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CONICET, Departamento de Geologa Universidad Nacional del Sur, San Juan 670, 8000 Baha Blanca, Buenos Aires, Argentina
Facultad de Ciencias Exactas y Naturales, Universidad Nacional de La Pampa, Avenida Uruguay 151, 6300 Santa Rosa, La Pampa, Argentina
c
CIC, Divisin Paleontologa Vertebrados, Museo de La Plata, Paseo del Bosque s/n, 1900 La Plata, Argentina
d
Museo Municipal de Ciencias Naturales Carlos Darwin, Urquiza 123, 8109 Punta Alta, Argentina
b
a r t i c l e i n f o
a b s t r a c t
Article history:
Received 29 March 2013
Accepted 8 August 2013
The Monte Hermoso Formation, cropping out at its type locality of Farola Monte Hermoso (Buenos Aires
Province), is a classical fossiliferous unit of the South American Neogene, highlighted by the abundance
and diversity of its vertebrate remains. However, its biostratigraphy and age have been largely debated,
and numerous discrepancies and controversies have been stated. In this regard, the result of the analysis
of new materials recovered from the different levels of this formation, following a strict control of
stratigraphic provenance, is here reported. As well, the provenance of specimens of previous collections
has been evaluated. The studied assemblage consists of Osteichthyes, Amphibia, Reptilia, Aves and
Mammalia. These latter are the most numerous and belong to the Didelphimorphia, Polydolopimorphia,
Rodentia, Notoungulata, Litopterna and Xenarthra. The recorded taxa suggest no important faunistic
variations among the different levels of the Monte Hermoso Formation that would imply signicant
chronological differences, and hence, justify the recognition of two biostratigraphic units. The analysis of
the rst and last records as well as the taxa considered as exclusive, does not support the validity of the
biozones of Trigodon gaudryi and Neocavia depressidens previously proposed. On this basis, a new scheme
for the Monte Hermoso Formation at its type locality is proposed, including a new single biostratigraphic
unit. This unit is the Eumysops laeviplicatus Range Zone, which represents the biostratigraphic base for
the Montehermosan Stage/Age of the early Pliocene.
2013 Elsevier Ltd. All rights reserved.
Keywords:
Biostratigraphy
Monte Hermoso Formation
Eumysops laeviplicatus Zone
Early Pliocene
Argentina
1. Introduction
The biostratigraphic units of the late Cenozoic of South America
are mainly based on the taxonomic content and distribution of the
fossil mammal assemblages (Cione and Tonni, 2005; Cione et al.,
2007; Deschamps et al., 2012). In this regard, numerous deposits
including levels bearing a rich vertebrate fauna, especially mammals, of the late MioceneeHolocene are exposed at the Pampean
Region (Argentina). These deposits, which sometimes include the
type sections of the biostratigraphic units of the late Cenozoic, are
among the most complete sequences in the continent (Zrate,
2005; Cione et al., 2007).
Farola Monte Hermoso, located at the south-west of the Buenos
Aires Province (Pampean Region, Argentina; Fig. 1), is the type locality of the Monte Hermoso Formation, formally dened by Zavala
32
R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
Fig. 1. Location map showing Farola Monte Hermoso, Buenos Aires Province, Argentina.
R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
33
Fig. 2. Stratigraphic section of the Monte Hermoso Formation at Farola Monte Hermoso locality.
climbing ripples. The Fsc facies includes dark brown to lightyellowish brown siltstones, with ne lamination and structures of
gravitational deformation (slumping). These deposits rest transitionally over the Sr facies. In both cases, fossil remains are very
scarce (Tomassini, 2012).
The Fcf facies includes massive muddy siltstones, lightyellowish brown, that overlie the Fsc facies transitionally (Fig. 2).
Fossil remains are abundant, as well as small mammal burrows,
these latter with similar features as those of Fl facies. Traces
attributed to T. barretti were also recorded (Tomassini, 2012).
3. Previous biostratigraphic framework
As mentioned above, several biostratigraphic schemes were
proposed for the deposits of the Monte Hermoso Formation.
