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Abstract
While a number of models have been developed to assist managers of deciduous fruit tree
crops with specic aspects of decision making, most are non-optimising predictive models and
few employ detailed mechanistic models of fruit-tree growth that would enable the simulation
of any orchard system from planting to maturity. This paper details the complex biological
and economic relationships present in an apple orchard system and describes a dynamic
simulation model based on these interactions. The model is bioeconomic in nature, and may
be used to investigate a range of issues of relevance to the commercial apple orchardist. These
issues include understanding how biological factors inuence apple-tree productivity, and
how to choose among a diverse range of apple orchard systems. Each system, consisting of a
particular combination of cultivar, rootstock, tree spacing and training method, has implications for fruit quality, quantity and ultimately prot. The choice of system is made at planting, while an important annual decision is the optimal rate of thinning, both of which
determine potential yield over the lifetime of the orchard. These decisions also inuence costs
and revenues per hectare and, by necessity, are made in the context of unknown future prices
of inputs and outputs. The bioeconomic model is used to maximise net present value of one
orchard system by selecting optimal thinning strategies over a 15-year period.
# 2003 Elsevier Science Ltd. All rights reserved.
Keywords: Horticulture; Bioeconomics; Modelling; Thinning; Apples
1. Introduction
Orchard systems are complex. Managers of deciduous perennial fruit crops must
consider both biological and economic relationships in determining preferred orchard
138
design and life-time orchard management strategies. Decisions about tree size and
varietal mix made at planting determine potential yield over the lifetime of the
orchard. These decisions also inuence costs and prot per hectare and, by necessity,
are made in the context of unknown future prices of inputs and outputs. Once trees
are bearing fruit, the decision concerning the amount of fruit to leave on the tree
until harvest (the thinning decision) remains one of the few ways of inuencing the
annual orchard yield.
A simulation model representing the apple orchard is a particularly useful means
of evaluating the eects on yield and protability of alternative orchard systems and
other decisions under the direct control of the orchardist. Given the importance of
biological and economic components in orchard protability, both elements should
feature in the model if the orchard system is to be adequately described and simulations meaningful. Lack of data on yields of many important apple varieties overtime means it is necessary to adopt a bioeconomic modelling approach.
Simulation models of agricultural systems have grown in popularity in recent
decades due to their usefulness in tackling the inherent dynamic and/or stochastic
nature of agricultural problems and due to increased computer capacity (Oriade and
Dillon, 1997). Simulation may substitute for large-scale physical experimentation,
which could otherwise take decades, especially in the case of perennial crops.
The biological simulation models that underlie bioeconomic models may vary considerably in their complexity and depend critically on the purpose for which they are
constructed. The biological simulation models developed by biologists and agricultural scientists are often too detailed to be used in optimising economic models
designed by economists. Conversely, simulation models developed by economists tend
to be too simple by biologists standards with models consisting of linear or simple
non-linear functions to allow for solution of optimisation problems (Cacho, 1997).
In this paper, an attempt was made to nd a balance between the complexity
demanded by biologists and the simplicity required in dynamic optimisation problems. The paper starts by presenting a review of past modelling work in perennial
fruit trees and identifying further research needs. A brief description of fruit tree
management is then presented, followed by the development of a bioeconomic
model for an apple orchard. The model is used to understand the inuence of thinning decisions on apple yields and prices and, consequently, on the net present value
(NPV) of the operation. The model is then incorporated into a dynamic optimisation algorithm to determine the optimal thinning path over a period of 15 years,
starting from orchard establishment. The paper ends with concluding comments and
future research needs.
