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SUCCULENTS

- THE ILLUSTRATED DICTIONARY -


Succuient.oJ:, with their huge diversity of forms
and habits, are one of me most fasdnating - lUI
well as one oflhe lar gest - plant groups (ound
growing wild throughout the world. Many of tho
genera - Agave, Hawarlhia, Kolunchoe and
Yucca , for example - include some of the best
known and most often cultivated houseplanL8.
Other specie8, howcver, are exotic:: and rore,
some seldom seen in cultivation, others now
rarely seen in the ....ild.
Neve r before has there been a single work of
reference in which examples of so wide a range
of genera have been described and illustrated in
colour. Succulents: The Illustrated Dictionory.
the companion volume to Cacti: The lIlulltrated
Dictinmlry, includes morc than 1,200
photogrnphsofspcciesand varicticsfrom 195
difTerent genera, and t herefore itconstitutcs 8
unique work of reference for succulent
enthusiast8 and CQllectors.
For case of use, the dictionary is organh:ed
alphabe~ically by ~pe<:ies. Each cntry indudcs a
full de&<:ript ion of the ~pecies as well as ~he place
of origin . Commonly used alternative names are
given, and these are also listed separately for
oonvenientcross-rcferenoo. In addition, the
introduc~io n summarizcs the charllcteriatica of
lhefamiHcH and genera.
A special feature of the text is the indusion of
the CITES (Convention on International Trade
in Endangered Spccics) status for all those
species that are indudcd in the Convention's
Appendix I and Appendix II of protected pla nts.
There is also a de&<:ription o[ the way in which
CITES is organbed to monitor the ~rade in
collectnblc spccies, as well as a summaryo[the
codeofconductofthelnternationnl
Organiza~ion for Succulent Plants Study for
oollect.orfl.
With such a wealth of information, Succulent.s :
The Illustrated Dictionary is assured of a place
among the standard ....orks of rc ference of all
succulent growers and ooUectors..

TIMBER PRESS
Portland, Oregon $29.95
ISBN o-aa]''l~'3''1a-~
SUCCULENTS
THE ILLUSTRATED
DICTIONARY

Maurizio Sajeva
'od

Mariangela Costanzo

limber Press
Port/and, Oregon
R'·,>rinu'<.'998 .lOOO.2001

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CONTENTS
ACKNOWLErx:;EMENTS

FOREWORD

INTRODUCTION

SUCCUlEl\'T Fk\4IL1ES ANDGENEKA 11

ILLUSTRATED A - Z OF SPECIES

CONVENT10;.J ON it-.'TERNATIONAL TRADE


IN ENDANCEREDSPECIES 228

INTERNAT IONAL ORGANIZATION FOR SUCCULENT


PLANT'S STUDY: CODE OF CONDUCT

CI IECKLISTOF ALTERl'JATlVE NAMES 233

FURTHER READING 237

CRFOIl'S 239
ACK NOWLEDGEMENTS
w~ would like to thank LucOl Magilgnoli for his encouragement and for his h~Jp
in (ocOlling interesting plants; Andrea Catlabriga for many useful suggcstions;
AnnarOS1l Nicola and P<lsquale Ruocco for supplying transparmcies; Sara
Oldfit'ld for pennissiOll to reprint extracts from the lOS Code of Conduct;
ROSt'mary Anderson of Casse:1i ;md Lydia D.ubyshire for their ilssiSlanCt! in
preparing the manuscript; and all the nun>erymen, private collectors and k<,<,pefS
of public collections who gave pennission for plants in their collections to be
phOlographL'<i. We would also like to thlmk Dr GeT va n Vliet, !'lant Officer.
CITES Secretarial, for the chapler un CITES, which unfortuniltcly had to be
shortened for rt'i1sons of spacc.
The plants illustrated are shown in hJ.bitOl\ ;md in the authors' collections.
"Tht-y hil\'c also been photographed in the public oollt!Ctions of thc Botanical
Garden of the University of Bologna, the Botanical Garden of the Unin~rsity of
Palermo, the Garden of the National Herbarium of Namibia, Windhoek, and the
Succuk'l1t Cnllcction of the City of Zurich; the private collections of Andren
Callabriga, Vivi,ma Didoni, Annarosa Nicola and Pasqua le Ruocco; and thc
nur.;cries of Luciano Cre"enna, Bergamo, Cactus Centre, Florence, Lalla
Pelliconi, Ra,'enna. and Anna Peyron, Turin.
All the photographs w('re taken by Pierfranco and Daniele Costanzo unless
otocrwisc slated on page 240, and without their help the publication of Ihis
diCiion.'\ry would not have bo......'Tl possible. Thanks IIlso to Pierfranoo Costanzo for
dr,Hv;ng the line iIIustraliofls.
Special thanks are due to Marina Di Stefano Saje"ll for her plltience nnd
support during the preparation of this book.
F OR E WOHD
The decision 10 write this dictionary was prompted by Cacti: The Illustrated
DicliOl/Qry by Rod and Ken Prcston-Mafham, which so successfuUy mel II need
among C"dctus enthusiasts. Succulents other than cacti are described and illustrat-
ed in St.>vcrai monographic ~tudi\.'S dealing with a single genus or a specific goo-
graph ic,,1 ~ion, but there is no single book with a Jarge number of colour
illustrations co~'ering a wide rllngc of genera. It is hOpl-d that Ims dictionary w ill
go some \V<ly towards filling this gap, helping collectors to identi fy their plants
and cl1 cou raging ncwcumcrs to scloo interesting species to study (lnd g row.
We would tikI' to e mphasb:c that we do not pretend to have found a solu tion
10 the chaos afflicting the dassifi(alion of succulent pla nts - rather, we are curn-
pletely involved in th.1.t chaos. The classification of both succulcnts and cacti is
affecled by several factors thai lcad to frequent change> and 10 the CT{'alion of
very small genera and new species. The main problem is probably t~ horticul-
tural value o f new species. Nurserymen often offcr unidentified specimens as sp.
11000 to stimulnte the potential (u ll(.'(;tor's interesl. If a plant is described and pub-
lish .... d as a new species in nccordnncc with the rok'S of the International Code of
Botanical Nomcndature but without the plant's variability in habitat having
bcocn verified, a new species could be established, which, although va lid in
formaltCTms, has no biological mcaning
It is essential that a group of Clfpcrts attempt toddine a system that will bring
som .... s tability to the naming of succulent p\;mts and will CUI the pk'lhora of
generic nnd specific names that a(f1ict~ this group of plants. The International
Organization for Succulent !'llmts Stud y (lOS) is aiming to complete a proje<:1
tha t will find a consensus among cactus experts on the naming of the Caclaceae.
It is to be hoped that the [OS will be able to carry out a similar proje<:t for succu-
Icnts.
In this dictiona ry WI' h,wc used Ihe names currently found in private and
public collC(;tions. Including available rcvisions would have resultlld in a partial
work with no cohcrent criteria that would confuse the average collector. Somc
revisions a re noted in the descriptions of th e succulent spt.'Cies and for those
readers who <Ire interested in pursuing the subject fu rther thcre arc book.~ lisk>d
in Further Reading.
INTRODUCTION
Sucrulcnts i1T(' defined ilS plants thai are able to withstand drought bcc.luse of
the water stored in some of their orgrms. They constit ute a widespre"d group,
~ing represented in severil! filmilies of flowering plants and, to some extent, in
the Gymnosperms as well. The definition is somewhat problematic OC'CilUse
several plants, although not true succulents, ilTC included among th eir number
for aesthetic reasons. The most precise scientifi c definition is given in Lift!
Strategies of SIICCU/ClIls ill Deserts by D.). von Willert, B.M. Eller, M.J.A. Werger, E,
Brinckm.lnn and H.D. Ihlenfeldt:
A succulent (or sUCCQphyte) is a plant po~ing al least one succulent
tissue. A succulent tissue is a living tissue that, besides possible other
tasks, serves and guaran tees an at least tcmpori'lry storage of utilizable
water, which makes the plant temporarily independent of an e)(ternal
water s upply when soil water conditions have so deteriorated thai the
rool is no longer able to provide the ncccsS<lry water from soil.
This definition implies that succulence may be present in one or more plant
organs. The specializa tion of an organ is determined by the particular sek>ctive
prCS/iurein the spccics' habital.
Leaf succulence occurs in several familil'S, and it is usually associated with
e nvironments that do not have a long dry SCilson. When the dry scason is very
long the succulent leaves arc highly specinlized. Litho/IS is, perhaps, the best
example: the stem is absent and the leaVe!; grow hidden in the soil, thus reducing
water loss.
Stem succulence, which is seen in the cactus family, is also familiar to succu-
lent enthusiasts. Succulen t stems have leaves that are reduct.>d in size or arc
caducous when well develo]X>d, ,!lId photosyntht.'Sis is performed by the grCCll
s tems. Stem succulents may inhabit more arid environments, but the critical
point is their sire: gig.lIllic spcctes need regular water to s upport growth, and in
very dry environments stems are reduced in size.
In root succulence the organ of reserve is subterra nean, which is thus protect-
ed from the stresses of wind and predation. Root succulence is often associated
with annual stems, which desiccate during the dry ~ason
In addition to thesc visible characteristics, succulents have severa Ilcssevidenl
features that give them strong selective advantages in arid habitats. The number
and size of the stomata (the holes that allow air to enter leaves and leaves to
cilpture carbon dioxide) are usually reduced. Not only docs this reduce water
loss but it also reduces the amount of cMbon dioxide taken in. Moreover, the
opening of stomat" during the day, in the presence of st rong sunlight and dry
air, would lead to the evaporation of water without a compens.1 tory intake of
wilter in the form of rain. A p<1rticular type of photosyn thesis, Cr.,ssulilcean Acid
Metabolism (CAM), has evolved in several families. [n CAM the stomata of
plants open at night, when the temperature is lower and the humidity of the air
is higher. These plants s tore ca rbon dioxide, using o rgilnic ncids - nilmely malic
acid - during the night when the stomata are open. Carbon dioxide is released
during the day when Ihe stomata are dosed but light is available for photosyn-
thesis to transform the carbon dioxide into sugars. CAM is not only efficient in
the economic use of water but is also highly effective in (apturing (arbon dioxide
from the air. CAM phOhJSynthesis evolvoo independently in several families,
and it is also present in some ;1qUJtiC plMltS, when thesclective advJntage arises
from the effiden(y of (apluringcarbon dioxide that is not abundant in water.
tn arid habitats suc(Ulent phmts may be the only SOUT(e of food for wild
animals. Several features have been cvolvt.>d to rope with this predation, the
most common being the presen(e of spines, whi(h make the pl,mts less appetiz-
ing, and mimicry, which makes them difficult to see against the soil. Another
defence against predators is the prcscn(e of toxic or repellent chemicills - on(C
Ihe predator hilS tilsled the plilnt il will not try to cat J similar specimen
The different adaptive characleristi(s are present in various groups of plants
Often spt.ocit.'S that Me distant in geographic or phylogeni( terms may look
si milar. The most striking example of this is probably the (olumnar or globulilr
euphorbia and cacti. The volume 10 surfil cc ratio is (Titical from the point of view
of storing water, and it is ~ible to identify a trend in stem shape, from the
cylindrical to the illmost spherical, th"t is the ideal for the volume to surf;1(C
ratio. These plants mily hilve very si milar sh"pes even though they are native to
differen t (onlinents and h.we different ancestors. The resemblan(e is caused by
the s.lme selective pn.'ssurc that (auSt.'S (onvergent evolution.

The problems of nillure (OnServation are widely ilpprcciated. Those who are con-
sidering growing, collecting and studying succulent plants must be "ware of the
impact that they can have on the wild populations of the plants they wish to
study. Most succulents grow in habitats that have a fragile coologi(al l'quilibri-
um, and the environmental conditions often lead to slow r.,tes of growth and
low reprodU(tive rates. For example, only 0.1 per cenl of the seeds produced by
WI'/wilscilin mimbilis will raise a new specimen, even in favourable conditions. In
the plant's habitat. the Namib Desert, filvourable mnditions may not ocrur for
several years, "s rain is very rare. If mature plants were to be removed, the
reprodu(tion rate of the species would fall below a sustainable level. Similar oon-
ditionsapplytoscveralotherspt.>c!<.'S.
In some instances the demand from mllcctors h~s meilnt that the limit in sus-
tainable collecting of wild spcdes has been reached. In response to this, in 1973
more than one hundred nations signed the Convention on International Trade in
Endangered Species of Wild Fauna and ROTa (CITES). Some mllectors regilTd the
CITES convention as a disaster. It must be borne in mind, however, that CITES
dOL'S not prohibit the trade in plants and animals; it merely reguliltt.'S and moni-
tors international trade in endangerL>d spt.ocies with the aim of preserving them in
theirh"bitats.
TIlt!re are three Appendi(es in which specit.'S endangered by tr"de are listoo:
• Appendix [ includes species that are threatenlod with extinction. The trade in
lmy wild pllmt or animal con taintod in this Appendix is forbiddclI.
• Appendix !I includes species that are not necessarily threatened with extin(-
tion but that may become so unless trade is strictly rL'gulatoo, and the trade in
wild plants and animals is, therefore, subject to the issue of an export licence
• Appendix !II includes those species in which the regulation in trade is within
thejurisdiclion of the nation (On(erned
Scver.ll 5ucru]cnts are ruITently included in Appendices I and 11; none is at
prcscnt included in Append ix Ill. All artificially prop<1gated plants m~y be
SUCCULENTS-Ti l E ILLUS TR ATED DICTIONARY

legally traded, even if they Me lish..od in the Convention. It is necx.'Ssary, however,


to check with local rl1,'Ulations to avoid any problems. A useful reference is nrc
EVIJ/,,/iOlI ajCrrES by W. Wijnstekcrs.
It is important to rem ember that even species that arc not listed in CITES m~y
be protco::tcd by local Icgisl~tion. If you are going 10 coiled any wild plant, first
check the local regulations and ask for an official permit.
Collectors Ciln plilY an important role in conservation. The management of
well-documented collections can help in conserving the genetic diversity of
endangered species, and propagation helps to relieve the pressure on wild
plants. Collectors should, however, always be aware of the damage c,lused by
illegal colh..'"cting. and they should never buy wild-collected plants of endangered
species, even with the aim of saving the specimen. The aim must always be to
s.we Ihe species not the indil·idual.

