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Landscape ecology, land-use structure, and population density: Case study of the
Columbus Metropolitan Area
Jia Lu a, , Jean-Michel Guldmann b
a
b
Geosciences Program, Valdosta State University, 1500 N. Patterson St, Valdosta, GA 31698, USA
City and Regional Planning, Ohio State University, 275 West Woodruff Ave, Columbus, OH 43210, USA
a r t i c l e
i n f o
Article history:
Received 17 February 2011
Received in revised form
21 November 2011
Accepted 30 November 2011
Available online 3 February 2012
Keywords:
Population density
Urban modeling
Landscape ecology
Land use
GIS
Quantitative analysis
a b s t r a c t
Traditional population density models based on the distance to the major Central Business District (monocentric) or on distances to multiple employment centers (polycentric) are extended to include land-use
structure variables derived from landscape ecology theory. A comprehensive database is developed for
the Columbus Metropolitan Area (CMA) at the Trafc Analysis Zone (TAZ) level, using remotely sensed
land-use data, Census socio-economic data, and other local data. Fifteen landscape indices, organized into
four groups size, complexity, diversity, neighborhood are computed for each of the 1763 CMA TAZs,
using Fragstats. Models are estimated for each of the 7 CMA counties separately, yielding homogeneous
and consistent results. These county models are then pooled into a comprehensive CMA model, with
dummy variables and second-order terms. Overall, the results provide evidence of a polycentric structure, with both downtown Columbus and county CBDs acting as strong population attractors, and of the
importance of land-use structure in the determination of population density. Spatial indices representing
neighborhood and diversity factors signicantly impact population density in most counties.
2012 Elsevier B.V. All rights reserved.
1. Introduction
The spatial distribution of a citys population reects economic,
social, technological, and market conditions, as well as planning
policies (e.g., infrastructure expansion), with signicant impacts on
its social and environmental functionalities. Models of population
distribution and density are therefore central to forecasting urban
futures, and of great signicance to planners and policy makers. Not
surprisingly, population density patterns have been studied extensively. Early research has focused on monocentric models, with
distance to the Central Business District (CBD) as the major determinant of population density. The negative exponential model
with a constant gradient, as pioneered by Clark (1951), became the
standard theory (McDonald, 1989), under the assumption of a
long-run equilibrium between utility-maximizing consumers and
prot-maximizing suppliers of housing, with all jobs located in
the CBD. Newling (1969) proposed an alternative model by adding
a quadratic term in the exponential equation, and addressed the
existence of a density peak away from the city center. A review of
these monocentric population models is reported in Smith (1997).
Starting in the 1980s, this model gradually evolved into a polycentric one to account for increasing suburbanization, the decline of
Corresponding author. Tel.: +1 229 333 5752; fax: +1 229 219 1201.
E-mail addresses: jlu@valdosta.edu (J. Lu), guldmann.1@osu.edu
(J.-M. Guldmann).
0169-2046/$ see front matter 2012 Elsevier B.V. All rights reserved.
doi:10.1016/j.landurbplan.2011.11.024
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
75
76
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
the area among patch types becomes more even. For a given number of land-use types, SHDI is maximized when all land uses have
the same area. The Shannons evenness index (SHEI) is equal to
SHDI divided by logarithm of the number of land-use types. SHEI
ranges between 0 and 1. SHEI = 0 when the landscape contains only
1 patch (no diversity) and approaches 1 as the distribution of the
area among different patch types becomes increasingly even. The
Dominance Index (DO) measures the magnitude of the dominance
of one or a few land uses over the others in a given area. This index
has been developed by ONeill et al. (1988), based on information
theory. Large values of DO indicate that the area is dominated by
very few land-uses, whereas small values suggest a relatively even
distribution of land-uses.
The assumption of this study is that the higher NP, RPR, SHDI,
and SHEI, and the lower DO, the more land-use mixing, probably the
more competition for land, and hence the more restrictions on land
development. These restrictions imply a higher development cost,
and, therefore, a higher development density to offset development
costs.