139
140
Table 1
A chronological sample of published tree crop models
Tree type
Biological model
Optimisation/analysis
techniques
Decision variables
Objectives
Apple
Apple,
pear
Apple,
cherry,
pear
Apple
Dynamic linear
programming
Simulation
Survey-generated
yield information
Mechanistic,
(FRUPRO)
Survey-generated
yield information
Apple
Empirical yield
estimation
Mechanistic
Peach
Apple
Empirical
Empirical
Groot (1996)
Empirical
Apple,
pear
Apple
Smulation/dynamic
programming
Inference procedures,
simulation
Simulation
Dnamic simulation
model
Simulation
Empirical
Smulation
Apple
Yield estimation
Cost-benet analysis,
simulation
Single/multiple period
linear programming
Author(s)
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(1987) denes an orchard system as the integration of all the horticultural factors
involved in establishing and maintaining a planting of fruit trees. Many decisions
must be made before planting occurs, including choice of cultivar (variety), rootstock (tree size), tree density and pruning and training (tree form).
The apple cultivar determines fruit variety and fruit features such as size, shape,
colour, avour, rmness, ripening season and pest and disease resistance. The age at
which an apple tree rst bears fruit (its precocity) will vary according to the cultivar/
rootstock combination.
A range of apple cultivars are normally grown in commercial apple orchards. A
variety of reasons exist for choosing a particular cultivar: each type of apple may
face distinct market situations and may potentially receive dierent prices; sequential harvest dates allow for more ecient picking and packing operations; some
cultivars have a greater susceptibility to pests and diseases than others; and dierent
cultivars may be necessary for pollination.
Tree size is central to an orchard system because of its economic implications.
Trees that are large at maturity result in increased labour costs and often take many
years to bear fruit. An objective of modern orchard design is to maintain smaller
trees that are planted closer together. These smaller trees have the additional
advantage of earlier bearing habits. Tree size is most often controlled physiologically
through rootstock selection.
Horticultural techniques used to manage vegetative growth and fruit production
are pruning, training and thinning. While the level of thinning is determined each
year, pruning and training are usually predetermined by choice of density, rootstock
and cultivar when the orchard is established. These latter choices are made at
planting and assumed to remain unchanged during the life of the orchard.
4. The model
4.1. Conceptual biophysical model
In this paper, apple tree growth during a growing season and over a lifetime is
described using a carbon-balance model. Seasonal growth patterns that are peculiar
to deciduous woody perennials, such as apple trees, form an integral part of the
model. The carbon-balance model is described in detail in Hester (2000) and only a
brief overview is given here.
Fig. 1 shows the conceptual biophysical model. The critical environmental variables in the model are day length, temperature and lightall three aect the growth
potential of the tree. Light (radiation) and daylength aect canopy photosynthesis,
while temperature aects both photosynthesis and respiration. Light interception is
critical to the amount of photosynthesis undertaken by an apple tree, and the level
of interception is determined by shape of the tree which, in turn, is inuenced by
chosen pruning and training techniques and tree age. Net photosynthesis describes
the amount of carbon available for tree growth after accounting for carbon lost
during leaf respiration and gained through photosynthesis.
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Photosynthesis provides energy used for growth by each tree component: roots;
wood; fruit; and leaves. Respiration of these components results in energy losses.
The fruit load on the tree is the major determinant of how energy is divided
between each tree component and is inuenced through thinning.
Fruit load also inuences size of individual fruits and has an impact on future fruit
production through its eect on the biomass of root, wood and leaf that grows in
the current and following seasons. Fruit is harvested annually and its quantity and
quality is taken into account by the model.
Apple trees follow temperature-driven patterns of growth and non-growth over a
12-month period. During late autumn, apple trees enter a dormant period where
vegetative and reproductive buds require a period of chilling temperatures if bloom,
growth and development are to occur in the spring. Dormancy is broken once the
chilling requirement of the particular apple cultivar has been satised. Growth,
however, does not automatically resume at this point, rather, it occurs following
higher daily temperatures that result in accumulation of a certain number of heat
units. These events are accounted for in the model by a set of triggers associated
with heat and chill unit accumulation.
4.2. The bioeconomic model
The economic model describes the costs and revenues associated with fruit production in an orchard system from planting to maturity. Annual fruit production is
143
determined by the biophysical model and used in the economic model to simulate
the annual protability and net present value (NPV) of a particular system.
