Professional Documents
Culture Documents
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lnlm ilancc l!f Horns and Face Colour in Sheet) fi om IVood, 19u9 )
IVlim a /JII ll n! black-faced bieed 1 j is a ossed unth 1I hornedu.hite-faced breed
( 2) spcckled-faced horned males (3' and polled fe male: (1) are produced.
When the latter are bred together, among other iohite-faced polled (G}
nnd blark-horned (7j ani mals are produced
J. C. Ewart
7
first mated. Ewart repeated the experiment, but the quagga as a species
having become extinct, he employed a Burchell's zebra stallion, and this
served mares of several different brceds, and got striped hybrid foals.
The mares were afterwards put to stallions of their own breed, but the
resulting 'subsequent foals' never showed any sign of having been affected
by the pr vious zebra sire, Ewart carried out other telegony experiments
with various species of animals and they were all uniformly negative as
regards any evidence of the occurrence of the phenornenon. Accounts of
this work, together with the results of other breeding experiments, are
given by Ewart in his book on The Penycuik Experiments (1899), in his
presidential address to the Zoology Section of the British Association in
1903, and in numerous papers contributed to scientific and agricultural
journals.
In 1913 the University of Edinburgh rented a farm at Fairslacks,
Midlothian, and here Ewart kept a considerable number of sheep of
different breeds and started experiments designed to improve the fleece,
This work brought him into contact with the woollen industries of Great
Britain and they realized the importance of the work, In 1923 he went
to Australia at the invitation of the New South Wales Govcrnment and
visited many important and some very outIying sheep stations. He then
proceeded to New Zealand and conducted similar work there. There can
be no doubt that these investigations lcd to the further recognition of
the economic importance of scientific breeding, In 1924 the Worshipful
Company of Woolmen in London struck a gold medal, which they
presented to Ewart in recognition of what he had done.
WALTER HEAPE
In the meanwhile Walter Heape (1855-1929), whose name is mentioned
at the beginning of this s h o ~ t history, had formulated and conducted
a scheme of inquiry into certain other matters connectcd with sheep,
and more particularly those dealing with feeundity. The inquiry was
conducted under the auspices of the Evolution Committcc of the Royal
Society, ofwhich committee Heape was an original member (1896). The
investigation was carried out with the co-operation of the Royal Agri-
cultural Society, and a full report was published in 1899. It dealt chiefly
8 Tlu Application 01Science to Animal Breeding
with the incidence of abortion and barrenness in the different English
breeds, and the causes were shown to be different, depending both upon
breeding and feeding, and to vary with local conditions. The general
fertility of the breeds was also compared and many useful suggestions of
genetical value to farrners were put for ward. At a later date a somewhat
similar inquiry was carricd out in Scotland by the Highland and Agri-
cultural Society (MarshalI, 1908). Among thc practical conclusions which
resulted from these studies was one on the effects of 'flushing' shcep ; that
is, supplying thern with extra corn, cake or turnips or putting them upon
superior pasture or a good new ley at the approach of the tupping or
breeding season. Sheep brcedcrs had already found that these practices
generally increased the fecundity of the ewes, but the matter was not
clinched until it was shown that under the conditions of flushing the egg-
containing follicles in the ovaries ripened more rapidly, and that at the
periods of oestrus or heat two or more eggs would often be discharged
rather than one, the final result being an increase in the number of lambs
born. Another matter of interest concerned the inheritance of fertility.
The evidence collected showed that rarns whi ch were born as twins and
not singles could transmit the fertility of their dams to the next generation
of ewes, and consequently that the practice ofbreeding from singles rather
than twins was one to be deprecated, notwithstanding that the rams which
were single lambs were often better developed owing to their having
obtained more nourishment from their mothers in the early days of their
lives,
Heape also published an important papel' on artificial insemination
(1897). This practicc, although there is evidence that it may have been
known to the Arabs, was first elearly demonstrated by thc Italian biologist
Spallanzani (1784) in the successful insemination of a bitch. But it was
not takcn much notice of until Heape called attention to it by publishing
the experiments by Millais, in which by this method he effected crosses
between Blood-hounds and Basset-hounds (two breeds differing consider-
ably in size). Heape al so emphasized its probable importance to horse-
breeders, qu oting the reports of the Royal Commission on horse-breeding
in which it was shown that the percentage of sterility in thoroughbreds
in any one year might be as high as 40 per cent. One of the earliest
Walter Heape
9
successes in inseminating horses was when the thoroughbred mare Sandi-
way was successfully impregri ated by injecting semen obtained frorn the
stallion Trenton and afterwards produced a first-class foal named Sand-
flake. The more recent deveIopments and applications adopted in the
practice of artificial insemination are referred to below.
Heape's best known and probably his most important work was his
memoir on The Sexual Season oj Mammals (1900), in which he gave a
comparative account of the oestruaI or breeding cycles in all the different
animals for which any data existed. This was soon followed by fuller,
more detailed accounts by others on the oestrual cycles of the sheep, the
dog, the ferret, and the rabbit, and in recent years similar studies have
been made for a large number of other animals.
Heape's book on The Breeding Industry has been mentioned aboye in the
opening paragraph. In this book, which contains much statistical informa-
tion, he stressed the great economic importance of animal breeding in
Great Britain and the heavy losses which were annually incurred through
failure to apply scientific methods to animal production for commercial
purposes. In particular he stressed the importance of breeders keeping
records so that their experiences should not be lost but be made available
to the community.
GENETICS IN CAMBRIDGE-WILLIAM BATESON
Another Cambridge man who played an important part in promoting
and taking part in the study ofbreeding was Wilam Bateson (1861- 1926),
whose first book entit1ed Materials for the Study oj Variation, by its insistence
upon the phenomenon of a discontinuity in organisms, gave a fresh orienta-
tion to the way in which problems of reproduction and evolution were
regarded. Then in 1910, with the disinterment ofGregor Mendel's Versuche
ber Pflaneen-Hybriden (originally published in 1865), there appeared the
clue for which Bateson had been seeking, and he and his co-workers in
Cambridge made much of it in' working out the inheritance of many
kinds of animals and plants. In the meantime, Bateson brought out in
Cambridge an English edition of Mendel's work in the book entitled
Mendel' s Principies qf Heredity (lg0g).
The original experimenta of Mendel, as is well known, were upon