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and H
+
, so it is acidic
carcinogenic: causing cancer
cardiac arrest: a heart attack, when the heart muscle
stops contracting
cardiac cycle: the series of events that take place in the
heart during one heart beat
cartilage: a strong but exible tissue, found for example
covering the ends of bones at joints, and in the form of
C-shaped supporting rings in the trachea
Casparian strip: a band of suberin in the walls of cells
making up the endodermis of plant roots, preventing
water from passing between the cells
cell cycle: the sequence of events that takes place
from one cell division until the next; it is made up of
interphase, mitosis and cytokinesis
cell signalling: communication between one cell and
another by means of chemicals
cell surface membrane: the plasma membrane, made up
of a phospholipid bilayer in which protein molecules
oat
cell wall: a structure that is present outside the plasma
membrane in plant cells, fungal cells and bacterial cells;
in plants, it is made of criss-crossing layers of cellulose
bres
cellulose: a polysaccharide made of many -glucose
molecules linked together
centrioles: a pair of organelles found in animal cells,
which build the microtubules to form the spindle during
cell division
centromere: the place where two chromatids are held
together, and where the microtubules of the spindle
attach during cell division
channel protein: a protein that forms a pathway through
a membrane, through which molecules that would not
be able to move through the phospholipid bilayer can
pass
chloroplast: an organelle found only in plant cells,
where photosynthesis takes place; it is surrounded by an
envelope of two membranes, and contains chlorophyll;
photosynthesis takes place here
cholesterol: a lipid (or a lipid-like substance) that helps
to maintain the uidity of cell membranes
259
Glossary
chromatid: one of two identical parts of a chromosome,
held together by a centromere, formed during interphase
by the replication of the DNA molecule
chromatin: the DNA in a nucleus when the
chromosomes have not condensed
chromosome: a structure made of DNA and histones,
found in the nucleus of a eukaryotic cell; the term
bacterial chromosome is now commonly used for the
circular strand of DNA present in a prokaryotic cell
chronic illness: an illness that lasts a long time
chronic obstructive pulmonary disease (COPD): a long-
term illness caused by damage to the lungs and airways
ciliated cell: a cell that has cilia on its surface, which are
able to move in unison and cause movement
ciliated epithelium: a layer of cells covering a surface in
an animal, in which some of the cells have cilia
cisterna: a space enclosed by the membranes of the
endoplasmic reticulum
class: a group of similar orders
classication: putting things into groups
clone: a group of genetically identical cells
closed circulatory system: a circulatory system in which
the blood is always within blood vessels
coenzyme: a non-protein substance that is required for
an enzyme to catalyse a reaction
coenzyme A: a coenzyme required for many metabolic
reactions to take place, including respiration
cofactor: a non-protein substance, often an inorganic
ion, that is required for an enzyme to catalyse a reaction
cohesion: the attractive force that holds water molecules
together by hydrogen bonding
collagen: a brous protein found in skin, bones and
tendons
community: all the organisms of all the diferent species
living in the same place at the same time
companion cells: cells that are closely associated with
sieve elements; they have a nucleus and many organelles
and provide energy to enable translocation to take place
competitive inhibition: inhibition in which an enzyme
inhibitor has a similar shape to the substrate molecule,
and competes with it for the enzymes active site
complementary base pairing: the pattern of pairing
between the nitrogenous bases in a polynucleotide; in
DNA, A pairs with T and C with G
concentration gradient: a diference in concentration of
a substance between one area and another
condensation reaction: a reaction in which two
molecules are linked together, involving the formation
of water
condenser lens: the lens on a microscope that focuses
rays of light from the light source onto the specimen
continuous variation: variation in which there is a
continuous range of values between two extremes
contraction time: the time interval between phases Q
and T in an electrocardiogram
convergent evolution: the development of similar
structures in organisms that are not closely related, as a
result of adaptation for similar lifestyles
coronary artery: one of the arteries that branches of
from the aorta and supplies oxygenated blood to the
heart muscle
coronary heart disease (CHD): a disorder afecting the
blood vessels (coronary arteries) that supply the muscle
in the heart walls with blood
coronary thrombosis: a blood clot in a coronary artery
cortex: the area of a stem or root between the surface
layers and the centre; it is made up of parenchyma
tissue
covalent bond: a bond formed by two atoms sharing a
pair of electrons
crenated: the shape that red blood cells become when
they have lost a lot of water by osmosis
cyclic AMP: a second messenger substance, which is
afected when particular molecules bind to receptors in
a plasma membrane and which, in turn, afects another
molecule within the cell
cytosine: one of the four nitrogenous bases found in