US IN( ; TI·115 DI CTI ONA It Y


The succulents included in this book are organizt!d in alphabetical order, by
genus and by species within each genus. Each dt>scription is laid out as follows·
The author who pobli5hro thc
prcscntnamc;whcrcapplic.:Iblcthe
Authur;'r..r"Tl'tlcenurnber name or the ao thor who originally
The family to which thctilXon bclongs described thisspccies in.:l different
Nameorthe~pt'Ci"lj genus Or rank isgivcn in br.lckcts

0350
Aloe humilis var. echinata (Willd.) Baker
LILiACEAE
A smaller va riety with fleshy spines on upper surface
of leaves.
Brief dO'SCription Qf Aloeechil1ata WiUd .; A. tllberCll iata Ha w.
''''''pOO~ South Africa (Cape Province) CITES App. ll

Oihernamesinu,;.,todeno\"th"
sameSpt'Cle5
Status of the species under theClTES
(up...:latroas~ tMan::hl992):
Rangeofdistribution of the specit!S
App.l =spcciesHsled inAppendi~!
App. 11 =spt'Ci..slistoo in Appt'nuix 11

The colour photographs werc t.:lken in a number of public and private collections
and comrnerdi11 nurseries or in the habitat. The colour and shape of individual
plants depend on their geograph ical position and the type of greenhouse or
glasshouse in which they are grown and on the attention they receive from the
grower. Plants also vary according 10 the time of year. During resting periods
they may shrink, hilve fewcr leaves or aClluire a reddish tinge. Plants grown in
sunny positions and plants that receive little water may also have .1 reddish
tinge, while those grown in shndc and thnt receive plenty of waler are greener.
For these r(,.:Isons the co100r and fonn of the plants illustrated may vary slightly
from the descriptions of the type species.

10
SUCCULENT FAM ILIE S AN D GENERA
The famili('S and the genera illustrated in this dictionary arc described bricny in this chapter. Some referenre books, must
of which nrc monographic studies, thai may be of use to interested readers are includoo in Further Reading. The most
com plete work on succulent plants is undoubtedly the three-volume A Hum/book of Sliccult'Jls Plants by 1'1. Jacobsen,
which covers Illust succulents in cul tiviltion, although it does not indud(' the most recent discoveries. Several periodicals
sp•.,'cializc in succulen t plants, imd they arc the best SQurce of up-to-date informa tion.

M;,\ V,\ CE,.\ E E.,dl. (.\ IQNOCOTl' L E IJ OM;) FurcroeaVL'Ilte


The 2Q or so species are similar to agaves but have taller
The Agavaceae family includes Soeveral plants of horticul - stems and longer leaves. The tall inflorescence is branched,
tural interest. It consists of about 18 genera, of which and the flowers arc campanulatc. The fruits conlain many
sc,'cn ;ITC illustrated in this book. The siems of the plants black SC«Is. The plant dies after flowering.
in this fami ly arc short or even 3bsent. The leaves are ~"T1!1 1;IU11ON: Mexico.
usually arrangLod in rosettes, and theyarelong-1ived,rigid
and often hlwe denla te margins. The rOOl5 are fibrous and NoJiulI;\olichx.
stoloniferous. The nowers are arranged on tall inflorb- There are ;tbout 20 species of small trees. Each plant has a
cenccs. Thc fruit is 11 capsu leora bcrry. succulent. swollen caudcx covered w ith thick bllrk. The
fibrous leaves are long imd na rrow. The till1 innorescence
G ene ril lll us irilled is branched and bcilrs numerous small flowers.
Agaut1.. DlSJl(Jlllfl"K»,l: Mexico to Guatemala.

This genus includes several species of use to humans. The Sallsevit riaThunb.
lea" es of Agaut s;salana, for example, are a source of sisal, There are approximately 70 species of rhh:omalOus or
which is used for milking ropes, S<lcking. insulation and so stolonifcrous plants with fibrous, succulent leaves. The
on. A. SiSlllllllR used to be cultivated ill Sicily untiI 19-W, but elongated innorcsceno.- is unbranched and is seldom pro-
it was not l'COnomically viable, and now the remains of duced in cultivation; the flowers are white or grL'Cnish.
~bandoned si5ll1 orchards may be S<'en. Other species arc Several species are tolerant o f shade and may be grown as
nsed in Mexico to produce alcoholic beverages by fer- house-plants.
menting the central part of the stem. D I!m< IHlJ110N: tropical Africa, Ind ia a nd Madllgascar.
The leaves uf agaves. w hich are very va riable in size,
are arranged in rosettes. The inflorescence is a ta ll spike-
1010 m or more - and usually has thick, modified leaves Between 40 and 50 species of smal1 trees bear simple or
(bracts). The tubular flowers, which are borne in clusters, branched stems. The leaves, which are arranged in
have six sepals. The fruit is a capsule wilh bl.1Ck seeds. rosettes, arc flexible to rigid. The brilnched innoresceno.-
All ngowes nre monocarpic - i.e., the plant dies after bears noctumill flowers. During the day the flowers do nol
nowering and seeding - but it takes several years for an close fully, gh'ing refuge 10 the moths that pollinate them.
inflorescence to be produood. The fruit is a dry o r fleshy capsule with black SC«Is. Some
DISTI!IHtJT1ON: I\ merica; a few species are natu ralized in species llre hllrdy and can be rn ltivated outside the green·
Mediterranean cou ntries and tropical regions. house if the cold season is not too long.
DISTRIDlJ110N: Nurth America and West Indies.
Cali/NlIIliS Rose
A mOllotypic genus.
D!STRJ8t.mON:Mexico.
/)QsyliriOlIZ UCC. M IAnl' U . IJ),\ CE,\E J .S t. lli 1.
(MONOCOT \' LE J}O NS)
There arc approximately 18 species of Si mple-s temmed
plants with long. lancrolate leaves with spiny margins. There are about 70 genera of bulbous plants, with corms or
The infloresceno.- is branched, and the flow~rs are cam· rhizomes. Only the genus Ammochari5 Herb. is included
panulate. here.
DlSTRI8UTJI)N: Mexico and southern USA.
11
the stilpcliile group pollen grilins adhere togethl'r to form
wilxy pollinia (Fig, 1). The frui l is a follicle con taining
About 70 g('nCrOl of IR'O.'S or shrubs with resinou$ b.uk <Ire
!;everal seeds with a Icrminal luftof hairs. Thegcnera con-
found in IropicOlI and tempcrMe regions of the northern
laining sucCtlk'1ll plant~ have a complex and confusing
hem isphere. The f,lmily cont,lin$ several species of E'CO-
taxonomy. The species with ~ucrulenl stems are prone to
nomic interest, includ ing M(lIISlft'rtI illdim (mango),
rotifover·watered.
AnflCflrdiulII OCCidffilfll.: (cashew nul) and Pi~tocia /culiscus
DISl"RJIIUTlO:-.l: lropical and temper.,It! regions..
(pistachio nut).
Only the genera Porllycontms Coville and Op.!rmiycarin
H. Perrier are considered ~uccu lcnl and arc included in
this dictiona ry.

,\ I'OC\'"\'\CI-:.\ E J,, ~~. (Dl eOT' 1.t: IW\S)

The family contains about 215 genera of lactiferous herbs,


lianas, shrubs and tTl.'CS. The simplc lca\'cs havc p.uallcl
~·eins. The nowers, whic h may be solitary or borne in d us·
ters,havefivcpctalsanrlfivcscp.l1s;lheblld5art'conlorl.
ed. The frui t is divided inlo two fo!liclt.'S. 5t'veral sJX'Cics
are sources of pharmacologi(.ll substil1lCCS - <llkaloirls,
gluoosidcs and so on. The plants belonging to Ihe Ihrl'C
gener<1 illust ra ted in this dictionary are very hmder and
need a warm position, and Ihey rl'quire plenly of W<l tcr
du ring Ihegrowing season.
D1STR1BllTlClI\::widl'Sprcad.
Genera Illus trat ed
Admil/III Roem. & Schult.
Succulent shru bs or tR'('S with swollen caudiccs and l<1n· Fig. I A dingrrwr of1/rejluWf!ro{"/>lanl irr tlr.:StapdiatgTOllp,
Ct'Olate leaves, cruwdlod althe stem apices. baS<'d 011 Orbea. P<II/t'n grr/i//:! adhert: 1~lrl'r to form waxy
DISTRJIIUTJ()N: tropical Africa, Ar"bian Peninsula and South poIli"ul.
Yemen (Somlra).
Gene ra Illustra ted
l'ac/rypodiumLindl.
Bro(hys/f/maR. Br.
Spiny trees or subtl'rranean, caudidfonn stems with
caducousleilves. Plants have luberullS or fusifonn rool~ with thin, caducous
~Iems. Leaves arc va riable in Sh,1J'C. The flowers, which
D1STRlUUTION: Angola, Madgascilr, Namibi., and South
Africa. may be solitary or borne in clustcrs, have a round corulla
and five lobes; Ihelobes may be free or unitl>d at Ihelip;.
P/umerUl L DlSTRlUUTIO:>I: southern and tropical Africa.
shrubortreeswithcylindricalstemsandbrilnchcs,l"ncro-
Caraihmur R. Hr.
late, d ark reddish·green leaves and showy, fragrant
flowers in variollscolours. Stoloniferous, clllmp. furmin); plants, with four· or five-
angk>d Siems bearing reduced, c.lducous leavcs. The
DI~"TRIBlfI10.'1: Mcxicoand tropical South America.
flowers are very variable in si1.c imd 1ll1ly be campanulatl;'
or have an open corolla; the corona h:'5 two whorls; the
five outer looc'S may be free or fusL'Ci, and tht! five inner
,\S C: LEI'I ,\I MC I·:,\E II . IIr. ( UI COTVLE IJU [\'S) lubes are fuSl.'Ci 10thCI)uler whorl.
DIST"IIllIlJf[or-: Arabian Peninsula, north and eastern Africa,
The family conlains ovcr 2,800 species of Hanas or low India, Mt'diterranean countrit'S and Soulh Yemen
shrubs, some of which arc succ:ulenl. The [ca\'es are simple (Socotra).
and caducous in most succulent species. The flowers have
five sepals and five petals and are ofll'Tl malodorous. [n

12
Ctrop·'s;aL. Dllvalrallaw.
These climbing or erect plants often h.1.ve swollen tuix:rs or The four- to Six-angled stems have spre.lding teeth. The
fusiform roots. The calyx has five Sl'pals, a nd the five leaves art! small. The flower.;, w hich appear on younger
pcLlls of the corolla arc united to fonn a tube, ..... hich is stems, are solitary or bome in clusters on s talks 1- 3 em
almost spherical at the b.,sc; the tips of the lobes arc uniloo long.
to fonn a lantefll-like structurc (Fig. 2). D1sTR18lJTlO:"II:eastemand .....esternAfrica.
DrSTlUIlIJT1ON:ccntral and southern Africa, Canary Isla nds,
EdudllopSisl look J .
Indi~i1nd Mad ei ra.
The stems hilve between 6 and 20 ilnSll'S,lhe ribs being
d ivided into hexagonallUbcrcl($. Flowers i1re borne in
clus ters of two to four; they have no tube, triangular lobes
andan inner whorl with fivereflexl>d lobes.
DISTRIBIITION: tropical Africa, Arabian Peninsula and Sou th
Yemen (Sorotra).

EJillleoII'llN.E.6r.
TIle few species have fiv~.lngled stems to 30 cm high.
Large, hemispheriCilI flower.; are borne at the stem apices.
DlsTRrBUTIO'l: Kenya, Somalia, South Yemen (SocotT3) and

FockeaEndl
These c,l udidform plants hllve tuberou s roots lind thin,
twining branches. The flowers, which Illay be solitilry or
borne in clusters, a re s tarfi sh-like
DlsTIlIIIUTJO'l: Angola. Nilmibia, South Africa and

FrereoDalzell
A monotypic genus very similar to Carol/lwlIl but with per-
sistent lea\'es to6cm long.
DlsTRIIIUTION:lndia.