2.1.4. Neighborhood indices
The Mean Edge Contrast Index (MECI) represents the mean
contrast between a given polygon and all polygons within a userspecied neighborhood, based on contrast weights assigned to the
edge lines between polygons. The average of all contrast values of
all edge lines is MECI. MECI = 0 if the landscape consists of only 1
polygon or the boundary contains no edge. In addition, MECI = 0
when all polygon perimeter segments involve polygon adjacencies
that have been given a zero-contrast weight. MECI = 100 when the
entire polygon perimeter is the maximum-contrast edge (w = 1).
The Mean Nearest-Neighbor Distance (MNN) refers to the distances
between polygons with the same land-use type. MNN equals the
average distance to the nearest neighboring polygon of the same
type, based on shortest edge-to-edge distance. MNN can be 0 if
there is no nearest neighbor. Polygons must be non-adjacent to be
included in the search for the nearest neighbor.
The Interspersion and Juxtaposition Index (IJI) is the only measure that explicitly accounts for the spatial conguration of patch
types. Each patch is analyzed for adjacency with all other patch
types, and IJI measures the extent to which patch types are interspersed, that is, equally bordering other patch types. IJI represents
the ratio of observed interspersion over maximum possible interspersion for a given number of land-use types, and ranges between
0 and 100. IJI approaches 0 when the distribution of adjacencies among unique land-use types becomes increasingly uneven.
IJI = 100 when all land-use types are equally adjacent to all other
land-use types (that is, maximum interspersion and juxtaposition).
The assumption of this study is that the higher MECI, MNN,
and IJI, the more fragmented and complex the area in terms of
land-use mix, probably more competing demands for land, and the
more restrictions on land development. Such restrictions imply a
higher cost of development, and, therefore, a higher probability for
a higher development density to offset higher development costs.
2.2. Applications of landscape indices
2.2.1. Natural environment
Kareiva (1990) shows that theoretical ecological models (island,
stepping-stone, continuum) make it clear that the spatial subdivision of habitat can alter the stability of species interactions and
opportunities for coexistence, but suggests that the empirical challenge is to investigate more rigorously the role of spatial subdivision
and dispersal in animal and vegetal communities. The following is
a short overview of such empirical studies.
Miller, Brooks, and Croonquist (1997) nd that several indicators of diversity and contagion are most effective in explaining the
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
(1)
77
(2)
(3)
As discussed in Section 2.1, land-use structure, measured by spatial indices, is likely to have an impact on population density. Let I
be the vector representing these indices. A generalized population
density function is then:
DP = F(D, X, I)
(4)
78
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
Fig. 1. Population density in the Columbus Metropolitan Area (person/million ft2 ; 1 ft2 = 0.09290304 m2 ).
Source: CTPP (1990).
website. The source is leaves-off Landsat TM data, nominal1992 acquisitions, with a 30-m precision. The NLCD includes
22 land-use types, 14 of which actually exist in the CMA, as
illustrated in Fig. 2. All of the 15 spatial indices described
in Section 2, except the Dominance Index DO, have been
Fig. 2. Land use in the Columbus Metropolitan Area (NLCD) (1 mile = 1.609344 km).
Source: NLCD (1992).
Orchards/
vineyard
0
2
1
3
0
1
2
2
1
Transitional Deciduous/evergreen/
mixed forest
Shrub
land
0
1
1
Grassland/
herbaceous
0
1
Cultivated Wetland
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
2
2
1
2
1
1
1
1
1
2
3
4
3
4
5
0
0
1
0
2
4
Bare rock
Quarried/strip
mines/
gravel pits
2
2
2
2
4
2
2
2
2
3
2
3
2
3
1
3
3
3
3
4
3
2
2
1
2
2
3
3
3
3
4
1
2
2
4
5
4
3
3
3
5
3
2
3
0
2
0
3
4
Open water
Low intensity residential
High intensity
residential
Commercial/industrial/
transportation
Quarried/strip
mines/gravel pits
Bare rock
Transitional
Deciduous/evergreen/
mixed forest
Shrub land
Orchards/vineyard
Grassland/herbaceous
Cultivated
Wetland
Low intensity
residential
Commercial/industrial/
transportation
ln(DP) = a + b DCOL
Open
water
High intensity
residential
6. Empirical results
Land use
Table 1
Weights for the computation of MECI.