DNA and RNA
cytoskeleton: a network of microtubules and
microbres that provides support in eukaryotic cells
denatured: when a protein molecule, for example an
enzyme, has lost its molecular shape, so that its function
can no longer be carried out
deoxyribonucleic acid (DNA): the genetic material
contained in chromosomes; a polynucleotide in which
the ve-carbon sugar is deoxyribose
deoxyribose: a ve-carbon sugar found in DNA
diaphragm: a dome-shaped sheet of tissue which
separates the thorax from the abdomen; contraction of
its muscles pulls it atter, which increases the volume
inside the thorax and helps to draw air into the lungs
diferentiation: the development of a cell to become
specialised for a particular function
difusion: the net movement, as a result of random
motion of its molecules or ions, of a substance from
an area of high concentration to an area of low
concentration
difusion gradient: a diference in concentration of
a substance between one place and another; if the
substance has particles that are free to move, their net
movement will be down the difusion gradient
diploid cell: a cell that possesses two complete sets of
chromosomes
disaccharide: a sugar whose molecules are made of two
sugar units
discontinuous variation: variation in which each
organism belongs to one of a few clearly dened groups
Glossary
260
disease: a condition that impairs the normal function
of the body
dissociate: to break apart; for example, carbonic
acid dissociates to form hydrogen ions and
hydrogencarbonate ions
dissociation curve: a graph showing the percentage
saturation of a blood pigment, for example
haemoglobin, at diferent concentrations of oxygen
dissolving: the separation and dispersal of molecules
or ions of a substance in between the molecules of a
solvent
disulde bond: a covalent bond formed between the
sulfur atoms of two cysteine molecules
domain: one of three major groups into which all living
organisms are classied; an alternative classication to
the ve-kingdom system
dopamine: a neurotransmitter found in the brain
double circulatory system: a circulatory system in which
there are two circuits, the systemic circulation and the
pulmonary circulation; the blood returns to the heart
after being oxygenated and before owing to other parts
of the body
ecosystem: the interactions between all the organisms,
and their environment, in a particular area
elastic bres: microscopic bres containing the protein
elastin, that are able to stretch and recoil back to their
original length
electrocardiogram (ECG): a recording of the hearts
electrical activity
emphysema: a disorder caused by the breakdown of
the alveolar walls, making it difcult to obtain sufcient
oxygen in the blood
endocytosis: the movement of bulk liquids or solids
into a cell, by the indentation of the plasma membrane
to form vesicles containing the substance; it is an active
process requiring ATP
endodermis: the outer layer of the stele in a plant root
endoplasmic reticulum: often abbreviated to ER; a
network of membranes within a eukaryotic cell, where
various metabolic reactions take place
endothelium: a tissue lining the inner surface of an
organ, for example the inside of a blood vessel
envelope: two membranes surrounding an organelle;
nuclei, mitochondria and chloroplasts have envelopes
environmental factor: a feature of the environment that
has an efect on an organism
environmental impact assessment: a study of possible
environmental efects that must be carried out before a
new development is approved
environmental variation: diferences between organisms
caused by their environment
enzyme: a protein that works as a catalyst
enzymesubstrate complex: the temporary association
between an enzyme and its substrate during an enzyme-
catalysed reaction
epidemiology: the study of patterns of disease in
populations, in order to work out the causes of the
disease
epithelial tissue: a layer of cells lining a surface in an
animal
equator: the mid-line of a cell, where the chromosomes
are lined up during metaphase of mitosis or meiosis
erythocyte: a red blood cell
ester bond: a bond linking a fatty acid to a glycerol
molecule in a lipid
eukaryotic: a cell containing a nucleus and other
membrane-bound organelles
ex situ: away from the place where an organism
normally lives
exocytosis: the movement of bulk liquids or solids out
of a cell, by the fusion of the vesicles containing the
substance with the plasma membrane; it is an active
process requiring ATP
experimental evidence: data obtained from planned,
controlled experiments
expiratory reserve volume: the maximum volume of air,
over and above the tidal volume, that can be breathed
out with one breath
extracellular: outside a cell
eyepiece lens: the part of a microscope through which
you look; it contains a lens that focuses light onto your
eye
facilitated difusion: the difusion of a substance
through protein channels in a cell membrane; the
proteins provide hydrophilic areas that allow the
molecules or ions to pass through a membrane that
would otherwise be less permeable to them; it is an
active process requiring ATP
family: a group of similar genera
fatty acid: a molecule containing a hydrocarbon chain,
with a carboxyl group at one end
bre: a grouping of brils of a substance, for example
in collagen or cellulose; bres are generally large enough
to be seen with a light microscope, whereas brils are of
an order of magnitude smaller
bril: a grouping of several molecules held side-by-side