The relationship between the biophysical and economic models is described in
Fig. 2. The yield and NPV of each orchard system will depend fundamentally on the
variety, tree density and pruning/training regimes chosen when trees were planted.
In addition, yield and protability may be signicantly modied each year through
changing the amount of thinning undertaken. Thinning is used to modify the number of fruits a tree bears to maturity and thus it determines their size and inuences
the price they receive. The length of the time horizon, T, also inuences system NPV
and can be included as a decision variable. The time horizon is important in assessing orchard rotation strategies, however, is not considered as a decision variable in
the current model.
The decision variables can be viewed as being of two types. Those pertaining to
the establishment of the orchard, and those management decisions made annually.
Thinning falls within the latter category.
Annual yield is used in the economic model to determine annual revenue from the
system. Apple prices are given exogenously. The cost of establishing the orchard
system as well as costs related to tree density and harvest enter the NPV calculation.
When the model is used in optimising mode, decision variables are adjusted so that
NPV is maximised.
The main features of the model are detailed more formally in this section. The net
present value (V) of the stream of annual prots obtained over the planning horizon
t=1,. . .,T is dened as:
144
T
X
t Ft fxt ; ut g1 rt CE
t1
where the annual prot t is a function of fruit production per hectare (Ft), r is the
discount rate and CE is the cost of establishing the orchard starting with bare
ground. Fruit production is, in turn, determined by a state vector xt and a control
variable ut representing thinning. The variable ut is inversely proportional to thinning intensity, it takes on a value of zero when thinning is 100%, so that no fruits
remain on the tree, and it takes on a value of one when no thinning occurs, so that
all fruits remain on the tree. Values of ut between zero and one indicate intermediate
levels of thinning. In the economic model, time (t) is a discrete variable measured in
years. Annual prot is described as:
t Pa fFWt g CFFt fxt ; ut g NCNt fAt g
where Pa is the price of fruit (A$/kg) which is a function of average fruit weight
(FWt ) and measured in grams of dry matter, CF is the cost of harvesting the fruit
(A$/kg), N is the planting density (trees/ha), and CNt represents costs per tree.
Labour is the main component of harvesting cost (CF). Cost per kilogram harvested
depends on variety and orchard system used, because small, compact trees require
less labour per kilogram harvested than large trees. Density-related costs (CNt)
depend on age of the orchard (At) and consist of labour, materials, chemicals, fertiliser and other expenses, such as irrigation costs per tree. These costs are assumed to
increase at a decreasing rate, hence:
dCN
> 0;
dAt
and
d 2 CN
40
dAt2
The choice of orchard system determines pruning and training requirements and
ultimately aects the shape of the trees which, in turn, may have a bearing on time
and eort required to apply chemicals, fertilisers and irrigation. In summary, all
costs (CE, CF and CNt) depend on the orchard system selected. The model represents a single orchard cycle starting with bare ground, thus At=t.
Annual fruit production (kg/ha) is determined by photosynthesis and respiration
rates during the year:
t1
Ft N
G L C
F; xt ; ut d
t
145
production and growth. The parameter converts carbon to fresh fruit weight. Thus
the integral represents fruit production per tree in the interval from the beginning to
the end of year t, and this is multiplied by tree density (N) to obtain yield per hectare.
The biophysical model consists of four state variables: leaf, fruit, wood and root.
Each of these variables is associated with a dierential equation for daily gain in
mass of each component. Dierential equations are numerically integrated on a
daily basis in the simulation. From the standpoint of the economic model, leaf and
fruit are not treated as state variables since they follow an annual cycle that, for a
given orchard system and climatic events, depends on the amount of carbohydrate
reserves available in wood and root at the beginning of the season. Thus, the state
variable vector for each year is dened as:
xt Rt ; Wt ; Dt
The state of an individual tree at the beginning of year t is described using dry
matter content in root (R) and wood (W) and the presence of pests and diseases (D).