79
(5)
80
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
81
Table 2
Pearson correlation coefcients of spatial indices.
MPS
MEDPS
DMMA
PSSD
SC
MPFD
MSI
NP
RPR
SHDI
SHEI
DO
MECI
MNN
IJI
MPS
MEDPS
DMMA
PSSD
SC
MPFD
MSI
NP
RPR
SHDI
SHEI
DO
MECI
MNN
IJI
1.00
0.41
0.59
0.78
0.23
0.15
0.07
0.14
0.08
0.53
0.62
0.42
0.52
0.39
0.54
0.41
1.00
0.06
0.07
0.26
0.03
0.18
0.10
0.32
0.33
0.36
0.27
0.36
0.12
0.02
0.59
0.06
1.00
0.92
0.19
0.21
0.12
0.52
0.26
0.27
0.40
0.61
0.42
0.25
0.54
0.78
0.07
0.92
1.00
0.16
0.22
0.07
0.34
0.18
0.34
0.46
0.63
0.44
0.34
0.56
0.23
0.26
0.19
0.16
1.00
0.00
0.56
0.30
0.23
0.06
0.14
0.08
0.13
0.05
0.16
0.15
0.03
0.21
0.22
0.00
1.00
0.04
0.12
0.16
0.01
0.06
0.18
0.18
0.09
0.10
0.07
0.18
0.12
0.07
0.56
0.04
1.00
0.25
0.25
0.07
0.15
0.09
0.08
0.07
0.01
0.14
0.10
0.52
0.34
0.30
0.12
0.25
1.00
0.57
0.16
0.09
0.45
0.27
0.05
0.39
0.08
0.32
0.26
0.18
0.23
0.16
0.25
0.57
1.00
0.64
0.22
0.53
0.14
0.12
0.16
0.53
0.33
0.27
0.34
0.06
0.01
0.07
0.16
0.64
1.00
0.85
0.23
0.46
0.28
0.54
0.62
0.36
0.40
0.46
0.14
0.06
0.15
0.09
0.22
0.85
1.00
0.52
0.49
0.41
0.71
0.42
0.27
0.61
0.63
0.08
0.18
0.09
0.45
0.53
0.23
0.52
1.00
0.12
0.44
0.73
0.52
0.36
0.42
0.44
0.13
0.18
0.08
0.27
0.14
0.46
0.49
0.12
1.00
0.24
0.49
0.39
0.12
0.25
0.34
0.05
0.09
0.07
0.05
0.12
0.28
0.41
0.44
0.24
1.00
0.23
0.54
0.02
0.54
0.56
0.16
0.10
0.01
0.39
0.16
0.54
0.71
0.73
0.49
0.23
1.00
this series. Expanding to the second order improves the approximation of the true functional form. The results are presented
in Table 4.
Model 1 includes only the four independent variables used in
the separate county analyses: DCOL, DCTY, MECI, and DO. DCTY is
the distance from each TAZ centroid to the CBD/center of the county
where it is located and to Delaware city in the case of Franklin, and
DCOL is the distance from that TAZ centroid to the Columbus CBD.
All the estimated coefcients are signicant at the 5% level, and
with negative signs consistent with the previous results, except
for MECI. Model 2 expands Model 1 by adding county dummy
variables:
IC045 = 1 if the TAZ is in Faireld, =0 otherwise; IC049 = 1 if the
TAZ is in Franklin, =0 otherwise; IC089 = 1 if the TAZ is in Licking, =0 otherwise; IC097 = 1 if the TAZ is in Madison, =0 otherwise;
IC129 = 1 if the TAZ is in Pickaway, =0 otherwise; IC159 = 1 if the
TAZ is in Union, =0 otherwise.