Hoodia5weet
The miln y-angkod stems, to I m tllll, Me covel"(.od with
Fig.2 A side "ii'll! of lilt'scilcmatie f/oM'rofCeropegia. Tilt' conic.11 tubercl es and hard teeth. Flowers, which arc borne
corollalrlls/ifJe/lClals.wlricilarclIlliledl%rmmlllimos/splrL'T-
neiIT the apex of stems, have a flat corolla, very small lobes
icallll""lItll1t'ba54'wllill'llrt7Ii//s(Jfllll'lobt'sll~lmilt'1il/l(Ja
and a fiv~lobt.od corona in two whorls.
III/Ilerl/-/rtt strllellm. This type of f/UWfr call ~ ,1OI/iIlMt'd oll/y D~'IIII11JTIO'l: Angola, Namibi.lll nd South Afric.l.
bysptCi/icpoJlilllllors.u'lric/lllrellbleIO~tlclr t/rrpollillillduwn
Iltc/ollglu"". HoytrR.lk
These epiphytic plilnts hilve climbing stems and branch($.
Cy"mlchllm L The lc.wes are vilriilble in s hape and size, and in some
Climbing shrubs with fl($hy brilnch($. The enrollil lobes of species are thick and sucrulent. Inflorescences i1 re borne in
Ihe small flowers form a pentagon. pendent clusters wi th severill waxy, stilr-like and fragrant
DlsTRIHUTIO'l: central and southern Africa and Madagascar. flowers.
DISTRI61IT10N: Asia, Australia and ]'olyul'"!ii il.
Thl'SC arc epiphytic plants with smilll, lVilxy leavl'S, solne Hrr;:nriaR.Br.
of which arc modified into large, inflatod pitchers inhabit- The short siems, which branch from the bilse, are four- to
ed by ants (see also MyrmmJill under Rubiaceile). The six-a ng k>d and have large teeth. Flowers a rc produced
flolVers i1 re smaiL from the b., sc of young stems and have a camp.lnulate
DISlRlIll1ll(N: Australia and India. corolla.
DlSIl!111l1TlO.-";: sou thern and eastern Africa, Arabian
Peninsula and Ethiopia.

13
Notechidllll//Sisl..wr"nos&B1cck deep lobes arc triangul"r. The corona has two, five-lobed
Similar to Echidnopsis bUl leafless al all SIJgt$ of grow th . whorl s, an inner and outer whorl
DtsTRI1K1l'1(»II: lIQulhern Africa. DlsrRJIIU'IlOI\': tropical and sout hern Africa.

Orbolllaw. SlaprlilmlhllS Chaux


The plants al't.' similar 10 Staprlia and are often included in This genus is closely related to HI/emia.
that gen us. IJIsnu!:llJTlOI\": Madagascar.
DISTKIIIUTION: south ern Africa.
Tat"re5i~Wt:1w.
Orbellllllms L.c. Lc~ch Aow~rs wi th a long coroll a arc produced from the b.1se of
The stems spread horizontally. The flowers a rc very the s te ms.
showy a nd h~vea hairy corona. D 1STRJ1l11110N: cen tra l and southern Africa
D~I8VT1ON: southern Afri ca.
TricJuxQullmN.E. ar.
OrbtopslsL.C. Leach Cylind rical stems are s imple or branching from the base.
Aower.; are borne in cluster.; from Ihe baS(' o f s tems; the The small flowers arc borne betWCCl\ the tu bercles toward s
corolla is flat ,md there is no annulus. the stem ,lpiccs. The fI"t corolla has acute lobes.
DISIlUIIIJlIQII>: southern Africa. ~IIlU1ION: southern Africa. Madagal>Car and Somalia.

Padrycymbillm L.C. Leach TromolricJrt' I law.


Rhiwmatou s s tems. The corolla may be ca mp-lnulate or A genus th"t is very si milar to SIIIp.'lia and that was, in
flat. fact, formerly included in it.
DIS,'IoiIHlJTION:southernAfrica DISTRllIlJnON: southern Africa.

Pummlhus R. Sr.
The flowers are s mall; the corolla is flat, a nd tht' lube is
absent or campa nulat\'; Ihe lobes are lanct.'Ola te.
DtsTRl1K1l'1(»II:soulht'mAfrica.
A family of abou t six genera o f rh iwmatous, climbing
PSt'udoiilll('J$ P.R.O. Bally
plan ts with s im ple. often succulent leaves. The b ra nched
The unbranchlod stems a re stone-like. The inflorescence
inflOre5O!llct" bears small flowers. Only one gen us,
hasman ysm" Ufl owers.
BolIssirlgaliltin H .B. & K., is iIIustra tocl here.
D1!>'TKIIIUTlON:Soma lia.
D1STRII!UIION: tropical and subtropical Ame rica.
QllaqWl (N.E. Br.) Bnlyns
This genu s is related to Cllmllrmm.
DISTRIBUTION: sou th ern Africa.

Ra"iliollacml' Harv.
BO.\ IBACA n :.\I·: "'11m" (JH COTYu: nO 'lS)
A genus d istinguished by its tuberous roots, cli mbing There are 30 genera of very l<lrge trees with soft wood. The
s tems and loose pomn ia. leaves, w hich may be simple or compound, a re covered
Dr.muBlJTl()l\,: eastern Africa. w ith hairs or hairy scales. The flowers are large. The two .
genera illustrated havc woolly fruits with numerous seeds
RJly/ulOOlUll/Il P.R.O. Bally
~11IlI1"ION: tropica l Africa and America.
The unbranched s tems are papillose. Solitary flowers a rc
borne 0 11 short sta lks; the corolla has spreading lobes. Ge n era Illus tra te d
DISTRlBUTlON:easternAfrica.
8ombaxL.
Sa rroslemmllR.Br. These large trees o ften have spiny trunks and palmate
Thin-stemmcd shrubs with clus ters of flowers. with s mall, leaves. The very large flowers appear before the leavt.'S.
projectingrorolla lobes DIS'Tl!IOUT101': tropica l regions.
DtsTRJOIJTlO:'<.l:t ropicaIAfrica.
CllOrlsia H.B.&K.
The trees have swollen trunks that are usually spiny. T1le
lbere are about 100 species. The flowers, which are borne large floWl'l'S appear before the p.llm.lte leaves.
on long st,llles, ha\'e a five-lobed, osuall y fla t, corolla; the Dls nU BlJIIQN: tropical America.

\4
III HHI I·: U ,\( :E.\ E JII ~~. ( MO~O C OTYLEUO .~S) CO .\IMEI.I~J\( ; I·: M: II . Ur. ( MO~OCOT\,I.E I )O"\~ )

The family cont<lins <loout 50 genera of terrestrial or epi- These tropical plants have jointed stems. The flowers nrc
ph)'lic plants. The bas.11 Ic,1\'es, which m<ly hilvc spiny usuillly blue, iI colour thnt is seldom seen among succulent
margins, are oftcn arranged in rosettes. Flowers are borne plants.
in spik..'S "'ilh colouwd bracts. The epiphytic genera grow
Gene ra Illus trated
on the trunks and stems of trecs.
Dislribution: tropicaJ America. Cal/isiaL.
These plants, which have succulent leaves, are similar to
Gener~ Illustrated
TmdcSCfwlia
Abmmdlirlla Mez DISTRIIIl1ll0N: Mexico, tropical South America and south-
Small, terrestrial rosettes form large clumps. The greenish eastern USA.
flowers are usually SOl itary, and the petals Me much
Cymwli5 D. Don
longer than the sepals.
A genus of pl'lllts with tuberous roots and succulent
OISTRml1J·loN: Argcnlinil and Dolivia.
leaves
DeuleroclmiaMez DISTRlBllTlO.\I: tropical Africa and Asia.
These short-stemmed plants have rosettes of leaves with
Tmdt'SCauliaL.
spinose margins
These phlllts have fibrous or tuberous roots. jointed stems
DJSrRIBUT10N: South America.
and leavcs that arc often covered with hairs.
Dyckia5chuiLf D1STRIt\UTION: North and South America.
The stemless rost..ttes have thick rhizomes. The leaves are
rigidnndh.1Vespinosemargins
OIS1"RJ6llTIO:-/: Sou th America
CO\ IPOS ITAE Cisf'kl' (1lI COT\' I.EIlO NS)
HedlliaKlot~_o;c h

Stemless or short -stemmed rosettes arc formed from grey A very large fJmily containin g both annual nnd percnni"l
or red-brown leilves with spiny margins plants. ranging from herbaceous plants to tr~ and
DISTRfBllTIO:-/: lvlcxico and southern USA. including epiphytic and aq uatic SpOOL'S. The inflorescence
m.ly bear from one to mnny heilds of clustered, stalkless
I'llyaMolinil
flowers, and the calyx has many bristles and scales
The toothed and spinose leaves arc arranged in rosettes,
(pappus), which remain attach ..>d to the sa>ds and facilitate
which may be stemless or long-stemmed, to 10 m tall
their dispersal by the wind
wheninnower.
DISTRlIIlITlO\l: widespread.
OISTRIBllTlO.... : Argentina, Bolivia and Chile.
Gene ra Illus trat ed
Oll1ollilaL.
These small shrubs have either entirc or lobed Ic"vcs. They
IlU II S EII ACI':,\ E KUUlh (U1 COTYLEOOi\S)
are winter growing and should be kept dry in summer.
A family of about 20 genera of shrubs or large trees. The DlsrRIBl1ll0.\l: Namibia and South Africa.
trun ks, b.1rk and wood arc resinous. The leaves arc com-
Sl'IIrcio L.
pound.
This very large genus contain..,; several succulent species.
Genera Illus trat ed The genus Klciuia h"s bco:::n included in this genus for the
purposes of this dictionilry.
BurseraJacq
DISTRlBllTlO:-/:widespread.
These shrubs have caudiciform trunks and compound
leaves.
D!STRlBllTlON: Mexico and southern USA.
Commipllom Jacq.
Shrubs with a very thick. tuberous Colude>:; simila r to
Burscm.
DISTWIIIUTION: Namibia.

15
COi\VULVL LM : E,\E Ju ss. (1J ICUTY L E UO:"lS ) Gene ra Illu strated
This family cont"ins pl.lnts of very diverse habit. including Adromisc/ms Lcm.
some parasitic spl."(ies. The showy, c"m piU1UIJte nowers Succulent herb.1CCOllsnr shrubby plants with fit'!;hy, per-
are usually whitcorpink sistent leilv<'S. The innorcsccnccs have noweTS at right
DISTR18UTION: widespread. auglesto thepeduncIes.
DISfR1BU1l0N: Namibia and South Africa (Cape Province)
Ge nera Illustrated
ACU!lillm Webb & Herthel.
IpolllrJelll.
Shnlbs with simple or branchl'<l stems and leaves
A large and variable genus of trees, shrubs and herbaceous
arranged inrosettesilt the tips of thcbrilnchcs. The chilrac-
pl,lnts. The species cultivated by succulent enthusiasts
teristic rosettes die after nowering.
have tuberous roots iUld slender s tems. Several
DISfRlBL"llOK: north Africa, Canary Islands, Madeira and
spt'Cies' are being imported.
MediteITane.lnrcgions
DISfRIBU1l0N: tropical and subtropi cal regions
Aichryso!!Webb&Bcrthel.
MenemillDelllUit.
Very close to Ai!O!!iullI.
These sm,lll. climbing pl,lnts h,lve lobed or compound
DISfRIDU1l0N: Azores, Canary Islands and Madeirn.
leaves. They ilre very simil'lT to Ipomoea, from which they
may be distinguished by differences in the pollen struc- CO/Y/I'dOHL.
ture. Branching shrubs with opposite waxy leaves. The campan-
DISTII.lIILIllO\J:tropical regions. ulate flowt!TS, which may be red, yellow or or,lIlge, are pol-
liniltedbybirds,
DISfRIULmON: Arabian Peninsula, Nil mibin nnd South
Africa.
Ctl ,\S SU I ,,\( :I':,\E DC :. (I., . COTYLE DUNS)
CmsslllaL.
The family of Crassulace<"1e contains <"1 large number of Ht!rba ceuus plants or shrubs with succulent leaves of
widely found mlllual, biennial and perennial species, with various shapes. There are 250-300 tipecies, which are found
more ur It'!;s succulent leaves, which grow in a wide range in habitats ranging from wctlnnd to desert. The southern
of cl imatic conditions (from wetlands tu d~rts). Th ... African tipt'Cies are widely cultivated.
innorescence usually beJrs sm"n nowcrs (Fig. 3). The DI<;TRIIIU1l0:-.l: southern and tropical Africa; a few species
seo..'(\s are dust-like. but plants an' easily prop.lgiltcd from arewidespreild.
stem and leaf cu ttings,
DlldleyaBr.&R.
low-growing plants with leavcs arrangt>d in rust'lles. The
branched inflorescences bear star-shaped nowers.
DISTRIBU1l0:-l: Mexico and USA (Arizona, California,
Nevildil)

Ec/U!I>eriIlOC.
low-grow ing rosettes bear erect stems uf l<"1teral inflort!s-
ccnceswithnumerousbracts.
D/STR1BU1l0:-l: (t'ntral America and Mexico

Crap/opelalllll! Rose
A genus that is closely related to Ecitl'l't'ria, from which it is
distinguis hed by its sl.lT-like nowers illld the wd spols on
thepetills
DISfRIBLIlloN:Mexico.
Grl't'lloviaWebb&Berthel.
The rosettes are very similar to those of Srmll("roiullm. The
nowers :ITe golden yellow, and the rosette dies after now-
Fig. 3 Tile f/uwers of members of 01., Cmssulllcene [amily lire ering
,>erg simpk IImi lire anllllged in inf/oresct'llces. Tire Illlmlx' of DISTRIBU1loN:Canary Islands.
stllmells iseqlllllt()tItCl1umbt.>rofpetIIISI!riS<lmllltipleo{titem.