by bonds between them, for example in collagen or
cellulose
brous protein: a protein with a linear three-
dimensional shape; brous proteins have structural roles
and are insoluble in water
lling time: the stage in the cardiac cycle during which
the heart is lling with blood
accid: a plant cell is said to be accid if it has lost a lot
of water, and its volume has shrunk so much that the
cell no longer pushes outwards on the cell wall
261
uid mosaic model: the universally accepted model
of membrane structure, in which proteins oat in a
phospholipid bilayer
fossil: the remains of a long-dead organism, preserved
in stone
freeze-dried: (food) frozen in a vacuum, so that most of
the water in it is lost as water vapour
gamma radiation: short wavelength electromagnetic
radiation that destroys living cells
gene: a sequence of DNA nucleotides that codes for a
polypeptide
genetic variation: diferences between organisms caused
by their genes
genetically diferent: containing a diferent collection of
alleles of genes
genus: a group of similar species; plural genera
geographical isolation: the separation of two
populations of a species by a geographical barrier, for
example a stretch of water
globular protein: a protein with a roughly spherical
three-dimensional shape; many are metabolically active
and soluble in water
glucose: a hexose sugar; the form in which carbohydrate
is transported in the blood of mammals
glycogen: a polysaccharide made of many alpha
glucose molecules linked together; it is the storage
polysaccharide in animals and fungi
glycogen phosphorylase: an enzyme that catalyses the
breakdown of glycogen to form glucose, especially in
liver cells and muscle cells
glycolipid: a molecule made of a lipid to which sugars
are attached
glycoprotein: a molecule made of a protein to which
sugars are attached
glycosidic bond: the bond that links sugar molecules
together in a disaccharide or polysaccharide
goblet cell: an epithelial cell that secretes mucus
Golgi apparatus: a stack of curved membranes inside
a cell, in which protein molecules are packaged and
modied, for example by adding sugars to them to
produce glycoproteins
G-protein: a protein that is linked to a receptor in a
plasma membrane, and that brings about a reaction
when a signal molecule links with the receptor
grana: (singular: granum) membranes inside a
chloroplast, containing chlorophyll
guanine: one of the four nitrogenous bases found in
DNA and RNA
habitat: a place where an organism lives
haem: a group of atoms with an Fe
2+
ion at the centre,
found in several proteins including haemoglobin and
cytochrome
haemocoel: a space within the body of an animal, for
example an insect, in which blood is contained
haemoglobin: the red pigment found in red blood cells;
it transports oxygen and carbon dioxide between lungs
and body cells
haemoglobinic acid: the compound formed when
hydrogen ions combine with haemoglobin
haploid cell: a cell that possesses only one set of
chromosomes
HDL: high-density lipoprotein
health: a state of complete physical, mental and social
well-being
herd immunity: immunity possessed by a large
proportion of individuals in a population, lowering the
chances of even those individuals that are not immune
from getting an infectious disease
heterotroph: an organism that requires organic
nutrients; all animals and fungi are heterotrophs
hexose: a sugar with six carbon atoms, C
6
H
12
O
6
hierarchy: a series of groups in which each group is
made up of several smaller ones
histones: protein molecules that are associated with
DNA in chromosomes
homologous chromosomes: a pair of chromosomes in a
diploid cell that have the same structure as each other,
with the same genes (but not necessarily the same alleles
of those genes) at the same loci; they that pair together
to form a bivalent during the rst division of meiosis
humidity: the quantity of water vapour held in the air
hydrogen bond: an attractive force between a slight
negative charge on one atom, for example oxygen, and a
slight positive charge on another, for example hydrogen
hydrolase: an enzyme that catalyses a hydrolysis
reaction
hydrolysis: a reaction in which two molecules are
separated from each other, involving the combination of
water
hydrophilic: attracted to water; hydrophilic substances
have small electric charges that are attracted to the small
electric charges (dipoles) on water molecule
hydrophobic interaction: a weak bond formed between
two groups of atoms that are repelled by water
hydrophobic: repelled by water; hydrophobic substances
do not have electric charges, and are not attracted to
water molecules
hypertension: high blood pressure
immune: able to destroy a pathogen before it can harm
the body
immune response: the way in which lymphocytes
respond to infection by pathogens
immune system: the organs and cells in the body that
help to destroy pathogens
immunoglobulin: an antibody; a small protein molecule
that is able to bind with a specic antigen
in situ: in the place where an organism normally lives
Glossary
Glossary
262
induced t: the change of shape of an enzyme when
it binds with its substrate, caused by contact with the
substrate
inhibitor: a substance that reduces the rate of an
enzyme-catalysed reaction
initial rate of reaction: the rate at which substrate
is converted to product right at the beginning of a
reaction
inorganic ions: charged particles of elements or their
compounds, such as potassium, K
+
and nitrate, NO
3