Model 3 expands Model 2 by adding the squares of the variables DCTY, DCOL, MECI, and DO. Model 4 expands Model 3 by
adding the following interaction terms: MECI DO, DCTY MECI,
DCOL MECI, DCTY DO, and DCOL DO. DCOL DCTY turned out
to be insignicant, and is not included. The formulation of Model 4
is:
ln(DP) = exp(a0 +
i=16
(6)
where ICi is the dummy variable for county i. The R2 increases from
0.600 for Model 1 to 0.630 for Model 2, 0.654 for Model 3, and 0.659
for Model 4. In the following analyses, the focus is on Model 4.
All the variables in Model 4 are signicant at the 10% level,
except DO2 , DCOL MECI, and IC129. The dummy variables help
differentiate the intercepts of the density curves, and thus the
CBD densities, among the seven counties. The regression intercept
(6.824) characterizes Delaware county (all the dummy variables = 0), and, in exponential form, is equal to e6.824 = 0.001087.
IC049 (Franklin) has the largest value (1.611), and thus an intercept
of (0.824 + 1.611). Faireld, Licking, and Madison have relatively
close values (0.760, 0.819, 0.613), followed by the cluster of Pickaway and Union (0.242 and 0.380). If there were only one CBD, and
the distance to this CBD (DCTY) was the only independent variable, then this intercept would measure the density at the CBD
(distance = 0). This, however, is not the case because of the other
distance to the alternative CBD (DCOL) and variables MECI and DO.
Nevertheless, the ranking is not unexpected, and reects differences among counties in terms of their urban/rural characters.
The effects on density of other variables (DCTY, DCOL, MECI,
DO) cannot be captured by simple visual inspection, as would
be the case in a linear functional form, because of the squared
and interaction terms. To measure these effects, the elasticities of
the dependent variable (DP) are computed with respect to all the
Table 3
Population density models for the seven counties of the CMA (dependent variable: logarithm of population density).
Variable
Franklin
Delaware
Licking
Faireld
Pickaway
Madison
Union
Intercept
7.170
(15.10)* , a
0.00002
(8.58)*
0.00004
(12.04)*
0.042
(5.41)*
1.356
(8.62)*
11.360
(11.43)*
0.000009
(1.77)**
0.000001
(0.27)
0.080
(4.48)*
2.516
(6.32)*
8.880
(10.90)*
0.000025
(6.41)*
0.000015
(5.34)*
0.083
(6.77)*
1.650
(5.26)*
11.790
(17.25)*
0.000007
(2.06)*
0.000007
(2.53)*
0.111
(9.74)*
1.688
(7.46)*
10.050
(7.63)*
0.000013
(2.00)*
0.000011
(2.37)*
0.078
(2.94)*
2.026
(3.74)*
11.100
(11.45)*
0.000008
(1.43)***
0.000006
(1.30)
0.093
(5.10)*
2.053
(6.33)*
10.200
(9.66)*
0.000009
(1.27)
0.000001
(0.18)
0.037
(1.67)**
2.495
(5.76)*
1117
0.39
105
0.63
208
0.67
86
0.84
67
0.61
103
0.59
77
0.58
DCTY
DCOLc
MECI
DO
Number of observations
R2
*
**
***
a
b
c
p < = 5%.
p < = 10%.
p < = 15%.
Numbers in parentheses denote t-values for each variables.
DCTY denotes the distance to the center of the major city of each of the peripheral counties. In the case of Franklin, it denotes the distance to Delaware city (1 ft = 0.3048 m).
DCOL denotes distance to Columbus CBD (1 ft = 0.3048 m). The original datasets from various sources were mostly measured in ft; thus, ft was used in this analysis.
82
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
Table 4
Pooled metropolitan-level population density models (dependent variable: logarithm of population density).