Among them, the most recent one is that of Cione and Tonni (1995,
1996, 2001, 2005) who stated that the different levels of this formation showed evidence of important faunistic variations of temporal signicance. On this basis, they proposed the existence of two
biostratigraphic units, chronologically successive, characterized by
the presence of taxa, considered as exclusive. According to their
proposal:
1 T. gaudryi Biozone: is the biostratigraphic basis for the Montehermosan Stage/Age. It was assigned to the late Miocenee
early Pliocene and coincides spatially with the Hermosense
tpico of Vignati (1925), correlated units showed in Fig. 3, and
the lowest part of the Monte Hermoso Formation of Zavala
(1993). Exclusive taxa: T. gaudryi, Parahyaenodon argentinus,
Notocynus hermosicus, Myrmecophaga caroloameghinoi, Palaeodaedicurus antiquus, Diplasiotherium robustum, Alitoxodon
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R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
Fig. 3. Stratigraphic interpretations of the Farola Monte Hermoso locality. Modied from Zavala, 1993.
R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
Fig. 4. Pie diagram showing the percentage of vertebrate remains recovered in each
facies of the Monte Hermoso Formation.
35
36
R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
Table 1
List of vertebrates represented in the facies of the Monte Hermoso Formation and NISP values for each taxon.
Taxa
Class Osteichthyes
Osteichthyes indet.
Siluriformes indet.
Trichomycteridae indet.
Percichthyidae indet.
Characidae indet.
Class Amphibia
Anura indet.
Bufonidae indet.
Rhinella cf. R. pisanoi
Ceratophrys ameghinorum
Lepidobatrachus laevis
4
13
2
Class Reptilia
Chelonoidis cf. Ch. australis
Tupinambis sp.
Colubridae indet.
2
3
1
Class Aves
Aves indet.
Dendrocygninae indet.
Eudromia cf. E. elegans
Mesembriornis milneedwardsi
Chunga incerta
2
1
1
1
1
Class Mammalia
Mammalia indet.
Thylatheridium pascuali
Lutreolina sp.
Hyperdidelphys inexpectata
Sparassocynus sp.
Argyrolagus palmeri
Macrochorobates chapalmalensis
Holozaedyus laevisculptus
Doellotatus inornatus
Plaina cf. P. intermedia
Glyptodontidae indet.
Plohophorus guratus
Eleutherocercus antiquus
Proscelidodon cf. P. patrius
Lestodon sp.
Rodentia indet.
Pithanotomys columnaris
Neophanomys sp.
Actenomys priscus
Eumysops laeviplicatus
Paramyocastor diligens
Lagostomus incises
Caviidae indet.
Palaeocavia sp.
Neocavia sp.
Orthomyctera sp.
Prodolichotis sp.
Caviodon australis
Caviodon sp.
Phugatherium cataclisticum
Cricetidae indet.
Sigmodontinae gen. et. sp. nov.
Necromys bonapartei
Auliscomys formosus
Paedotherium sp.
Paedotherium bonaerense
Paedotherium typicum
Tremacyllus impressus
Pseudotypotherium sp.
Toxodontidae indet.
Toxodon chapalmalensis
Proterotheriidae indet.
Aff. Diplasiotherium robustum
Eoauchenia cf. E. primitiva
Macraucheniidae indet.
Facies Fm NISP
26
1
1
1
1
1
1
2
2
1
7
4
2
6
1
18
2
1
14
3
2
2
1
2
1
2
1
1
2
2
Facies St NISP
52
2
5
24
Facies Sr NISP
30
3
3
4
1
1
1
1
3
1
1
1
1
1
18
48
1
2
33
8
5
2
11
6
1
1
1
2
1
1
1
3
2
2
4
1
1
2
1
1
1
1
1
1
1
5
36
2
11
13
3
3
3
8
2
13
2
14
1
1
2
2
1
1
1
1
2
1
1
26
2
1
1
2
1
2
1
1
27
6
4
2
3
1
1
1
1
1
1
17
7
2
1
4
1
1
R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
37
Table 1 (continued )
Taxa
Promacrauchenia antiqua
Total
223
Facies Fm NISP
Facies St NISP
Facies Sr NISP
87
279
49
16
95
Fig. 5. Holotype of Eumysops laeviplicatus (MACN-A 1623), from the Monte Hermoso Formation. A. Dorsal view. B. Ventral view. C. Lateral view. Scale 1 cm.
38
R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
39
Fig. 7. Biostratigraphic scheme of the late Cenozoic of the Pampean Region, including the Eumysops laeviplicatus Zone dened in this paper. Modied from Cione and Tonni, 2005;
Cione et al., 2007.