16
/oviborl>aOph: Tacitus Mor,ln &. J. Meyr5n
A very small genus, closely allied 10 5<'ml't'rtJivlmr. The 5..>eGmptopt'lnlulII.
flowers arc camp.1nulah.'.
Tylccodou T oclken.
Dls'ntllllfl'l()fl.: ea~t Europe (H.1Ikans,md ('astern Alps).
Distinguished from COlyi.'I1ol1 by the lion-waxy leaves
KII/;mdwt Adans. arranged in spirals and by the presence of bracts on the
A vC'I'y \'ariilbl(' genus con taining hcrb.1(COU~ plants, flowering stems.
shrubs and climbers. The leaves in some sp<'Cies produce DlsnU61JTlO:'<1: sou them Afric,l.
pl,mtlcts. The lenni"al inflorescence bears showy flowers.
Vililulinilaw.
DIsnl,lll;Tl!.JN: southern and tropical Africa, AS;il and
A genus that is dosely related 10 Starlm, from which it
Madagascar.
differ.i in hln'ing pct"ls unikod int() a di!>tinct tube.
M011Ill1lht!< llaw. D,5f1UHU1l()f.;: Mexico 10 Peru.
A genus of small pl,lIlts with fleshy leaves and hairy in-
floresomre;.
DI51RIIlUTIlN: C.1nary 1~lands and Madf'ir,l.
CU(: Lltlll'!",\CE,\E Ju ~~. (1) ICOT \' I. EI)U I\Sj
()ro;;/"chys(OC.) Fbch.
The small T05cttcs die after pmcluring ,1 1011 1in fl orescence. This is a f<lmily of f,1st'growing, climbing plants with ten-
LJISTRI!lUT10'i:Asia. drils. Se\,eril] species produce edib](' (m ils. '1111' species
cull i \'al~'<l by succ ul ent colll...:tors usually IUlI'e tuberous
Polc/1Y/1l1ylllm Link, Klot7$ch &. Otto
TO<Jtstocks(>T sw,,]lcI1 b.ls,!S.
The fleshy-leaved roS('itl'S may be distinguished from
DI!>TJ<IUU1loN: tropicfl ] find warm temperate countries.
[c/U!VI!riu by the pTl'sence of a pilir of sca l~ inside each
pctJI. Ge nera Illustrated
LJISTRlDUTlO'i: M('xico.
Cel'Jrn/<I)lelltalldraChiuv.
R05uiaria(OC.)Stapf A monotypic gellus.
The pl<lnts in Ihi~ genus <lfC similar to Sedlllll aod DlsnI,HLIT1OI\: Ethiopia, Kenya and Ug.lnda.
Srnrpm!iVlmr, but the roscttl'S 141Velateral inflorescences.
COl1lllOCllrplj5 Welw. ex llook.f.
DlSI1UIIU1lO'i: cast Europe and Asia (Caucasus and
A genus of ilhout 20 speci('S of pliln ts with climbing or
Himal,,},as).
trailing stems arising from a tuberous root. Very few
StdumL specics are of intercst tu the succulenl gmwer.
There <lfC approximat('ly 600 spcde; of hcrb.1CCOUS or DlsnIrHLIT1OI\: Afric.l, India nnd M,ldaga<;car.
shrubby plants, with erccl or dccumbt>nt stems. The star-
Garo,dllllllrusi larv.
~hap'-od f]OWl'fS arc various colours. European and Asiatic
A genus of plants with a tuberous rootstock from which
spcci('S are hardy and suitable for the rock garden.
climbing stems wil h tendrils are produced.
Dl5T1!lBlJT1O'ol: Asia, Europe, north Africa and I\'orth
DISTR18LIT101\:ccnlral,eastcrn,U1d stlulhem Africa.
America.
/wroilil:t! ~.D.Grcelle
StIllpt'rtJh",lInSlapf
The glabrous :.tems arc swollen at the holSt.', and the climb-
TIIest' small plants have leaves arranged in rosettes and
ingbranch('Sh a~·elcndril s.
white or pink flowers.
DISl'RIIlUTIO/\.: north Mex ico and sout hwest USA.
DISTRJUlJTIO'l:Asia(]-limllllly,1S).
KcdrostisMcdik.
5<:mpl"rlliVWHL
Theseclimbing plllnts have swollen, cll udidform bllS(.'5.
The lell\'es of these Sioionifcrous plfl nts flfC nrrMlged in
LJISTRIBtmOl>l: Africa to trupical Asia.
ro*lk~, anu th ... flowl'rs arc star-s hapo.'(\. There are about
40 ~p'-'("ies and mnTl~ than 250 cul ti vars, allllf which are MeiOlliriaL,
hard y. A genus runlilini n); " pprOximlltcJy 10 Spt....:il'S of climbing-
DISTRIIlUTlO'l; north Africa, Asia and Europe. or trailing-stemmed plan ts with tend rils, very f('w of
which <In' of int('r('St 10 th(' succulent collector.
Simxrassll/a A. Berger
DlsTR181.!11OK: tropical and su btropica l America.
A :.mall genus, :.imilar 10 Sed",.,.
DIsTR,6l1I1C)"II: A:.ia (I-limalaya:.toChina).

17
MomordiCllL. EU I'IIOIIIII ACEA E Ju ~~. (llt r.OT\'I. E IlO NS )
About 60 species of climbing plants with fleshy rootstocks.
Th(' Euphorbiacca(' family contains about 320 genera and
The fruit s a re more attractive than thc flowers.
o,",er 8,<XXI species of geographically widespread ph'mts,
DtsrRJ8UTlON: Africa.
which range from annual herbs to la rge trees. All
Sf'yrigia Kl>r;lUdren Euphorbiaccae have a milky sap that may be hannfu l to
A gCllus contai nin g a fcw species of climbi ng plants, with th(' touch. The innorescence has a complica ted st ructure,
slcndcrorsucC1.l lt'nt, little-branched s tems wi th tendrils. based on the cyathium, which consists of nn involucre con-
DISTRlBlJTlON: Madagasca r. taining one roouccd female flowt'r and sevt'ral male
nowers. Th('re lire protective bracts and ncctaries. The frui t
Xerosicyosl'[umberl
is a capsult", which explodes on ",aching maturity.
A gen us of cl imbing plants wilh gl~ brous or hniry st('ms
and thkk,su<xulcnt leaves. Ge n (' ra Illus tra ted
DISTICIHlJTlOl<l:Madagascar.
EupllorbiaL.
Zygosicym Ilumbert Tht'''' art' over 2,txXJ species of very di\'ersc habit, from
Climbing siems with tendrils are produced by tuberous annual plallts to large trees, and including several succu-
roo< •. lent species. All species contain a poisonous, irri tant white
D1STIt18lJ1lCY.'11: Asia and Madagascar. latex. Cyathia are endosed in a five-lobed im'oluCfC
J:xoaring nectaril'S and subtendcd by enlMged, colollrro
bracts (Fig. 4). Some spcci<'S make sui table house-plants.
DISTltI~U1ION: widespread but the succulen t spt.'Cies Me
1J1IJIEIIE,u; E,\E \)r"k,' (IH COTYI,EI>O NS) more common in Africa and Madagascar.
This family, which is related to the Cact.lcc.le, contains
:o;crophytic spiny shrubs and small trees.
Gen e ra Illus tra ted
AllmmdiaDrake
A genus of spi ny shrubs with ('rect, spreading branches.
OtSTltHIUl'ION:Madagascar.
Dirlierra&ill.
The stems ha\'e thick, tuixorculate branches, which bear
narrow leaves and spines
DISTRl8UTlOl<I:Madagascar

IJ tOSCO Ht;,\ Ct :,\t: It. IIr. ( i\ 1 0~O(: I )T \' I , E Il O~S )

The plants in this family have large rhi7.om<'S o r tubers


v.ithtwiningshoots.
DISTRllJUTIOl<I: southern Africa and Sou th and Central Fig.4 I" F.ul"lOrbinCfdtIIJl,'itlf!ornce"ctCOlisistsofwmpliCi/l-
America. tdf!ofl>trs/rI/Clrlr6clllltdcya/hia(siIlXlllar,cyathillm). TIII'SI!
CO/lS;51 of III! illoo/ucre (n sericsofbrncts), which Cllulllill~oue
Ge nus Illus trat ed reiluced felllllir f!mI4'r 111111 St'Vt'r111 ";IIIt' ours. Tile f!Ofl~'rs lire
DioscorrnL. IISI/ally ill(tlllspirliOlIs. l'ol/iuators lire Ilttmc/t~1 by l/ie prolre-
The very IMge, spherical caudex is covered wi th b.1Tk, tivebmcls(mOlIi{ielllcaves),wlrichcall/tcvcrysJll1W1j,llIl1lby
which cracks into polygonal warts. Scv(,Till species have lire ~'dum of /Ill' lire/aries. TIle illlls/mliOIl is ba ....t 011 the
edibte tubcrs, 3nd some have pharmacoiogicaluses. cyatllia of Euphorbia milii.
OtSTlmMJl')()f.;: sou thern Africa, Central and South Am('rica Ja/ropha !..
" genus of trees or shrubs wilh simple, palmately veined
leaves, sometimes coven.<d with hairs. The inflorescence
has man y branches and bcarssc.lTlct flowers.
DtsrICIKU11OS: tropical and temperate regions.
18
,\.1o,mdeulllmPilX Ge nera Illustrated
These succulent shrubs h(lv~ !>everill spt'(:ies with spirally
l'ellirgoniumL'Her.
tuberculate ~tems. InfloT\"'SCences nrc borne at the stem
A genus of low-growing shrub species, some with succu-
apiCl'5, lind th(' lobes . md glilnds arl' fuS<'<! togl'thcr.
lenl or swollen stems and roots. The leaves, which may be
DlsTRrBUTlOK:tropicaIAfrica.
simple or compound, aT(' of len aromatic. Some of the com-
mooly culth'aled geraniums art' included in Ptlargollium.
DlSTRI6Ul"J()N: temperate and tropical regions.
SarcvCllJllon{DC.)5weet.
The succulent stems of the plnnts within this genus are
covered with protective b.uk. The spn-ading branches are
spiny and ha\'e small leaves.
DISTRIBUTION: temperate and tropical regions; most succu-
lent species aT(' common in southern Africa, especially
Namibia

C, ES\, EIII Af.EA E n Ulllurl . (J)I COT \' I.EI H)NS )


Fig. 5 A tytliclll Monadenium {louII'r.
A family of shrubs, herbs or Iianas, rilrely tr('CS,wi thoppo-
PcdilllllllwsNcck. site leaves and showy flower.;. Very few spt'cies are of
A gl'llus of shrubs with scver,ll br,lnch('S :md smallleavcs. inteTcst tosucQllent growers.
DtsnW.lUTlQl\: Central Al11l'rica. DISTRIUlITION: Widespread.

PhylllllltilusL. Gene ra lIIus lra ted


Trees or shrubs that often hnvl' l('nf·likl' stems. The leav('S
R.et:hstrinrriuc. Kegel
themselves arc of variable ~izl'l>. The inflorescence> are
The species within this genus have tuberous roots, vch'ciy
bome from the ~xils, and the fruit is a capsule containing
or hairy leaves and flowers borne in short p.lnicles.
Iwoseeds.
DISl'RIBt.'TlON: South America.
DIsnuBIJnoI.,;:tropical~ions.
SimliJlgiuNecs
Sy,rmlrniu/ll Boiss.
The shrubs in this genus are tuberous and have opposite
A genus of shru~ with thick, succulent stems and large,
leaves. The flowers are borne al the axils of the leaves.
p<lle gl"C("fl leaves. The lobes <lnd glands are fused together
DtsnuBl.lT1(lI\: Mexico to Argcntin."l.
inloa single uni t.
DIsTRlIlUllON: tropical Mrica.

If.Af. I,'\i,\ c' I:,\ E ' li" T." (1l 1c'OTYI.E n ONS)
,,'OLTQ UI EIII ,\ CE,\ E DC. (O ICO'!''' 1 E J) O~S) The tI"C'CS,shrubs nnd li,lnilsin Ihis family have stems that
may exhibit peculiar growt h forms. Only one genus,
fOllqll;l'rifPH.ll&K.
Pyn7lucanllw Wright. which is native to ~astem Africa, is
This is the only genus of this f.lmily. There are a few
illustraled in thisdictionnry.
species of spiny shnlbs or tr('CS with stems over 10 m tall.
DISTRIBUTlON: tropical and temperate regions.
The gcnu~ Idrill is considerL'(1 to be a sy nonym.
DrsrRIBlTtlO'I: Mexico and southln'Stcm USA.

LABI ATAE JII ~~. (O If: On' I.I\ I) O\'S)

GEM \ 1,\ CJ.: ,u·: Ju u . (OICOTl' tEI)ONS) The shrubs and herbaceous plants within Ihis f<lmily
usunlly contain fragrant oils. The leaves are simple.
The shrubs or herbs in this family usually have aromatic
Several plants are of interest in horticulture, but few of
oils in gl;mdulllr hnirs, jointed stems and spirally ;lITanged
Ihem are succulents.
leaves.
DlsnuIllJT1ON: w idesp~d.
DlsnuBUl1ON: tcmpcrate and lropical regions.
19
Genera Illustrated QlstrriuC.-J . Duval
Stemless pla nts wi th succulent leaves th"t are arranged in
OcillllllllL.
roset tes in malure sped mens; younger plant s have dis-
There are abou t 35 species of a romatic shrubs and heTba-
tichous leaves, an arrangement that may persist in mature
reous plan~ w ithin the genus. Many spt.-'cics are important
specimens.
for cooking - O. basilicum COOsi]) is one of the best known
DlSTRlBtJTKX\: south('rnA frica .
cu lintlry hcrbs - or for medici"",l purposes. Very f('w
species are su itable for collections of succulent plants, Iluworlhia C .-J. QuVtl!
howt'Vcr. Planls In dwarf rosettes Ihat ]}lay be solitilry or clu~leri l1g.
DIST~IHunoN: trnpica l regions. The s ucculen t Ic'lv esan~vt'rydil' .. rs .. s hapt'S
DISTIUBtmON: southern Africa and Mada gascar.
Plt'Ctrullllms L· J-! er.
A s('nus of shrubs or herbs of which only a few species l'iJtllllil~ inUitewaal
havesucculenlleavt"Sorslems. A monotypic genus.
DtsTR!8UT!ON: Afri ca, Asia and Austral ia. Dtsr~I6U1IO'I: South Africa.