The annual objective of thinning, to achieve the most protable fruit size, is tempered by the eect of current fruit load on potential fruit size and yield in future
years. If a tree bearing a large number of apples is thinned only lightly, it is probable
that the tree will only be able to support a small number of apples in the following
year, which may negatively impact on prots. This tendency towards biennial bearing is inherent in many apple cultivars and may be modied by thinning. The orchard manager must therefore take into account both immediate and long-term
fruiting prospects of the tree when determining the appropriate level of annual
thinning to maximise NPV.
Following picking, apples are graded and packed into various count sizes that
are based on individual fruit weight and reect number of fruit per carton. The value
of the apple harvest is closely related to fruit size. Generally a crop of small apples is
worth less than the same weight of larger apples, hence the incentive to improve fruit
size through thinning.
In the model, a biennial bearing pattern is imposed on fruit growth that sees a
large crop followed by a light crop. Data on natural fruit bearing patterns is scarce
although the biennial bearing pattern simulated by the biophysical model does
compare reasonably with the fruit mass data from an Australian trial (Middleton,
1984; Middleton, unpublished data). The biennial bearing pattern is simulated as
follows:
EFLt BFLFLt1 BFL
where EFLt is expected fruit load when no thinning takes place, BFL is fruit load
that would lead to minimal or no biennial bearing and FLt is fruit load in the previous year. Fruit load is dened as the number of apples per m2 of leaf area. The
value of EFLt depends on the amount by which actual fruit load exceeded BFL in
the previous year. The estimate of BFL is set at a level that appears reasonable given
data from an Australian trial (Middleton, 1984; Middleton, unpublished data).
146
The process of thinning leaves a proportion of the original amount of fruit on the
tree. The actual fruit load that results from thinning (FLt), is calculated as:
FLt EFLt ut 0 4 ut 4 1
The energy for growth produced as a result of photosynthesis, in the form of dry
matter, is allocated to fruit, leaves, wood and root components of an apple tree. This
allocation process is known as partitioning, and is aected by whether or not a tree
is producing fruit. One noticeable eect of the presence of fruit is a reduction in dry
matter partitioned to root and hence a reduction in tree growth (Heim et al., 1979).
A heavy fruit load (FL), dened as fruit number per m2 leaf area, reduces photosynthesis through a reduction in leaf area (shoot growth) when compared with trees
having no fruit (Faust, 1989).
Heim et al. (1979) found that levels of partitioning of dry matter to various tree
components were caused almost entirely by dierences in fruit load. Using data
from Heim et al. (1979), the proportion of assimilates partitioned to tree components during a growing season was estimated in the model using the following
equations:
j j
lj FLt
j FLt
R 1 L W F
for j L; W; F
7
8
where j is the proportion of dry matter partitioned to the various tree components
and lj ; j and j are constants.
The pattern of partitioning is important in perennial tree crops such as apples,
because the relative dry matter allocations to tree parts in the current year have
implications for growth and fruiting potential in following years. At the extreme, a
tree may not bear a regular crop every year but instead a heavy crop may be followed by a small crop because of fruit overload in a particular year (Monselise and
Goldschmidt, 1982). Inuencing the partitioning process in an apple tree through
the adjustment of fruit quantity may correct these biennial bearing patterns if a
cultivar is prone to them. One important outcome that normally results from lowering fruit load of a tree is improvement in average fruit size.
Once FLt is determined, fruit number (FN) is calculated according to:
FNt FLt : LAmaxt
where LAmaxt is the maximum leaf area (m2 leaf area), calculated from the biophysical
model. Thus the units for FN will be a number, representing the total number of
apples on a particular apple tree. Finally, average fruit weight (FWt) is calculated as:
FWt
YF t
FNt
10
147
where YFt is the yield per tree measured in grams of fresh weight, calculated in the
biophysical model. Average fruit weight is used in the economic model to determine
the protability of a given thinning strategy.