Variable
Model 1
Model 2
Model 3
Intercept
9.556
(32.62)*
2.10E05
(11.6)*
9.929
(32.58)*
2.41E05
(13.58)*
9.96E06
(8.62)*
1.14E05
(9.06)*
9.313
(15.67)*
7.32E05
(13.63)*
5.41E10
(9.31)*
3.02E05
(4.13)*
7.43E11
(2.67)*
0.137
(7.2)*
1.35E03
(4.52)*
1.171
(2.79)*
0.152
(0.67)
DCTYa
DCTY2
DCOLb
2
DCOL
MECI
0.083
(18.44)*
0.060
(12.22)*
2.004
(18.52)*
1.584
(14.15)*
MECI2
DO
DO2
MECI DO
DCTY MECI
DCTY DO
DCOL MECI
DCOL DO
IC045
IC049
IC089
IC097
IC129
IC159
R2
*
**
***
a
b
0.597
0.757
(3.84)*
1.397
(9.57)*
1.093
(6.38)*
0.412
(2.11)*
0.229
(1.08)
0.400
(1.93)*
0.630
0.736
(3.86)*
1.673
(11.56)*
0.835
(4.91)*
0.454
(2.35)*
0.178
(0.86)
0.312
(1.52)***
0.654
Model 4
6.824
(5.47)*
9.62E05
(8.47)*
5.09E10
(7.42)*
4.01E05
(3.44)*
5.53E11
(1.81)**
0.105
(3.38)*
1.74E03
(5.24)*
3.440
(4.45)*
0.267
(1.07)
0.029
(2.72)*
3.93E07
(1.86)**
1.48E05
(3.03)*
2.01E07
(1.21)
8.97E06
(2.29)*
0.760
(4.00)*
1.611
(11.02)*
0.819
(4.80)*
0.613
(3.13)*
0.242
(1.17)
0.380
(1.85)**
0.659
p < = 5%.
p < = 10%.
p < = 15%.
DCTY: distance to county CBD.
DCOL: distance to alternative CBD (Delaware city for Franklin county, and Columbus CBD For the surrounding six counties).
DP DP
X X
(7)
(8)
(9)
MECI = (a5 + 2a7 MECI + a9 DO + a10 DCTY + a12 DCOL) MECI (10)
DO = (a6 + 2a8 DO + a9 MECI + a11 DCTY + a13 DCOL) DO
(11)
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
83
Table 5
Basic statistics for model variables (1 ft = 0.3048 m).
County
Variable
Mean
Minimum
Maximum
Standard deviation
CMA
DCTY (ft)
DCOL (ft)
MECI
DO
36,354
125,110
33.80
0.89
0
59,231
0.00
0.00
11,1631
235,065
58.26
1.89
22,480
32,660
9.37
0.37
Delaware
DCTY (ft)
DCOL (ft)
MECI
DO
42,458
109,471
28.57
1.08
1755
65,110
10.84
0.32
98,679
175,109
50.99
1.77
23,975
28,803
6.86
0.30
Faireld
DCTY (ft)
DCOL (ft)
MECI
DO
42,251
129,349
27.37
1.02
432
59,805
9.64
0.24
93,476
197,781
43.77
1.57
25,030
32,314
7.10
0.33
Franklin
DCTY (ft)
DCOL (ft)
MECI
DO
34,448
117,824
37.50
0.77
1037
59,231
0.00
0.00
81,154
193,514
55.70
1.83
18,617
26,924
7.96
0.31
Licking
DCTY (ft)
DCOL (ft)
MECI
DO
32,382
159,183
33.10
0.93
90
68,047
0.00
0.00
111,631
235,065
58.26
1.81
29,759
31,946
7.82
0.29
Madison
DCTY (ft)
DCOL (ft)
MECI
DO
44,991
118,972
22.47
1.28
0
69,113
0.00
0.13
110,020
195,609
44.12
1.89
24,563
29,217
7.86
0.43
Pickaway
DCTY (ft)
DCOL (ft)
MECI
DO
46,374
121,611
24.59
1.29
0
65,027
8.94
0.32
102,749
178,231
40.17
1.75
26,123
31,477
6.53
0.34
Union
DCTY (ft)
DCOL (ft)
MECI
DO
37,451
151,537
24.75
1.21
884
74,119
0.00
0.07
92,544
230,054
44.30
1.84
25,726
36,247
8.64
0.44
varies between 1.56 and 3.68, with a mean of 1.39, and DO varies
between 4.76 and 0.04, with a mean of 1.20. Again, there are
combinations of the independent variables that will produce signs
for MECI and DO that are different from the central tendencies.