Xii/ilL.
A gen us of bulbous plants wilh linear leaves.
J)rSTlUBUTtON:A{rica a nd Europe.

A very larlie family containing over 16,000 spt.'CiL~ with


very dh'el'!iC hnbits. Severn! species arc import,mt as culti-
vated crops. Only one genus, Dolklros L.. is induded in MESDIIlIt" ,\ NT II EM ,\ CE,\E Il aili.
this dictionary. ( 1)I CO 'l'\'tl~ J)UNS )

DISTIIJ~: widtosprcad.
The family of Mt':ot'mbryanthemaceac consists of about 100
gt'llera with apprOXim.l tely 2,000 species, all of which h.1\'e
succulent ]('avcs. The species range from sn1.o,1I s hrubs 10
creeping and th .. extremely specialized stcmll'SS plants.
ULlA (: EAE Ju ~~. (MUl\O(:OT\ U ;I)O M;)
The f]OWl'l'S arc usually s howy and have many peta ls. Th('
A family of la rgely herbaceous plants but containing a few fruit is a hygrosropic capsule, which opens when wet, so
trec-likespcdcs. Scveral genern hnvesucculent lenves and releasi ng Ihe seeds, and closes when d ry. nlis adaptation
nre adapted to live in dry condit ions. is a response tu il rid environments; the fru it protccls the
seeds until W;lIt'T is ava ilable for germination. The filmily
Ge ne ra Illust rated
is widespread in sou th ern Africa, lind S<'veral species are
Aluc L. nil l ur~li :wd in Mtoditerranean regions. The classification o f
A large genus of plants wi th succulent leaves a rranged in the genera isb.l scdon the cha ractcristics of the fnli t, bul il
spi rals. Th(' plants are stemless or have short, rarely is possible 10 distinguish some by their habi t. The great
woody, Siems. variablity wi th in thl;' family has led to the proiift'Tation of
DlST1lIIIUTION: southern Africa and Madagascar. gcll{'Ta and spt..-'d~, a nd any revision would considerably
rOOucethcirnumbcr.
Aslro/oW Ui lewaal
[n this dictiOl\iuy the names currently in use in pri"a te
A g('nus containin g a few species; closely related to
and public collL'Ctions as well as in trade Cillalogu($ have
Haworlhin .
been followed. More than 50 geneTtl and 296 species are
DISTRlDVTION: South Africa
illus tra ted a nd described in th ... alphabetical St.'Ction.
BllibitlcL.
Genera lllus trilled
A genus contilining planls wilh sucl'Ulent leaVl;~S and sub-
terran{';lnbulbs. Alllillop;;is Sch "'anl~
DtsT'~ull.rno:o.l;sou thernAfriCil. ArKyrodenml N.E. Br.
AsptlZOll/tl N.E. Sr.
[riospmnum End!.
AslritfiuDinter&Schwantes
Solitilry or stoloniferous plilnts with globose, tuberous
ikrxcrllJlt/IIl5SCh .......mtcs
rools li nd with rl;'duO;'<i, ;,calt>-like !caws ilnd one o r mo re
Bii/jaN.E.nr.
well-<levclopl-'dleaf.
GlmlllnlhusSchwantes
D ISTlUIIl,m ON: sout hern Africil.

20
SUCCULENT rAMI Ll ES AND GENUA
Ct'l1lmlophylllllllN,E. Sr. \1U1I \ CI: \1 : Lin k (1I1I:On U :UO\ S)
C.·roclmllysN.E.IJr.
ON'iridojl<is~.E.Br. The MOT.lce,lc f,)mll}, rncludes about 50 geller., ilnd over
COI/Oiollylium 5c"hw"ntes 1,200 species which ar~' vcr)' dh'erse in habit, ranging from
COIlUloI'ylllwl\.E,Br. l'lTgc \1\....." to !omall hcrb.I Ct.'Ous pl<lnls, usu<llly with milky
CylillrlrujlllyllllwSchw.1ntt'S latex. Flowers <lTeSIlI<llI ~nd. in most gencr~, arc wind pol-
Daclyl()P"'i~ N.E. IIr. linated. (FirJls nowrr's aI"(- polli n<ltl-..:l by inSl..'Cts.l The
Orolos'~""IIN.E.Br. nowers <lTC };rouped in innoTl,,<\,~, with thickened aXI$
DmlrrrmllmsSchwJntes forming;)n inv.'gin;)lcd TCCcpl,)cle. The most extreme form
Dmm/lilil1/5 Dinter & SchwJ nt('S is found in Fiw$: thl' f/uwcrs arl' 1"1C1U<llly insidc the fig.
Drm:DII/JII.'IImmSch",.lnles ;)nd the pulp we cat is romposl...1 of hundreds of fruits and
E~'llm:ia Schwant~'S
[uargall/iI,·N.E.Br,
""",
DISJ"RllIlJTIO"\I: tropic.11 nnd tem""rate Tl'gions.
FllllrariaSchw"ntes Ge ner') IIlus tr,)le d
FtlJi"!ilmria N.E. Br.
GiN"h,,,,, llaw. OvrsttniaL.
GI.>lIiphyll,m, Haw. A );enus of l()w-);rt>wing plan\'> with slender stems ;)nd
HI7.-rrn lJint~r &Schwanks luber-like rhi;~01l1es. Inllol"('SCCI1Ces nrcsolitilTY and may be
/onllllliillil H. H.1rtmallll, regarded aSil fig inflOI"('l>«nce that is not enclosed (Fig. 6).
w/lJl'nm/lwsN.E.llr. DtSTRIBI,.:nO'l: tropiC"ilJ Afrka and Anll'rka.

wJ!"larIllSchwant~'l>
LrJI>I1IdliaL.BoI.
ulllopsN.E.Br.
Machnmwilyllllml.lk>1.
Mnll·l'horaN.E.Br
M,-sloklfIlIllN.E.Br.
Milrr>I'hyllllmSchw,'nt~'S
MOlJllar;nSchwan l~'S
NlIllIIJllllmlllms l. BoJ.
Nllllld!inDinter&Schwant~'S
NdiaSchwilnles
Nrolimlldn L. Bol.
o.luIIIOI'hofllsN.E.Br.
OpIJlnlllw/lJ,ylluw D;ntcr& Schwilnles
PlfiOSI'JlosDinter&Sch",,,ntt'l>
I'vlylllllllL.lk>J.
RIIbi,-a 'I.E. Br.
IVwmoo/1l'ylllllll Schw"nll'S
RIIsrhlllSchwanlt.~
RJI;;cllimlllm<LBoI.
ScrIl'IIIIIIIN.E.Br.
Schu'IIIIINIllDinter
SmirrosllglllnN.E. Br.
SlomalilllllSchw"ntcs Fig. 6 TIll" III('mba'; of liJ,' Momml(' fllmi!./f IIIII~' $IImli f/OU'l'rs
Tflllqllllllll Hartmanu & Lit-de III 1110<1 g,'lwra fiJ,,,,, arc !I'mli I~JI/mlllrd, alliJ,mgh IhOSt, of lit/'
1·"lIlIopsisSChwantl.... Ficus IIrt 111$«1/>011111111(11 1111' f/ml\-rs tift SroJlIIt~11II mftorN-
Trich()r/illr/rmIlSchw.lnt('S crt/c"". u,jliJ liurk/,lIrd Q.ft'> [vrmill,l( QII illl'tlXillll/,'t1 n'Cfjltoclr.
VOl/iI,·ml,'IIL. Bolo Tile mo;;l erIn'''',' fimu i~ I<mlllllll Ficus: lill' f/mwrs arc acllllll-
Iy ill<;dr till" f(r. IIl1d III(' 1'111" JI~' 1"111 is formed by Iwudrrd~ of
frJllb IIlId ~I~. III Don.tenin /1/1' illf/UrtSCtllCl' Ifllly ~ rrgardl11
tI$ n'St'mblm.r. /itl' IIIf/O"~'-IIC" of II fig Ilml j< 1101 r'IIc/osed. Tile
illll<lmlirJII i, btl<"d 011 III,' flml~'r< af Dorstcnia.

21
FicusL. Sesnmot/lIl/1rnrlsWelw.
The trees or climbing vines within this go!nus con tain latex A few species of spiny shrubs or small trees, with short
The leaves are simple, and the flowers are very small, with leafy shoots in Ihe axils of the Ihorns.
up 10 several thousands enclosed in a single receptacle DISTRrlllffiON;· Angola, Bolswana. Ethiopia, Namibia and
DISTlI.ll:IUTlO:'>!: tropical and temperate regions. Somalia.
UI/CI/filli/Stapf
A genus of nine species found in the dry to arid regions of
sou th M~d agascar. The flowers arc yellow or maroon to
OXAl.l Ilt\CEAE IL Br. (1l ICOn ' LEUONS)
pink.
The small trees or herbs within Ihis f.1Il1 ily have tubers and DISffiJUlffioN: Madagascar.
trifolia te leaves. The fruit is a capsuJe.
DlSTRI6l1T10N; tropical and tempcrilte regions.

Ge nus Illustra ted


PIP EIlt\ CE,\ E C. Agllrtl h (1l ICOT \' LE UONS)
Ora/isL.
The Piperaceae family contains species ranging from
A genus of stemless herbaceous plants with tuberous
herbacrous plants to small trees. many of which are aro-
roots. The trifolia te, long-stalked leaves arc usually
matic. The leaves ilrc simple, ilnd there arc small flowers.
lowered at nigh t. The flowers are yellow. Fruits explode
Pepper is obtained from Piper lIigrulII, a member of th is
when they achieve maturity, d ispersing the seeds over
family.
considerablcdistanccs.
Dr5TRlBlffiOO: tropical regions.
DISTRI6l1T10N: soulhern Africa and South America; some
spedes are naturalized in Europe and have become pt'rni- Gen us Illustrated
ciousweeds.
Pel'erOlllil/Ruiz & Pav.
A genus of small succulent herbs with minute flowers
DISTRIButiON: tropical regions.

PASS IFLO II ACE AE J u ~s . (1)I COn /LEUO NS)

Lianas, shrubs or trees with lobed leilves arrilnged in


spi rals. I'O BT UL,\ CACE AE J u ~~ . (1) ICUT YL EDUi"S)
Ge nus Illus trat ed This family contains more Ihan 20 genera and 400 species
of shrubs and herbaceous plants with succulent le,wes.
Adl!llia Forsk.
The leaves are entire and often bear long hai rs at their
A genus of herbaceous climbers wit h tendrils. The species
base. The flowers are small, but they may be very showy
of interest 10 succulent enthusiasts have swollen roots.
in some species. The family is widespread in tropical and
DISI"RIBlITION: Africa to Asia.
tempcrateregions.
Ge ne ra Illustra ted
ArracampserosL
PEIM L! AC F. AF. IUk (DICOTYLE DONS)
The genus contains dwar f pl,lIlts that have papery stipules
A family of shrubs or herbs with opposi te, usually hairy, covering the small leaves or hair-like stipules among the
leaves. The flowers are campanulale, and the fruit is a succulent leaves.
capsule, often armed wi th spines or prickles. DISTRlBlJTION: Argentina (one species), Australia (one
DISTRlBlITION: temperilte ilnd Wilrm regions. species), Namibia and South Africa.
Genera Illustra ted eeraria Pearson & Stephens.
A genus of shrubs and branches with reduced leaves. The
Prerodisclis H ook
inflorescence has from two to six small pi nk flowers.
A genus containing species of small herbaceous plan ts and
DISTKrl:ll1TlON: Namibia and South Africa.
shrubs with succulent, swollen caudices and tuberous
roots. The leaves, which have undulate margins, are vari- LewisillPursh
able in shape. Flowers arise from the leafaxils. A group of low-growing plants with fleshy t~proots and
DrSTI<l8unON: Angola, Nilmibia and South Africa. leaves arranged in rosettes.
DISffiIBunON: west to north North America.
22
PvrtulllCilL. \'IT AC I':'\ I~ Ju ss. (1JICOT\' U :DUi\' S)
Trailing herbs with opposit(' l('il\'es ilnd tufts of bristles in
A family containin g ilbout 12. genera and 700 species of
the axils. The purple or yellow flowers open only in direct
lianas with tendrils or small trees. Only two genera are of
sunshine
interest to the succulent collector. Induded in this family is
D1STRlSunON: tropical regions.
the grapevine, Vilis vinifem.
PortlllllrnriaJacq. DtSTRIIIUTIO:-J: widespread in tropi cal regions.
A genus uf plants with branches cuvered with succulent
Genera Illus t ra ted
leaves. The sm,dl flowers are pink.
D1STRlllunON: Muzambiq ue, Namibiil ilnd South Africil CissusL.
A gcnus of climbing pli\nts with tcndril s and oppositc
Tali'lIIm Ad~ns.
leaves. A few species hilve succulent TOOts and leaves
Plants with fleshy rools and annual, shrubby brandIes.
DISTRIBUTIO~: tropical and sub tropical [(.-gions
The flowers have two-keeled sepals
DIS1~lblffIO:-J: tropical regions. CypJrQstellllllll{Plnnch.) Alston
There nrc approximately 150 spt.·dl.>s of caudidform shrub.;
or trees within this genus. Thc lei\ves nre ~Iustercd (1\ th e
apices of stems. Succulent species 1Th1y achieve huge
IlU lIl ACEA E Ju ~s. (!)!{;UTl' I.I·: ])() "\'S) dimensions.
DISTRIBUTIO:-l' southern and eastern Africa and
A large family of trees, shrubs and lianas, although very
Madagascar.
few genera Me of interest to succulent collectors. Coffell
species (coffeeJ belong to this famil y
DISTRIBlffIO:-l: /\frica. ,\si.. and Europe.

