR; ciuded, ciadud, ciedod, CIUDAD; premuo, primeo, prumao, PREMIO; cruzor, crazur, crezir, CRUZAR;

Experiment 3
The items are arranged in triplets in the following order: TL-prime, RL-prime, and target word. Consonant transpositions/replacements: revulocin, revao  lecin, REVOLUCION; evedinte, evadonte, EVIDENTE; o enomarado, enimurado, ENAMORADO; impisoble, impusable, IMPOSIBLE; inivetable, inovatable, INEVITABLE; amenacer, amonicer, AMANECER; genareci n, genuricin, o o  GENERACION; difuciltad, difoceltad, DIFICULTAD; escelara, escilora, ESCALERA; nataruleza, nateroleza, NATURALEZA; horozinte, horuzente, HORIZONTE; adalente, adolinte, ADELANTE; anamiles, anomeles, ANIMALES; li barecin, liburici n, LIBERACION; femineno, femonuno, o o

M. Perea, S.J. Lupker / Journal of Memory and Language 51 (2004) 231246 FEMENINO; comasirio, comuserio, COMISARIO; semajente, semijonte, SEMEJANTE; avineda, avonuda, AVENIDA; op  areci n, oporucin, OPERACION; ilisun, ilosan, ILUSION; o o o o  habaticin, haboteci n, HABITACION; unorme, unaferme, o o  UNIFORME; telivesi n, teluvasin, TELEVISION; amirallo, o o amorello, AMARILLO; cominudad, comenodad, COMUNIDAD; aperace, aporuce, APARECE; litaretura, litorotura, LITERATURA; amaneza, amonuza, AMENAZA; amirecanos, amoracanos, AMERICANOS; enimego, enumago, ENEMIGO; autirodad, auturadad, AUTORIDAD; velicodad, velucadad, VELOCIDAD; unevirso, unovarso, UNIVERSO;    unacimente, unecomente, UNICAMENTE; absuloto, absileto, ~ ABSOLUTO; se~ irota, se~ urata, SENORITA; imiganar, imon n  genar, IMAGINAR; expisocin, expasucin, EXPOSICION; o o  evulocin, evilecin, EVOLUCION; opisocin, opusacin, o o o o  OPOSICION. Vowel transpositions/replacements: silimar, sitinar, SIMILAR; gusbata, gusdala, GUSTABA; namiciento, navipiento, NACIMIENTO; cr cita, cr rila, CR ITICA; condiserar, conticerar, CONSIDERAR; dortimorio, dorlinorio, DORMITORIO; esdutio, esbulio, ESTUDIO; doroles, dovotes, DOLORES; cazebas, caredas, CABEZAS; coroles, covotes, COLORES; sotilario, sobario, SOLITARIO; hatibual, halidual, HABITUAL; hernamo, hersaso, HERMANO; prospito, o  prongito, PROPOSITO; anretior, ancelior, ANTERIOR; o  esm tago, esnfago, ESTOMAGO; descapio, desragio, DESo o PACIO; platena, plafema, PLANETA; rebplica, redglica, u u  REPUBLICA; tradegia, trabepia, TRAGEDIA; ansilis, a  anritis, ANALISIS; posatio, pozalio, POTASIO; sonsira, a  soncina, SONRISA; tracidin, trasibin, TRADICION; zatao o pos, zabagos, ZAPATOS; nodevad, nobesad, NOVEDAD;  trnimos, trvisos, TERMINOS; canimos, carisos, CAMINOS; e e prosefor, procetor, PROFESOR; ornedador, orcebador, ORDENADOR; juscitia, jusrilia, JUSTICIA; esrepanza, esnebanza, ESPERANZA; deyasuno, dejavuno, DESAYUNO; coditiana, cobiliana, COTIDIANA; dispiclina, disgiblina, DISCIPLINA; oldivado, oltisado, OLVIDADO; desduno, desburo, DESNUDO; cladirad, clatinad, CLARIDAD; con vidin, concibin, CONDICION; sercivio, sernisio, SERVIo o CIO. Note that the nonword primes in Experiment 3 were the nonword targets in Experiment 4 (single-presentation lexical decision task).

245

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