However, such deviations are very limited in the case of DO , with a
maximum value of 0.04. Similar across-sign patterns emerge for
some counties, but not all. This is the case of Franklin, which is not
surprising, as it dominates the CMA sample; DCTY changes signs in
all counties, but DCOL remains always negative in Delaware, Faireld, and Pickaway counties; MECI is always positive in Madison
and Pickaway counties; and DO is always negative in Delaware,
Faireld, Madison, and Pickaway counties.
Table 7
Basic statistics for population density elasticities.
County
Variable
Mean
Minimum
Maximum
Standard deviation
CMA
DCTY
DCOL
MECI
DO
DCTY
DCOL
MECI
DO
DCTY
DCOL
MECI
DO
DCTY
DCOL
MECI
DO
DCTY
DCOL
MECI
DO
DCTY
DCOL
MECI
DO
DCTY
DCOL
MECI
DO
DCTY
DCOL
MECI
DO
0.67
1.30
1.39
1.20
0.52
1.26
1.94
1.74
0.53
1.31
2.00
1.49
0.85
1.41
1.03
0.94
0.27
1.07
1.74
1.02
0.41
1.18
2.10
2.31
0.22
1.12
2.24
2.15
0.45
0.96
2.08
1.81
1.90
3.61
1.56
4.76
1.50
2.39
0.87
3.61
1.76
2.54
0.46
3.02
1.84
3.28
1.48
3.98
1.36
3.61
1.56
2.14
1.74
2.26
0.00
4.76
1.38
2.05
0.64
3.69
1.90
2.80
0.32
3.93
5.53
0.95
3.68
0.04
3.25
0.13
3.01
0.49
2.04
0.34
2.92
0.23
1.02
0.01
3.03
0.02
5.53
0.95
3.48
0.00
2.93
0.58
3.45
0.10
4.52
0.44
3.23
0.27
2.60
0.63
3.68
0.04
0.73
0.45
0.96
0.77
0.81
0.32
0.76
0.70
0.76
0.38
0.67
0.70
0.39
0.40
0.88
0.54
1.16
0.52
0.92
0.48
0.93
0.48
0.57
1.05
1.13
0.37
0.60
0.79
0.91
0.70
0.81
0.99
Delaware
Faireld
7. Discussion
Franklin
Licking
Madison
Table 6
Elasticities for each county and the CMA at the sample means.
County
DCTY
DCOL
MECI
DO
CMA
Delaware
Faireld
Franklin
Licking
Madison
Pickaway
Union
1.19
1.09
1.16
1.21
1.18
1.02
0.94
1.14
1.43
1.37
1.44
1.49
1.19
1.30
1.23
1.13
1.78
2.16
2.24
1.27
2.04
2.37
2.45
2.11
1.14
1.71
1.43
0.92
1.00
2.23
2.11
1.76
Pickaway
Union
84
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
8. Conclusions
As discussed in the previous section, this research is deemed
to have achieved its goals of incorporating land-use structure
variables into population density models. It could, however, be
extended in several directions. First, the analysis is cross-sectional,
with variables measured at one time period and with the implicit
assumption that the models are static and with non-varying coefcients. However, population and land-use structure are highly
dynamic, as demonstrated in the cluster analysis of residential density in the U.S. North Central Region by Hammer et al. (2004).
If data could be assembled over several time periods, dynamic
density models could be estimated. Second, in order to achieve a
more generalizable theory about the interactions between landscape indices and population density, similar models should be
tested in other metropolitan areas to see if specic characteristics
(size, shape, dominant land use types, etc.) impact these interactions, with a particular focus on the role of land-use dominance and
edge structure. Models could also be tested while including only
the surrounding suburban areas, and excluding forest and agriculturally dominated nearby rural areas. The increasing availability
of very detailed remote-sensing data and GIS land-use coverages
make such extensions feasible, but at probably large computational
costs.