Genus Illus trated WI·; I, WlT SI:lll ,\( :E,\E M;.rk)!.r. (' ;l'i\1 ,"\'()Sl'ElI ,\ IS)
MyrmcrodiilJack A monotypic family belonging to the Gymnosperms
A genus of woody s hrubs with large tubers, which are group, together with cycads and conifers. The pollen is
usually inhabited by ants in the wild. The tuber conta ins a dispersed by the wind.
series of cavities, filled with dead plant materiaL TIle ants D1STl!IIIl!T10": Angoli\ and Namibia
remove the dead milteriill ilnd use th .. cavities ilS n ..~ts.
Wdwitschia Hook.f.
There are several advantages to the plants for thi s symbi-
A monotypic genus.
osis, the main one being defence: the ants keep potentiill
predators away from the plilnts. Another advantage is the
re-use of the carbon dioxid", produced by the respiration
of the ants.
DtSTRIBlfflO~: Indonesia and New Guinea

STE Il CU I.I M~ E ,\ E II nr lH!. (])I COTV LE!)ONS)

A family of trees and shrubs, but with no true succulent


plants. Some species are cultivated for their bonsai or cau-
diciformaspect.
DISTRIBUTIO~: tropical regions

Genus Illus tra ted


BmchychitOlr Schott ilnd End!.
A genus of trees with swollen trunks ~nd entire o r deeply
palmatelcilves.
DISTRJBuno:-l: Australia and Papua New Guinea.

23
ILLUSTRATED A-Z
OF SPECIES
Abrome;t;dl~ IQrentzi~n~ (Mez) A" C<ls./
~,,~~,~~~~tl:,t~ Engl
...""...... ~t <.uJ~,. "1""""11 upw.nl. In.'' ..................... 1 m ,.11. ,."h Imdrils
~~:"~,"":,~'= ,':.:;:~~;:,,,...-~"'~;~~;~~;'" M-.dd~ilJl~IN ..... """'II"'... .... y...~ ...... flu"....,.,..

ArJ!"""'"
:::;:'~;:~~I'7:~;,.~. ~,.;I),o'
. )"; A"".-....."" H.ormo; A _I,..w.. ' Wt.

26
:"~~,~:~~!~~:~ Sdtinz
L",):;~,i,~"L>,&""U<"I""'"

~~~;~S~~~~;~~~S~~~
~~:'2.='~'~;;I~)~.~~~II~
Bo""·.,""SuulhMri,.
(Tr.... '·.l.ll)

Adcniakl'ramanthus
Ha rms

Ih ... k(d"d~,IS-IO<mm
~~'";,O:;':;;:":,:~,,!: ;:.~~; '"
roundPdk·.w ...;yello"'i<h
no"",,,,"

An,",""ti,"bul""f",u";do""fo,~Adt-o, .. f""nd",,,,~·er.oJ,,,,II,,..~io,,,,
~~':,~;;:,;. 10-15= in di.,moh". "'ith ..""".1 ~.fy ",oms; S"'''*''')''UOW

27
Adromischus alveoldlus Hutchison

~~j~b~~~~;o~\:,~~~;E,~~_S:;~,S~C~~Q"~~~J~~
Su"lhM'k.I(C~I"'P,,,\'\n,cJ
:l:~~~ i:~~~,scooperi (B.lkcr) A. Berger Ad romischushcmi (W.F.BMkC'rjrOC'lln,
K""""ocklh ... k;"""",,,·,h.ln7red·b,,,wn l•• ,, ... ,,-h ... h.,,· ,,,,,,,,J,.r'hw..
=;~rnn;::;:~7.~::;:'\:·;.;.7n!:';:h 'runW. I,'.,,,... ,m,bl.· In
nod fL.,,, 01">. C,.",iJen:d I" b" • ~,,,n ,,{ JI. JI.""',,,,,,, .r. ollli
~~~~~,:;,".,.;
,y""w""I'JI'c",,,,,',,' 1'"",lIn.;'" .""'''''''' l.'" ~m_:'" ".r"" ....."." U;r~w •• 1
Soorh Alrko (Col"'l'rmir..-,,) s"~lhAf..... (C·f",rr"\i",,,"1

29
Adromischusroaneanu sUilcw,1al

Ewc"I"'dybranch'I\I\"""";;;""'!\""'n 1",,,,.,,,3,,,,, I""S "','h nu "'t'"",,"


w"ym.,king<; IIO\.,..... ,;i\h'~I\""'",;t"p;n.bl'"
5ou'hAffi.:~(C.I''-·Pro\·Ul'-d

30
Aeonium glJndulosum W('bb & Berth.

R<N..1leS "ilh rhomboid.1 t.."","S','''·c"o.J by,.,r, ...;""ydl"w 110 .... ,""


;::;;::mryrr,;.... ",lIolk'

Al'Onium arboreum var. atropurpureum (\V.A. Nicholson) t~e::J:~~::;.~~~tinosu m (Aiton) A. BergeT


A. Berger
Lo"'shrub"'ilh"o.Jbr.,\CIl<"S.Ii);ht);"~... k,,,·<'S"ilhfi.,dy,iIi.I~ ' ''''I}; .....
EA~' ·n»<"l',.. 20<n,in d ... m"cr:k.",.,,d.,k
•••,,,,.'olmh;gh .... ilhdc., ... """ .... """""hrood;,;h,lnl""'.in"l".. ""'~ I.'<~nn;ndi.''''''" ••';;nllo,.,,...
~~lishIS"""' in '''''IYI'''''p«"i''I; in fJon:.<c<o'''''lo)O<m~'lI.ycllo''' ~~:.,~ cm tall; ~dlo,,' IOJ ",h'll' n"""",.

C.""ry l,Lond •. M =:"".u,.,li..-.:I(?) in ..., ...... IMoo;' "..,..""• ., rou.,It;.,"S

31
:a~~~~:..~~:dlryi WclJb & Berth.
H..... "ph."T".1 bu,h I" :10,,,, l"~h. "'lh,..:'~r.1 \hi" b'."d"~,,,,,,tj,,~ (" ,,,,,II
""'-it.". k." .. h.,;,.,. .....t Ii~ht!\""~. ,,,110,,"110,,"",,
C.",,,,,·I.I~nd,

AroniumsrdifoHumrit. &I'rou!.1 :a:~~~~:~~:thulatum l'TJI'gCf


~. "J i~<m tuxh. """~t.."ulll"'''''' ''''~'''''-'''''''''''''' Ie........·ith rod
..ri_yd ...... lk........
lu 8r~nr","" "'..., <m h'l\h ...·lIh ..... U....,<n~ Io:.,',~ )"' .m 10.>0'II; w,th OJliLo·
gi"'-""'!Ji:'"'' wi)"'"n."..""
Conu. 1,J..".J,ILoP~lm.J, T,""-"fll~, (&"''Y 1• Lond•

I
J2
L"w.""".wi,h L>.,!:~ ","",'~ ,,,50<m in diA_; Ie.".... s,...n; brAnching
",/Io""""'''''''t"60<n\ h ;Sh;)'<II,,wflu''·~rS.Th.'''''''tt.d,'''afl"rn O\wri ng .
.....,'"mMIa/'I..""m 1Io1l~; A. m<1<mI'l""" Webb
c.."'ryb"nd>IT~'''-·ri/~l

Aeonium urbkurn Webb & Berth A g~\"e ~rneric~ l1 ~ va r. Illcdiopida Trcl.

~=~',;'g~~~~~I;.b,,'1,i:"~h~~~/hl":h~~k~:~~'b:'~7n"~~~;;.~,:J~ in
CalU ,y ld.lnd.(f....,,;f.)

33
~~:::c~:I~llIdna Gentry
Sk<>n..,.lNnmM """'t<-o; g""... I""' .... 50--;0,", lonp.. with brown m.lf~lI," .nd
tk.:!'!"'Th,t:;:~~~!f;::.~~~~°;.t~;~rt~~,:;:;,:~I\~~~t,l~: r~~o:~,kr
~';::=~iI,,,og"''''A ms"""

34
~J:,V:CfEi!~era Salm-Dyck
<;I,'ml,,,,, ",,;pll,,, '"f0..5~m in di."n.~<.,.wi'h 1"<'1':11 ~h"o.s; num~I'O\" <hiny
K,,,,,nk."">lu25cm k",~ .. \d3cm w,J~ ",lh><h""I""".,,d fihf.ruu>
m.:Iq;iru.; inll"'""",,,", 103 m ",U;II"-"'I\ish-)",llow Ilo"w",

:;t:::"""" Sohn·[)y<"k

~!.:::~~e:,din~"di · rt'SiS A. Bc>rg('r ~!.:::C"~:~' i nif1 0 ra KerGawl


SmaIl",",-..
. ,.,J
".oir'>lk..J"''o\'<"'''13<mlon~''·''hho,ny
'I""" I" mm I..
bl.orl.l.~mi",,1 Inil"" .............•
n~
.. rip<Sinlo,,,<'fpa''
m ,.11; \·.nn,I,,,,,,,J
,,,~
SI<-mI<So,brdnchinll""""<$ofm"""hanIOOk-.>,-".IOSOcml""l\ondSrnm
w;d~ ,,"h4 mm long te""",.1 'p'ne,nJ fili~,mu ..... '1:''''' inllon"",,,,,, to
fIoM''''' lin~<-.I w"h ,,-.I ,or 1"''1'10.0. ("",,,,'1 <on, ~k",' l n" In b... 0 ",m'"ym oi 4 m hiSh; Ilo",,,,,.sn...,ni>h 1>.:low. Il".hc-.l will. ,oj .oo,.'c.
....,'.<!"'....")<,,,••: ~~~::.:ngr"I""''''''En&'lm
I M",ku CITESA f' l'li

35
~t:,~:c~.~:.e5b",ghlii c. Koch
S!emk""""rb..1(i~"""'Ul,>"',(hdJr~gn."I\,"'rung1yatn>NIo."",!040cm
~:::"nfl~oct'lOJm"II;I:""-'IU<h-bro"'n("purrh!ollno"•...,.

Ag~,'e k~rwi"skii Zucco Agave~leopoldii 110rt


St"""(o4mh'sh;,greo'lI,,,,"v""'toi'\)<mlongand4cmwid..,w,(hdArk
b",w""'rmi''''I,pll\''5<mIDn);;inllo.>"""""""(o6mhj):h;~,,,-·nj>htof''1o
~.... lIo,," tI""......
ASOt"h>urlR_;AromJomt.Il.:IIu.'r
M"",("<>
Aga\·emarmorat~RlI....:l
~~:-":c~~~i Engelm.
51,·"'k'»f_""''',1",inJ.,n''-"I.·' ..... 'h_,o-;o \c'J' ... ,olmk'''.~.mdlo ..-~, .... 50-80<01 in d;"mct~r; 11"-'1' '0 ligM II""'" IN,..,.tOJO('nI
Comp.!d
""'I):"" .•
3(I('nI"" de •• b,' .. •. Ih,'" rm.'(! woth,,,,,,'(! 'r,,,,,,.md h·rrr.,,,,,1 ~An<l ~1O('nI wid~: mf1on,scence 2-J m .. 1I:S"""".. ~·)Yllow Aow<"fS.
>Pi",,2 '<m 1""1";"(1",,,",,,,,,,.· ,,, " '" ~'II.);"tJ,·n ,~II~ " II"",·,...
~:::'I!"WI!.'k,.r :r:::~~~~:~~~~~:;~:'~:~:11
37
Agave potatorum z ucc. ~~:::C~..~idigera l£1lI.
rosct"" ,,·i(~ I\recn", )"llowish·!VN" ~~,.",,30--40 em kmgond 7 em
S"IiI~ry
~:';~rd~hg~~pig=':::=~~~;~7:,~,,;~~r;:i~I"=jJe
"""",Uml.ll;lrghlgn...,.,Iuyt'llo,,·i>hnu w eno. ;:,t:n~u:':~.(C=':j~~b)~l~hl~~t.'~~~':JIP'Ji,&,~~;Fm~:
=:rrrul<r.<uHook.f.' A .J<roIym~'KArw. ~;::lSapt'I~Drum'A.""'lrl.S.wal"'n

38
Agav~ strkta S.,lm-Dy(k Agavctriangularis /acobi

~~'~.';.":'~~,,~:.:?~~~,~'~;;;r~~~':~~~h.~~:~,~~~~~.~~,:::;~h
w;,.,,,,,,,i,,. 1,p' .... 2<mkm!>';nfl",,""""'""2m'.II;l'4'dt"purrli'h 11m,,,,""
~\o.».ironohu.Kln)

39
Agave utahensisvar.discretaM.E.Jones Agav~ victoria~·regin~e x asp~rrima
Similo'Tolyl""perlesbu'";lh,ligh,diff,-"",,,,",inrol"UTandf05ol'!"'h.ol""'
~:~~~T:':~h~V=~IC~~ .• y"""ym at A. "'ohtn ..... il ;"Slilt /uurul i~
USA (Arizon.ll
:~~?::;~ ~:thencourtianurn Webb Alluaudia dumosa Drnke

