Urena grandiflora DC. Urena trilobata Vell. Urena lobata L. Urena diversifolia Schumach. (and others: Institute of Systematic Botany 2003).
pubescent both above and below, ovate, angulate
or shallowly lobbed, 1 to 12 by 1 to 12 cm, and have serrate margins. Axillary flowers are usually solitary and have five pink petals. The flowers are about 1 cm broad. The fruits are 8- to 10-mm globose capsules that break into five fine-barbed (glochidate) mericarps. (Howard 1989, Liogier 1994, Stevens and others 2001)
Range.The original range of Caesar weed is
probably Asiatic. Today it grows throughout moist tropic and subtropical regions including American Samoa, Florida, Guam, Hawaii, Louisiana, Puerto Rico, and the U.S. Virgin Islands (Natural Resources Conservation Service 2003, Pacific Island Ecosystems at Risk 2003).
Ecology.Caesar weed readily invades disturbed
areas, especially poorly managed pastures, scarified and eroded areas, and perennial crop plantations but is much less of a problem in annual crops. The species does not compete well in tall grass swards and brushlands and does not grow under forest canopies. Ceasar weed grows on a wide variety of soils of varying fertility derived from most parent materials. It tolerates salt spray and a moderate amount of salt in the soil but does General Description.Caesar weed is also not grow in saturated soils. The species ranges to known as hibiscus bur, aramina, pink Chinese elevations of 1,500 m (Pacific Island Ecosystems burr, bur mallow, grand cousin, cadillo, carrapicho at Risk 2003). In Puerto Rico, it occurs in areas do mata, malva, mahot cousin, cousin petit, cousin that receive from about 1400 to 3000 mm of mean rouge, jut africain, cooze mahot, dadangsi, and annual precipitation, forming thickets in favorable mautofu. It is a subshrub 0.6 to 3 m in height and sites. up to 7 cm in basal diameter. The species usually has a single stem emerging from the ground but Reproduction.Caesar weed flowers and fruits normally produces several stems or major throughout the year (Stevens and others 2001). A branches low on the stem and many branches collection of seeds from Puerto Rico averaged throughout. The nearly smooth bark is tough and 0.0239 + 0.0003 or 41,800 seeds/kg. Placed on fibrous, brown on the outside and green within. Its moist blotter paper without pretreatment, just 3 pale yellow wood is of medium density. Plants are percent germinated in 9 months. Germination is supported by a tap and lateral root system of tan or epigeal (authors observation). Sulfuric acid ivory-colored, tough and flexible roots. Grayish- scarification was tested as a means of breaking green, discolorous, alternate leaves are stellate- dormancy of seed from Sierra Leone. The best treatment (18 molar solution for 90 minutes) gave Escolar Universitario, Mandiola, Philippines. 96 percent germination as compared with distilled http:// www.ceu.edu.ph/research1.htm. 3 p. water that gave 4 percent germination (Harris 1986). Seeds are dispersed by clinging to fur and Cmara de Comrcio e Indstria Portugal-Angola. clothing. Seedlings are common in disturbed sites 2003. Caracterizao econmico social de near seed sources. Angola. http://www.cciportugal-angola.pt/ caract.htm. [not paged]. Growth and Management.Caesar weed grows rapidly and can reach 0.5 to 2 m by the end of the Fagundes, M.H. 2003. Sementes de juta e malva: first year. In Puerto Rico, it can live 2 years, algumas observaes. Companhia Nacional de usually dying back to midheight after the first Abastecimento, Brazil. http://www.conab.gov. growing season. It is not known whether the br/politica_agricola/Conjunturasemanal/Especiai shrubs sprout from the roots multiple times. A s/SEMENTES%20JUTA%MALVA.doc. 10 p. fiber crop yielding 1,800 kg/ha is ready to harvest after 6 to 7 months and seed crops of 300 to 500 FAO. 2003. Food composition table for use in kg/ha are ready after 7 or 8 months (Fagundes Africa. Food and Agriculture Organization of 2003). Fiber crops are established with seed. the United Nations, Rome. http://www.fao. Because of the aggressive nature of the species, org/docrep/003/x6877e/X687705.htm#ch5. [not wildland plantings are not recommended. The paged]. author knows of no published specific control measures. Forest Research Institute of Malaysia. 