7. Conclusions
The discovery of new specimens, together with the detailed
evaluation of specimens of previous collections, all of them with
accurate stratigraphic provenance, allowed the proposal of a new
biostratigraphic scheme for the Monte Hermoso Formation, at its
type locality of Farola Monte Hermoso (Pampean Region,
Argentina). This scheme recognizes a single unit, the E. laeviplicatus
Range Zone, which represents the biostratigraphic basis for the
Montehermosan Stage/Age and corresponds to the early Pliocene.
Several authors (e.g. Cione et al., 2007; Zrate et al., 2007) have
pointed out the necessity of a strong revision of the biostratigraphic
units dened for the late Cenozoic of Argentina. In this sense, this
proposal provides a new framework for biostratigraphic comparisons and correlations with other fossiliferous localities of
Argentina, mainly in the Pampean Region (e.g. Caleuf, Balneario
Saldungaray, Dique Paso Piedras, Las Obscuras, Cascada Grande,
Indio Rico, Paso Otero), that contain deposits and faunistic assemblages related to this interval.
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R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
Monte Hermoso
Formation facies
Fl
Class Osteichthyes
Order Siluriformes
Siluriformes indet.
Trichomycteridae indet.
Order Perciformes
Percichthyidae indet.
Order Characiformes
Characidae indet.
Class Amphibia
Order Anura
Rhinella cf. R. pisanoi Casamiquela, 1967
Rhinella cf. R. schneideri Werner, 1894
Ceratophrys ameghinorum Fernicola, 2001
Lepidobatrachus laevis Budgett, 1899
Class Reptilia
Order Chelonii
Chelonoidis australis (Moreno, 1889)
Chelonoidis cf. Ch. australis
Order Squamata
Tupinambis sp.
Callopistes bicuspidatus Chani, 1976
Colubridae indet.
aff. Bothrops
Boa sp.
Fm St Sr Fsc Fcf
B
C
C
C C
C C
C
?
C
?
C
C
?
C
?
?
Class Aves
Order Anseriformes
Dendrocygninae indet.
Order Tinamiformes
Nothura parvula Tambussi, 1989
Eudromia cf. E. elegans
Eudromia olsoni Tambussi and Tonni, 1985
Order Rheiformes
Heterorhea dabbenei Rovereto, 1914
Hinasuri nehuensis Tambussi, 1995
Order Ralliformes
Mesembriornis milneedwardsi
Moreno and Mercerat, 1891
Chunga incerta Tonni, 1974
Order Ciconiiformes
Vultur gryphus Linnaeus, 1758
Dryornis pampeanus Moreno and Mercerat, 1891
?
?
Class Mammalia
Order Didelphimorphia
Thylatheridium pascuali Reig, 1958
Thylamys contrerasi Mones, 1980
Thylophorops aff. T. perplana Ameghino, 1904
Lutreolina tracheia Rovereto, 1914
Lutreolina biforata Rovereto, 1914
Lutreolina sp.
Hyperdidelphys inexpectata Ameghino, 1889
C
B?
?
?
?
C
C
C C
C
C C
C C
B
B
C
B
C
?
?
?
C
(continued )
Taxa
Monte Hermoso
Formation facies
Fl
B?
B
C
Fm St Sr Fsc Fcf
B
B
?
?
C
?
?
C
C
?
B
B
?
C
B
B
C
?
B
?
B
C
?
?
?
?
?
C
?
B
B
B?
B
C C
C C
B
B
B
C
C C
B?
C
C
?
?
?
B?
C
?
?
C
C
C
C
?
?
?
C C C C C C
C
C
C
B?
C C C
C
?
C
C
C
C C C
C
B?
C
?
C
C
B
C
C
C
C
B?
B?
C C C
C
?
C
B?
C
B?
C
B?
C
C
C C
C C
C C
C
R.L. Tomassini et al. / Journal of South American Earth Sciences 48 (2013) 31e42
(continued )
Taxa
Monte Hermoso
Formation facies
Fl
Fm St Sr Fsc Fcf
C C C
C C C
C
?
B?
B?
B?
B?
B
B
C
B?
?
C
C
C C
C C
B?
B
C
B
B?
?
?
?
B?
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