~.:;; ~~~:,,,,~:r:' ;:'il\~\~A:;~t;:':,~he<ffid;'\1I in.m,1I """""," Je..v.. A.ruubin\""juv...rule.t.l~eb ul ll'o"·ingloIOm 'I ""' lu ri l y."; lh .."·~",,, 1
=ngbrnnche<.aoof..... \hom<;leA'..,.v"'Y"""II.ndcoducou<;whi~h
C''''>yl.l.ond.
Modag"""c CrThSApp. 11

41
Al()!'b~keri ScotI- E.11iol

Grwn Wio,..,,&-IOcm \I>Ilg "'lIh ",hll,


.... _ _ ;"""';nK.,.,j ..·;lhrtdd ....
"""l.Jn", _rOJ W,"I~ """T"",I.",,-
l"n~

\.~., CrrESApp.D

42
Aloe brevifolia Mill Aloe cili.uis Haw .

51...,...,., m><1" .. S",n in dio""'l<>r "'jlh """"",1 olf,twx:.t. from "...., forming
"~'I"ml";I<'.",",6cmlo"g.'nd2<:mwjd~.tb.'se"·jthl;ttlc",n j,.t,,,,",n;
j"~n",.w'mt.n;l"le!i<".'k1il<"\·.,,. ..
III... ,.....~·'" 11.'\01
Svulh M",. (C>I"'I''''''i",d CITFSi\pp.1I

43
~J ~(~:~;ngsi i Reynolds
SI............. _·""""'mru,,_;',..20"",,"~r"'N ... rtl',hurtt«lh;
'...n..r-""""Io'iO'm .....~yd..,.,fkm·"'" =-,~cm"'::,'~"I:~,:r:~~~~~;'~';'.;~~~:";
\hd.>&O,...... CITE5i\I"P, II ~"'!::'.,..,.,j .. ;'h".h~,.,""";;n~'~<m ... 'I ...... ,~• ..,.._

crn~"'rr·1I
Aloeeru A.Bcrger

~~~::tI~~';;;'m~.:c.::.~~~:tI~~~:'A';,=:~"'~lt:~:~~ n::~';.
lII""oI'Y»i~;"'Il.ILo:r
I)!oop" CITES App. II

45
Aloehumi1i sv ar. ech i nat~ ( Willd-> Baker

"""" Ik-rv.lietywilhfkshyspi""""" ~Pi'<"'5Urf.\«!of"'. '·""


1\I««h""". WillJ.; A 1.,"" .... .,1. fl."
'iou,h Alricd (up<'l'rovlnul CITESApp.l

46
"1(11' mdoughlinii (hn)tl.m ~~.~( ~:~]Oltii Reynolds
*mk~'''r .1",,, _"',n"",1 """"'~;):'''''' I.".,...., .""",4(lom Ion/\ ,.·lth St''fI~20 25<mk>nl\.b..nchN1 from b.o..., k.,·",8-10cm km~.nd 7mm
:"'n:':"';.;!"!.,.,;'.."'n..'\.' ......... '>dm....:"'..II."1h.lnllo""""n<t'lmh,Sh. ~~7~:~",whi'" m...~i"l;'.nd ,,·h,le. c.ntl.lgi"""" ""''11, ...1"-'<'Ih; r,n~'"
~'f'" CITESApp.1l M.....I;""'.., CIT£SApp.1l

47
~~~c~::vu1a A. Elcrgt'r AlocrauhiiRcynolds

5rndllp"'nl>... ~'I"ryo,fumoinss"",ng,,,,,,,,,,blu"'h·S"'Y.lhi<:k.nd Oe>hy Sl<-m"",,<>T.h",f->temm..:t """"t",,10'm in di.lm•."k.,. fonningJo:n",,);f1><Op>-


~~~Ocm \on~ amu-d .... llh eanil.l~inous. white. m.J'lIin.Jllccth. rrod". ~"'Y1\'('('" le"v",- someli"..,. "ilh brownish IinSe.W<m lon~ 2 em "iokot
b...... . ndM'm ..ing"'"""" .. p"'nt.t.pt'X,.,.ithwh" ••JX>!".nd . m.,II .... ,.
AI"''''"'P'"'')'WiJN II.I'<"f1"icr ginal,pinn;influnx.'I'lCt."30<ml>oIl,,,,.rl<10,,,n.,..
M.daga."",,' CITES App. II )o.1.>d.g.>S<df un;sApp.II

48
Alocs.apo nanall,1"'. ~,I~~er~~ drrosa Ihlker
"UA(~"
<.mnJ.." n ...1"~.I,~I" ~M'" k"""~ 1~ 20"" long wah horn,. J.lT~ IPrown. Thin<icnl'.2Ilcm ~m~"'llh k>."·'... 'nl, ...."!"OiS('II'":I\I..'.'nl"'".'<~rm "'''SAnd
~lt.~.h",",,~.'Ik> ...·"'" 2cm ""I,·"·,,h'f"",.nd t."",I"",,1 <h."l' m"fJ;"..11,,·lh. ,nfkm'" ...,,,,.
...... M"'''''h;.I'',~((""I''·I'r'''''''"·.'.,"'I).Z""rul,,,.· ClTESApl' II
::~ ~:!::~.;;:,:;~;"" Cm;s;.pp.lI

St..ruo.."",,,,~,~ 12-lb I.·.,·.~; I'>rm"nl,h"");""" I~A\,...lI>oUt 211 '"'" "'n~


""h nun"~'~I"f"~' .1n,j rt"dd"h-\>n,wn m."~I,,",II.~1h; I"ln~ n.. "~,,
5om.I1... { 1 11~Arp II

~~:-c~~:nOS;'S; l11a I!orl. All)(' striata subsp. kara sbergen sis Gll'n & 0 .5. Hardy

~ hl"l>r>d. ~ ~",,,,'.,, ,1. om","'''''' ..... "h ''''m' ow,. I m hlj;h; I.·ow, u,"-'-'''''''''I>i'",'u'I''''''i",-d;fi<'''''·''I''I~'.'\d loppo.-d """h~n."'n.
l'-\lnn:''''''',lhho,n,.. ""''l'"",l 1"1h. ",.\I\):.""-.! [k,,, •. ,, '.m'~Id .'>ou lhAfr",.(C.f"·I''')\·'n«.'1 CfTESApp.11
OTESApr· 1I

49
Aloe I'era L.
~!~~.:~r;!~~~:~~:~~~~ L. BoJ.
~Id;':' ~~~';::~ ~"::::"h:Ct~~';~~~~~~:I~_dn~.~\,,"':.~":~~u~-
ETect..1d""""lat<.gJ<Iucou's"-",nl"",v,",2<rnlongmvere.lWl,h",,,a ll tubo>r-

,-atronim,lSplumna<.... ht.tp,op",h""
~~""=,e;:,~~~::ts,Ydlow n<> ....,,rs2.5crni"d"''''d''r
Ah-I»rb>.I."", Mill.;A onol(1lRoyJe;A . l.o""","Tod.;A . •n./}<orisLlm South AlricatCA tv;lU.llJi.'nc,)
Gonaryt,land •• CaI"'Ve,J"J,t.nJ.;w,JclynatlJ'ah,O'd 'n'c.".,P<.... ~·n"Sliln.
CITESApp II
50
Aloinopsis orprnii (N.E.BdL.Bol
..> >£"~.V~"''' '''~U-M
=~~~~~,~~~';'~'n ",0\"" 1S-20om 10"1\ wah Jdf~J"",. yellow

"""""'~""~J"'''' L 8,,1.
Swth AI",. ("J'o' J'r,w,,,,<,!

.1II... ,Ir ........",,,..,.,iLIJoI


SwlllAln,.,'C.'f"'" 1',,,,,,,,,,'1

=
~:~:f:!~ ~,t:~~::f;,·LloI.
~~~7n:I?;~.~::;~~~;~~:~;,::Vp-::r~ ~,~;~1ot;<:~f.':~,~~: l~~;'nd
)'do~·!I""""25cmrndi.1I""tcr
rd""L'''''~'I,,'''t"LBd
So<lh ... tn<. (C, p"I ·""·, ....@)
Anacampserosalstonii Schonland

51
~:~~:~~~~! crioit~ Oi nt~r ~:~~~~AS::~! papyr~cea E. Mey.
:;'~:~~i~:r!~': f5c;;';m~'~~"~f'jt~K~;i~~k~,f.~h\:::'~::'~ ~~t;I:~~~!h ;;~,r~"t:'~~o:~:,"~.~, Ion~.11<1 1 <m Ihkl-: I,,,m.
"'o",ibi./Suuth Af,ica (Nom"4uol.ondl CITESApp_11 "'.mibia/~th Alrico IN ......qu.l.nd). South Arne.. (K.]f"") ClTESApp.1l

52
=". .
s..·~
~~; r;'S lelephiaslrum OC.
...I .......,..105cmh,!\h.,i>in~f"'mlhid.<:n..,J1\>ct;8n....,nu,bru""ni,h
... V... 1 <m longdnd !\mm wid ... ""...,....J Wllh briSlly hoi ... , pink flower.;.
Argyrodenn~s(huldljiSchwan tes
"':S"''' ~~l"''i'; THl!d"n''E
;;:::~~ pl.n",: l<"3vl... 1_2 <m Iong.nd 'Inn wId., n",nd.-d • • "I"''' pink
Soulh Alrico (Cop"Provioce) CITESApp. U 5"ulh Afnr. (C'P" ['''''"'n<t")

53
54
~~~lj~!:~:~~;t~~~~ t ~.
Bowil'~ volubi li5 I I.uv.

:::::.."' ~ blN"""J"" Ioo~~rod 1.2",,_~,...uowno..-mJ.s<m'"

s.:..tIoAf".. tC••,,·Pro.... ""d

55
~~~~!~~~~~n~m.eum (Schlfr.) ~.E. Sr
~d:..,,:,r . '::"
. ..~~I:t:>=:~1:..'!-'ri=-.~.. _5 lOem""',,-
South Afnca f l ... "'.· ... 11

56
57
Car~Jlum~ IN'lraea
1..aI'TJnos
.....:;ll P'AU .. U ...
Fn.,,~ybran<~.·'''I!kd,

~~~~k~ft't:~~
CI:<"Ih; ..... ILc.d""""·Srftfl
IN ...... no....... d.<l:brownor
y.-I1noo.."'",,·,thbrown dob.
s"udl".obio.Ymwn

Card llum~priogo nium


Schum.
"S("I.rr'ADAn...
Drown .....".2IJ...Wrmlon&
.. i'~4"""e.ng""bN""g
W'I'" .. 1h;"'.'·"""'~mm
lon!loflo....... purp"' ... II~

~':::~::'"rI.ol. Chic,.
EIIuop ... K.:to,~,s.....wli.o,
T.", .. nl..>.Ul\Ond.

Cu~l1um~ socotr~n~
(8.llf.f.)N.E.Br.

F"",,,ylmlnch'ng.~·.ngIed.

=~~"::~~I:;:=
...."h,d.,kr.-dno..'"-
C."'II~_,.".,."It""N, E. Ik"
C.n' .... ("hKlV
~Ihk>ri •. Kony.,Somalia..
So ut~ v,,""'" (So<<>I")

Cmllum~lunlii i\'.E.Br.
~,u'r'AI'~~l~l
c..... ~,~. ~ .•,.p.-d .kn\> IO-:!Ocm h,~h ",lI! f<'<l do,...nd conlr,l-n;
1Ioo<...,~.)"ftIow>!_ .. nd",""".....bonT•• ~byM.G.

=y::V""''''''~~"'''

59
Carru.nthus ~Bii L B()1.
~:~~A~~:!~ ~gmiftr. C Fisch
""'''IIIYA'llll''''UA(
""""....,"'~ SI>c>n.br;uw;"'-"oI •.".,."... ""~<,,,Iurqi;."""'"""""""'·ud
,_h Oll ..... IlI'M,'....""'·o..w.n,
••F"""""""
s....lhAfriuK.".. Pr"'"ntt)
Ccmchlamys pachyphyllal. BoI
~:~,~;,I:~hj,~I,~~:~~~~~le L. Bol Mo,;sl"."'.\Nntl"AllA'
I..<>w-;:.ro"",~ rl,,,, "'Ihcl<m);ollod b"'ncb,-,>; l"'vn7_IO<m 1""Il.nd ~ mm St..-mIt'S>,<1ump-lllmllng I~.., ... bCm lung in 2I'<'i,.... purpl,-.w nO'WD.
.ro.. .... """'·"'I~"";pu'1'l ... plnk fIo"·~,, So.,tbAtn co{Opo"l'm,'inno)
5ou'h,\i'~.IC,po.·I'''''i''''')

61
~::r~ir~~~:Ir:~ndii R~nh
F.JW1 or rm''',lle S""'" "','m' m_l~cm lonj;.nd 2 em Ih,ck, ",lh blMx_,poI>;
...><Iu«",..... ,...,.5 7"""Iu"~~""'-~""nnu,,,,",,
\1"d.S"" .r CITESApp.1I

"
~!;~~~~;~~~liC. Kotschy
;:~:::,~~:':::~~\\~':1"~~r!:t~1~,r:'':''bn''!.~ :~lucktoroN roios;
K...y•• Sud.n l'TTE5App,U

63
~:,o,~~~~:(~~~ersollii JAocr>e. ex Hook.{.
Sucn,I.·",. ''''ini''g <I~m" I"".... I 5<m Ioo~ .,nO 3cm w;J~;I\"-.... n 110".......
\\u""''''b''luc.50u.kAlricd('1.....LTr.\l\,. ...,l) CITESApp II
Chriridopsis brown ii Tischef ~~i:~~~~H~i~~~t!~cra (A.Bcrgcr) N.E. Br,
~~["~~YAN1l<r'IAl!'~f

~~:~t~2~r~~~~:~:'; ~~:::~\~~':">~cr::~:~~cm
SouthAfn<~(C.I""'mv;~)
Cheiridopsismeyeriv ... r.minor N.E.ll r
MF.srM~~~ ~ "TH"""'cr~.
5imi1.>r lOlhelyl"'.pcci<sbul.m.ll....
SvulhMnc. (C~p"rru,joccl

~~s~:B:~~i~It~~r~~~.E. Br :~s~a:~~~It~~~~~£~: L Bot


~":~.n";grcY·5"..,nl"'v"'w'lhcl.'kerclo!llbu,.,..,j"2pa "~;Y"11,, ...
..
~I~ Ci!"'.!:'!!;,T~'t~;:~~~~; ~.~~~~::.Ie!~ ~t:::.=;.~
yellow How.. ",.
N.lnibJa/Svulh Africo (UUlcN.' .... 4ua!<lndl
Svulh Af,ico (Col"'l>""'jnn:)

66
Ch~iridopsisplllw rule nt ~ L. BoL
wU[WUIA'fIID,v.CEAr
Si>orI .... ncM "'i'" 2 f'"~oil"ll~~,,'''n luv.. ro""..,.j with mmut~l"'p'I"'e;
~Jo~''''''''''''''
s.,.",hllfoic.ICap<'I'"",inc,>1

W''''''.'A"m.''''UH
500rt <I"'"""'llh ~ '·~krl)". uf1l'<ju.1 ",e<l kit,-.", ';-' <nI kmg; }'d]o,," fiuwer>.
Sou,h Mri<"" (C'p"l'rm',nro)

67
Chorisiainsignis Kunth
TI"L~"" 15m ",1I;bottl<~

f~'X!,~~~~..!':',~~;O~m
~~'I~~~'~:!u';:~·:;~;~~'fS·
''''''r<.,udOci(orm~PI"w>n<~

"rs~n'i""

Cissuscacliformis Gilg

Clionbint;~.lJr:;'dnglru st"""
4 5cmindiam"'.........-'m.

::::::::.."'I~~:t~~~~rz~,'i
liou,rooI$devdop.lrnv,,"
<.ldun-.u>;i n~ ."",,20cm
kmg; Y'-~Io"·i'h·~"""'11I1u"'~ ..
<;ou •.....".r>d l mpwA1Afnra

CissusolearacuL.BoI
Subt,.... ,_n ,hi,,,,,,,,,, 'n
~,~a:~:~~~".:':=;;
>1"n .. to60cmlon~l<d'·"
20cm Iong ....d 15..", wide;
inflo~br.l>Cht.'d:S""-·n
ish n""'<.'f'S"'ilhpi"k'ips.
Southo:rnAfn.,:a

Cissus rolundifolid {Forssk.JVahl

68
~~~~~~r!~UT~:['::;!:US (I\.E. Ijr.) Schwantes
St,"",,2Q.;!O<mhi~h;2(yp<.""<oflc.w ... llyp<J.7<"111Ionf\andaJmoo(<om.