5. Model implementation
In this section, the protability of one hectare of Granny Smith apple trees is
simulated over a planning horizon of 15 years from planting. The trees are planted
on the semi-dwarng MM.106 rootstock, at a density of 1000 trees per hectare and
trained to the central leader system. The specic system chosen for simulation is one
of those outlined in Hester (2000).
Prices received by the apple orchardist are determined exogenously. For a given
variety, apple prices vary according to individual fruit size. Apple size is approximated by the fruit count or number of apples in an 18-kg carton. The higher the
count the smaller the apple size (weight) and vice versa. Larger sizes receive a price
premium; however, enormous apples receive a price penalty.
Data on prices received for each grade of apple variety are not regularly recorded
at fruit selling centres in Australia, rather, only maximum, minimum and average
prices for a variety are publicly available. Detailed price by grade data was obtained
for Granny Smith apples, sold during January 1999 and was used to estimate the
grade price dierentials associated with fruit size for all varieties.
For simplicity, it is assumed that the relative price dierences shown in Table 2
hold regardless of maximum price. The data were used to estimate the following
price-count relationships used in the economic model:
8
P75
if countt 4 75
<
Pt
Pmax
if 75< countt <105
11
:
c 0:2448countt if countt 5 105
Table 2
Price data used to develop price-count relationship
Count
70
80
90
100
110
120
135
150
165
180
198
36
43
43
43
38
31
25
24
21
16
15
148
where P75 is the price for counts less than 75, Pmax is the price received for counts
between 75 and 105 and c is an intercept term. Using the Batlow data P75 has a
value of A$36 per carton, Pmax is A$43 per carton and c is 61.29.
Costs related to tree density (CN) are presented in Fig. 3 and are specic to the
chosen system. Labour costs peak in year four after planting when pruning and
training requirements are highest. After this time, labour requirements remain
stable. Chemical and nutrient costs are lowest during the years immediately after
planting but increase as the tree reaches maturity. The peaks in nutrient costs
represent the cycle of applications that is adopted in this orchard. Capital costs for
training are minimal in the central leader system and are incurred in the 2 years
immediately after planting.
Establishment costs are estimated to be A$8770 per hectare for this orchard system, assuming a cost of A$6.25 per tree purchased from a commercial nursery.
The biophysical model, described by Eq. (3) is solved using numerical integration
at daily steps and is implemented using the specialised simulation packages Simulink1 and Matlab1 (Mathworks, 1996, 1997). The biophysical model is solved for
specic values of thinning and the resulting fruit quantities and sizes are used in the
economic model to calculate annual prot and NPV.
149
Fig. 4. The eect of thinning target (u) on average fruit weight (a), price (b), yield (c) and NPV (d).
150
The decrease in average fruit weight has implications for the average price
received (Fig. 4b). When fruit is thinned so that only a small number of fruits remain
on the tree (u=0.05) average price is slightly above A$2.00 per kilogram. At this
level of thinning average fruit weight, and hence size, is greatest. Highest prices are
not received here because the largest sized apples (count > 75) do not receive the
highest price per kilogram. Fruit price does increase as more fruit are left on the tree
and a drop in average weight (size) improves their value. Apples receive the highest
price per kilogram when thinning leaves between 20 and 25% of fruit on the tree
(0.24u 40.25). Beyond this level of thinning average fruit price decreases in line
with reductions in fruit size that put the fruit in lower-valued count sizes.
While leaving progressively more fruit on the tree does reduce average fruit
weight, it also has the eect of increasing total fruit weight on the tree, or yield
(Fig. 4c). As thinning increases, yield per tree also increases, but at a decreasing rate.
When thinning leaves only around ve per cent of fruit on each tree (u=0.05), total
yield is approximately 20 kg per tree, and increases to around 75 kg of fruit per tree
when thinning leaves around 55% of fruit on the tree (u =0.55). However, at high
yields, average fruit size is small and average price per kilogram of fruit is well below
the maximum.