2003. Plants information: Urena lobata Griff., pulut-pulut, Benefits and Detriments.Caesar weed Malvaceae. Forest Research Institute of colonizes disturbed areas and helps to protect the Malaysia, Kuala Lumpur, Malaysia. http://www. soil while furnishing cover for wildlife. It has frim.gov.my/tu/Urena.htm. 1 p. attractive flowers and contributes to aesthetics of areas it has colonized. Tens of thousands of tons of Freedman, R. 1998. Famine foods: Malvaceae. a jute-like fiber from Caesar weed called aramina Perdue University, Perdue, IN. http://www.hort. fiber and Congo jute are produced in Brazil perdue.edu/newcrop/faminefoods/ff_families/M (Fagundes 2003) and Africa (Cmara de Comrcio ALVACEAE.html. 3 p. e Indstria Portugal-Angola 2003). Various extracts of leaves and roots are used in herbal Harris, P.J.C. 1986. Dormancy of Urena lobata L. medicine to treat such diverse ailments as colic, seeds. I. Development of sulphuric acid malaria, gonorrhea, fever, wounds, toothache, and scarification techniques. Ghana Journal of rheumatism (Forest Research Institute of Malaysia Agriculatural Science 14-19: 79-84. 2003). A semipurified glycoside obtained from Caesar weed leaves was 86 percent as effective an Howard, R.A. 1989. Flora of the Lesser Antilles, anti-inflammatory as aspirin in rats (Bautista Leeward and Windward Islands. Vol. 5. Arnold 2000). The leaves and flowers are eaten as famine Arboretum, Harvard University, Jamaica Plain, food in Africa (Freedman 1998). Raw leaves are MA. 604 p. reported to contain 81.8 percent moisture, 54 cal, 3.2 g of protein, 0.1 g fat, 12.8 g carbohydrates, Institute of Systematic Botany. 2003. Atlas of 1.8 g fiber, and 2.1 g ash, 558 mg calcium, and 67 Florida vascular plants. University of South mg of phosphorous per 100 g (FAO 2003). Florida, Tampa, FL. http://plantatlas.usf. However, the plant is little browsed by cattle and edu/synonyms.asp?plantID=1364&genus=Urena can become a severe weed in pastures and &species=lobata. [not paged]. plantations. Burs that collect on clothing and in animal fur are a nuisance. Liogier, H.A. 1994. Descriptive flora of Puerto Rico and adjacent Islands. Vol. 3. Editorial de la References Universidad de Puerto Rico, Ro Piedras, PR. 461 p. Bautista, L.M.A. 2000. Inquiry into the anti- inflammatory activity of the syrup from the Natural Resources Conservation Service. 2003. glycosides of the leaves of kulutkulutan (Urena Plant profile: Urena lobata L., Caesarweed. lobata, Linn., family Malvaceae). Centro http://plants.usda.gov/cgi_bin/plant_profile.cgi? symbol=URLO. 5 p. Pacific Island Ecosystems at Risk. 2003. Urena lobata L., Malvaceae. http://www.hear.org/ Pier/urlob.htm. 3 p.
Stevens, W.D., C. Ulloa-U., A. Pool, and O.M.
Montiel, eds. 2001. Flora de Nicaragua. Monographs in Systematic Botany Vol. 85, No. 2. Missouri Botanical Garden, St. Louis, MO. p. 945-1,910.
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John K. Francis, Research Forester, U.S.
Department of Agriculture, Forest Service, International Institute of Tropical Forestry, Jardn Botnico Sur, 1201 Calle Ceiba, San Juan PR 00926-1119, in cooperation with the University of Puerto Rico, Ro Piedras, PR 00936-4984