References
Alperovich, G. (1983). An empirical study of population density gradients and their
determinates. Journal of Regional Science, 23(4), 529540.
Anderson, J. (1982). Cubic-spline urban-density functions. Journal of Urban Economics, 12, 155187.
Batty, M., & Kim, K. S. (1992). Form follows function: Reformulating urban population
density functions. Urban Studies, 29(7), 10431070.
Bockstael, N. E. (1996). Modeling economics and ecology: The importance of a spatial
perspective. American Journal of Agricultural Economics, 78(5), 11681180.
Bunting, T., Filion, P., & Priston, H. (2002). Density gradients in Canadian metropolitan regions, 197196: Differential patterns of central area and suburban growth
and change. Urban Studies, 39(13), 25312552.
J. Lu, J.-M. Guldmann / Landscape and Urban Planning 105 (2012) 7485
Herold, M., Couclelis, H., & Clarke, K. C. (2005). The role of spatial metrics in the
analysis and modeling of urban land use change. Computers, Environment, and
Urban Systems, 29(4), 369399.
Holloway, S., Bryan, D., Chabot, R., Rogers, D. M., & Rulli, J. (1998). Exploring the
effect of public housing on the concentration of poverty in Columbus, Ohio.
Urban Affairs Review, 33(6), 767789.
Huang, J., Lu, X. X., & Sellers, J. M. (2007). A global comparative analysis of urban form:
Applying spatial metrics and remote sensing. Landscape and Urban Planning, 82,
184197.
Hurd, J., Wilson, E., Lammey, S., & Civco, D. (2001). Urban sprawl using time sequential
landsat imagery. ASPRS 2001 Annual Convention, St. Louis, MO, April 2327
Jenerette, G. D., & Potere, D. (2010). Global analysis and simulation of land-use
change associated with urbanization. Landscape Ecology, 25, 657670.
Ji, W., Ma, J., Twibell, R. W., & Underhill, K. (2006). Characterizing urban sprawl
using multi-stage remote sensing images and landscape metrics. Computers,
Environment, and Urban Systems, 30, 861879.
Kareiva, P. (1990). Population dynamics in spatially complex environments: Theory
and data. Philosophical Transactions of the Royal Society of London Series B, 330,
175190.
Lloret, F., Calvo, E., Pons, X., & Daz-Delgado, R. (2002). Wildres and landscape
patterns in the Eastern Iberian Peninsula. Landscape Ecology, 17(8), 745759.
Luck, M., & Wu, J. (2002). A gradient analysis of urban landscape pattern: A case
study from the Phoenix Metropolitan Region, Arizona, USA. Landscape Ecology,
17, 327339.
McDonald, J. (1989). Econometric studies of urban population density: A survey.
Journal of Urban Economics, 26(3), 361385.
McDonald, J., & Bowman, H. W. (1976). Some tests of alternative urban population
density functions. Journal of Urban Economics, 3(3), 242252.
McGarigal, K., Cushman, S. A., Neel, M. C., & Ene, E. (2002). FRAGSTATS: Spatial pattern
analysis program for categorical maps. Amherst, MA: University of Massachusetts.
http://www.umass.edu/landeco/research/fragstats/fragstats.html
McGlynn, I., & Okin, G. (2006). Characterization of shrub distribution using high spatial resolution remote sensing: Ecosystem implications for a former Chihuahuan
Desert Grassland. Remote Sensing of Environment, 10, 554566.
Miller, J. N., Brooks, R. P., & Croonquist, M. J. (1997). Effects of landscape patterns on
biotic communities. Landscape Ecology, 12, 137153.
Mills, E. S. (1992). The measurement and determinants of suburbanization. Journal
of Urban Economics, 15, 117.
Millward, H., & Bunting, T. (2008). Patterning in urban population densities: A
spatiotemporal model compared with Toronto 19712001. Environment and
Planning A, 40, 283302.