Ultimately, the value of thinning hinges on its eect on NPV. As u increases from
0.05 to 0.45, NPV increases (Fig. 4d) to a maximum of A$1,210,102 and decreases
thereafter. Prior to the level of thinning that produces maximum NPV (u=0.45), the
increased yield per tree compensates for the reduced price per kilogram received for
the progressively smaller fruit. However, beyond u=0.45 average fruit size is
reduced to a level that receives dramatically lower prices, and the increase in average
yields per tree does not compensate for this price drop.
It is interesting to note, therefore, that maximum NPV does not occur where
prices are at a maximum. Nor does it occur where fruit yield or fruit weight are at
a maximum. Rather, the trade o between total yield per tree and individual fruit
weight results in the achievement of maximum NPV where prices are below the
maximum level.
This simple analysis provides insight into the operation of the model and the eect
of thinning. However, these results only apply when thinning is treated as a static
variable, taking the same value every year.
6.2. Dynamic optimisation
Initial attempts at solving the dynamic model with a nonlinear programming
(NLP) algorithm, showed that the solution was highly sensitive to the starting point
for the search. This suggests that the model tends to converge to a local optimum,
this problem is described by Cacho (1998). To overcome this problem, a genetic
algorithm (GA) was combined with the NLP in an attempt to identify the global
maximum. The GA was adapted from that reported by Cacho and Simmons (1999).
The GA was rst used to undertake a global search of the decision space and, once
the GA converged, the NLP was activated using the GA solution as the starting
point. This approach yielded a higher NPV than any solution of the NLP on its
151
own. Although there is no guarantee that the global optimum was found, the technique increased our condence that the solution was close enough to the best possible for practical purposes.
The optimal thinning trajectory is shown in Fig. 5. This trajectory gives a cumulative fruit yield of 967 kg/tree over the 15-year period with an NPV of A$1,265,000/
ha over the same time horizon.
The optimal trajectories of fruit weight, price, yield and prot are presented in
Fig. 6. Average fruit weight remains constant after year four (Fig. 6a) and maximum
prices per kg are received in all but the second year from planting (Fig. 6b). Average
fruit yield per tree (Fig. 6c) rises consistently over time, although the increases in
years 24 are slower reecting the thinning regime during this time, when relatively
large amounts of fruit were left on the tree. Annual discounted prots (Fig. 6d) are
aected by fruit prices resulting from the optimal thinning strategy. Before the trees
begin to bear fruit, initial establishment costs and variable costs result in a negative
prot. Prot continues to rise until year 10 when it levels out and remains reasonably constant for the remainder of the planning horizon.
Optimal fruit yield and fruit load results over 15 years are compared with a
situation of no thinning in Fig. 7. When no thinning takes place, the biennial bearing pattern that results from the model shows a very clear and consistent pattern of
alternation between the o year and the on year of fruit production. In the on
years, yield increases from 18 kg per tree in year two to around 200 kg per tree in
year 14. In the o years, no fruit is produced due to the eect on the tree of the
high yields in the previous year (Fig. 7a). This contrasts with fruit production per
Fig. 5. Optimal thinning targets (u*) using an NLP (dotted line) and the GA-NLP algorithm (solid line).
152
Fig. 6. Optimal trajectories for fruit weight (a), fruit price (b), fruit yield (c) and discounted prot (d)
under an NLP (dotted lines) and the GA-NLP algorithm (solid lines).
Fig. 7. Optimal trajectories (solid lines) for fruit yield (a) and fruit load (b) compared with the no-thinning option (dotted line).
tree using the optimal thinning strategy, where the cycle of dramatic uctuation is
eliminated. When no thinning occurs, the on years produce a fruit load of 20 fruits
per m2 of leaf area, and a fruit load of zero in the following year (Fig. 7b). The
optimal thinning strategy modies these dramatic uctuations and fruit load
appears to settle at a value of six fruits per m2 of leaf area. Small uctuations occur
in the earlier years but are reduced to almost no variation by year ten.
153
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