Munroe, D. K., Croissant, C., & York, A. M. (2005). Land use policy and landscape
fragmentation in an urbanizing region: Assessing the impact of zoning. Applied
Geography, 25, 121141.
Newling, B. E. (1969). The spatial variation of urban population densities. Geographical Review, 59(2), 242252.
Nott, M. (2000). Identifying management actions on DoD installations to reverse
declines in neotropical birds. Technical Report Summarizing Findings of the First
Year of DoD Legacy Project Number 00103. https://www.denix.osd.mil/denix/
Public/ES-Programs/Conservation/Legacy/Neo-Tropical/neotropical.html
85
ONeill, R. V., Krummel,. J. R., Gardner, R. H., Sugihara, G., Jackson, B., DeAngelis, D.
L., et al. (1988). Indices of landscape pattern. Landscape Ecology, 1(3), 153162.
Perfecto, I., & Vandermeer, J. (2002). Quality of agroecological matrix in a tropical
montane landscape: Ants in coffee plantations in Southern Mexico. Conservation
Biology, 16(1), 174182.
Pster, J., 2004. Using landscape metrics to create an index of forest fragmentation for
the state of Maryland (Master thesis). Towson University.
Riitters, K., ONeill, R. V., Hunsaker, C., Wickham, J., Yankee, D., Timmins, S., et al.
(1995). A factor analysis of landscape pattern and structure metrics. Landscape
Ecology, 10(1), 2339.
Rutledge, D. (2003). Landscape indices as measures of the effects of fragmentation: Can
pattern reect process? DOC Science Internal Series 98. Wellington, New Zealand:
Department of Conservation.
Smith, B. (1997). A review of monocentric urban density analysis. Journal of Planning
Literature, 12(2), 115135.
Schwarz, N. (2010). Urban form revisited Selecting indicators for characterizing
European cities. Landscape and Urban Planning, 96, 2947.
Steffan-Dewenter, I. (2003). Importance of habitat area and landscape context for
species richness of bees and wasps in fragmented orchard meadows. Conservation Biology, 17(4), 10361044.
Tilman, D., & Kareiva, P. (1997). Spatial ecology: The role of space in population dynamics and interspecic interactions. Princeton, NJ: Cambridge University Press.
Tscharntke, T., Steffan-Dewenter, I., Kruess, A., & Thies, C. (2002). Contribution of
small habitat fragments to conservation of insect communities of grasslandcropland landscapes. Ecological Applications, 12(2), 354363.
Turner, M. G., Gardner, R. H., & ONeill, R. V. (2001). Landscape ecology in theory and
practice. New York: Springer.
U.S. Geological Survey. (2000). National land cover data product description.
http://eros.usgs.gov/products/landcover/nlcd.html
Venema, H. D., Calamai, P. H., & Fieguth, P. (2005). Forest structure optimization using
evolutionary programming and landscape ecology metrics. European Journal of
Operational Research, 164, 423439.
Zeng, H., & Wu, X. B. (2005). Utilities of edge-based metrics for studying landscape
fragmentation. Computers, Environment, and Urban Systems, 29, 159178.
Zhou, & Kockelman. (2008). Neighborhood impacts on land use change: A multinomial logit model of spatial relationships. Annals of Regional Science, 42(2),
321340.
Jia Lu is an assistant professor at Valdosta State University. She received her Ph.D. in
city and regional planning from The Ohio State University. She also holds a master
degree in Community Planning from University of Cincinnati. Her research interests focuses on urban modeling, urban planning, GIS, environmental studies, and
geography. She has published a book and a few book chapters regarding these issues.
Jean-Michel Guldmann is a professor of city and regional planning at The Ohio State
University. He holds a Ph.D. in urban and regional planning from the Israel Institute
of Technology, Haifa, Israel, and a masters in industrial engineering from the Ecole
des Mines, Nancy, France. His research focuses on the quantitative modeling of urban
structure, energy and telecommunications infrastructures, and environmental planning, particularly air quality and water issues. He has published a book and over 60
articles and book chapters on these issues.