t:,~:~~~~~:~h":t;':lt;;,~;~: A:"t;i::;;:J-:i~:~in~;:'kll..;,·~:
)... llowf low.~
MII">rIr\tII"'"I'",..imu.NEBI
.... UlhAfri,~(c.!"'r1"(wln ....)
69
~:i~'~~:!!~~H~~!~:: L BoI ~~;~~:~!~~"!:t:;:~:n;-.l.E. Sr.
C......,.oboowni ... -5"""'bod .... 10-15mmlr>n"'w;!~ n u _ ...... lldol .. ObcorucaLbod ... 2<m!uAhondlcmu.,J ..........,.f<><....... "',....,lft.>b,butioI
whmoh-pinlcfloo,.oM. lighIgtftn...,!~ff'dd, ... ."...... I"'''.nd_<p<>01N'''I'.,...(IowI.,.....,( .. _

SOUlhMnc.(C.....·p"',.. ,""') _nO"'Uu..p«~IuC,,'''J''''r" .. ~E.Br


SuutJ.Afn<a(c..""r'mv."",,1

70
~:~~~~~~~,~:~~:~m L. Flo1.
Up.,Ssubron",.lbo.>.J' .. m rlu mp' .."ft... lvetybud;.... 2-.3 cmindiAmet.".
. lIIll.scmhigh;wh;",n~,,·~, ..
<outhAlnc.IC. f""I'rom....)

::~~~!!~~,~~S!::A~.E. B
r COlloph ylum exlradum Tisch .
.. f.5~ .. ""yA"Tllf'tACfAf
:=01 bod,e<1o 1 rm in diom..,,,,., green,,;,h d;o,k ~"'''n do ~ while 11",,,.. h.tpcd ,d;t' k.gbucoo ,g,,,,,, n bud i cs2 _)< mlong. n dl _ 1.5=wid~.

~.., ! hAIn<.(C~p< Pro.,in<c)


x::,,~:~ dr:'~YC.t\~;;-u~··;;:/o~~:k°.~~~':; ~,:!~pociet<m.lY~
Soulh Afric. (CaI"' !·r<Wi..,..)

71
~~S~~~~~!~;:'H~~~:~~ns Schwantes ~:<~~~~~!~~I~a~~::~N.E. Br.) N.E. Br.
Sm.ll1sh",bt<>20<mhi,o;h;'I<'1tI<""ith."""'di~b,anches.~,,,,-'I1bod"'" V.n.bl.bodi"'(whirish1l"""".gl.L><"OU<g'''''n.y.n''''·1l''''''''''ptnk)1.5cm
15-25cm m di.""",,", with liShteTdm.;ordnS.fio ...... ions.nd2cm w,Jewilhgn:yJ'J";nws,",,'" now",",
'bmibio/!:ioIJ.h Ame.(Wltl. N.m.qu. l.nd) Sou .hA frk.(C'I"'I'ro,· \~)

72
~~~:~~t!~~:,~~~c:::pense L.1301 ~~~~,~~;!~~,~~:~~~~des L. Bol.
\\1ut"",,.~""·n '0 pale g"",n bodi,.,.'o 1.5 em I""fi\~nd 7 mm in di.l""...,,, RuSly brown 10 y.. llow·S' .. n bOO.,. 10 15 mm long .nd 8 mm ,,·i d ~ ,,"ilk fine
,.llmdk,,,",,, IY,,,,.nd w ind,,,,.. ; O,,,,·,,,,,purple
\ .I.. ko(Bu>hm. n1,ndl N •• mibi.> (H" .h..... nlandl

I
73
~~<~~~~!!~~t~~~I:~~-E. Hr.
c.·.""p"o..... p ... nt;2.lob<-d.k""I,-d.s'''''",u w h''i.h"'!\"..... booJi<;l..3cmhrp
art<l2<m In d •• ",""" <11'-""'(/ withd",s;"·hit"o)'<'II,,"'II,,,,~......
Sou,hMrka(C.l"'l'ro,-;t>C<"

~~~~~~!!~~t::::~~~:tum Lavis ~~~~~~!!~t~:~~~I~mJ L. 6(11.


~~~:i.~t~O~;~~~~~~':':"n\ in didm""" usu~II" do"",l), A,."ri.,n,rnC.mty<T"d .. lln s,,,,h,-dby.,,,,,lIr"'UfC,,OO''''dUI'1"''U'.
Soulh Afnc. {C.,P" l'f<wi1l<"<')
".m,b.. (Bu.h",,,nl;>,,,j1

74
Co n(lph}·tum mi llutum (l1~W.l N.E. Br
~('I\t •• '''''n'''' t." ~~s~~~!!~~:,~~:~~~:llUI1\ (Haw,)N,E.lk
8!ui>n ~f\'ffi!u.l"" '" 12 mm ~i~~ ,,,oJ 1<m\\1<I,""""~,,.,J w,lh >put>; pol,'
I""llo",jj"n.m.>~'''Il,r>''·lvpu'''''·h,'~.'low ... ~~~~c~~:\~'::~~~~~'<~~':~;::"bl~Crl~~:r;:~'!~:I~~::t:""'"
S<oulnAtr",ll"f','I'n"U1,d
South AfriCd (CaI"'I'ro,·,,,,,,l

(onophylum mundum (N_E, Br,) N,E. ilr.


~1->""nA"TH~"M,~"l ~~:;'~~~;'!~~I~~:~~:~or Tisch
Gr.:-'"j;1W7\ bod,,,,, 12 mm h'6h wi'n mno<lu<i!n, d.,..; yellow fI",,·~rs. Green "'brow"'""',<ylindric.>l bodi<:> ,<> 1""In in oJ;'m.I., wilh "ch,~ md'"
Soo" th Alrir. (C'pI'l'<O"i"""J i~g<;"'m.lin.o/old"""""p""';""',,';wltii.'op;n\;nowen
N.mibi,/SoulhAfric.> (Liltlo:-l"''''''lu"I"",O

75
~~S~~~:~!~~(~~7:E:~rum I. wis ~~~~:~!~~,~~,r:'~~~.E. St
I'.leg""," bod"" d",.]o.'d inl,,2 .horl_~nd "".."....j wi,h d~rk .. dOl .. Shit.r';"",:~~I>g~. p.ok!~"-"'n'oblu!'h-&""",bool; .. 1-2cmi"d;"_
rinktowhit,·llow,.,..
SoulhMri<'l(Capo'Pruv,,,,,,,) "",u,h Afri<~(CAl"'"Y(... i""'J

76
Conophytumruschii Sch""antes
"n~'MUU'''TIl''''~('''f
A ..... lltorm ... C""'h..... '.. r>.£ 8,
".mit...:5ouIlIAIno:~ll.Jnl<-'.rn.>qu>i£".n

~~!'~~ ~:,:~~!~m TISCher ~~,~:~~;.!~~,!~~~I,~!~m L Bol.


c...~ .... I.-:z.~<m"'Jth .nd "em,nd ... """", .. "hdo r""" lI..,..,.dOb.: pon~ e ........ bod"""" I~ """ k"'K~nd I nn 'nd........... ","~ d.r~ dot.and I.......
~. \" ...... nfc,rat."" _~II, pon~tu""'Jl"'lt.lno..~Ar... mufC.',"""" '-t.o,,'.I"I.E.8r
s.,..' hA./ncoIC• .,.,rr."'..... .

77
~~'~~~:~'!~~H~:s!:r:~~gi~num lisehe!'
Ughlg"..,.. b.>docsl .. 2.!ICII\h'dI~0>d15""", .. d"'m<tn"·;'h~
~~t!~Z=","'~,n~ {~~tt..>ll.>m",.."'~. ...
5c>u l hAfric-~(C~ I ., .... Di$tn<l)

~~~~~~~'!~~H~~~~o:::e (Haw.) N£ Ik
Cn:y·[U ...... l..",Jd'''''bud....
l< ... hll.h.O>dwid .. ''''hdotl~ ....... Of~
doI..nd .....1""< ,,·h;tW,·)..tktw fl<>wfor<.
Soulh Afrio:.o (Cop<'"/'moI"""",
Conllocupussp. ~~e:O:::;bicul~l~ L
f ........... _ ~I I) I S"''' L ndi.uro<tc< .. ilh <lLmbLns>l<_'''''ryot. .... SbrublOl.5mhigll;lhi<k ...........llh ....... br.nd... boNori "ll!N....... 'ipt.
aod ....-...; ........... 2-'i<m Ior,g; ..... U.U""'" Ro..."ft$ 1e" ..... 4 Uan~I"6cmwidclU>d"".·......,.t"'lhprul_w.>.;lnfk>ro,o.
T~.IAfric-. <'ftIt"I''''1II<m .. ll w ithy.lI<>wf<.h· ..... fIowt.rs.
CdyIrdoot ..'. I"'w.; C""""'~H.ow. C . - - H....
~oIo.N.mibill,SnuthAfn<a(C.rel""',""'1

79
Crusub rupes trisThunb,
SmolLm .... hbranrhl',J.htubotuo;onnh,l'h;fbl....... , .... tui.5 ..... bag.of"'" Srcm.l 4cm~I"'nca'.k>o,..... 1<mIonIl-Ionn,"Il,."....,t.... whO\t""" ...
""lhrWJn.h""''P..... ptn ...... ~ Soulh A/n<iII (Ctl""'""",.:t"
SoulbAinnIC.p"''n,,·,nao)

86
(fjssul~lomcnlosaThu nb Cy"~nchum ~ phyllum (Thunb.) Schilr.
~cm h'~h. J""",ly I"",)" !,;"'y-g"-" n
1'1101 1,-","-' I ~<m ~"'g.amo"!,;ed '" F""'ly brooching "~m.' '''''''"gat nod,,,; "",1I1~~, .. ~ mm Iongaoo I mrn
""'~",wh'''''''I'',k>}" 'II,, ..., fIo'H'TS "'Jokg":,.nflo,,·~,,,,;thbrv"'n>tr;pco

~,":::'gMN<M" lb'V M.daSO""'" M"",mbique, S"uth Afn<a (Na t~t)

87
Cynan(hum
mimierii numRauh

Cynolnchumsp.

Anunid,..,tll ..-d "J'<"'mt"n ",'h round"" ,~u.k... 10- I~<m In d",,,,,,,,,... t>J
iu<'I!-'hlnt-<~",""".~m"·n.~..... ",,,,.....

88
C)'phostemm~ C l,lrrori
(HookJJDcsc.
~ITAn,.,

F"""'vbw.dli~_""""

=t.'.
4mh'gI> ...~.h""'''''''·,J-I'''''
b.o&;lnp.>n::~.;·:,:,:",'

n'rhol~c.Phtd
Angob,'l.mlt-l ••

C)'p"ostemm~l,Il fT
(bell & Mcnd.)Dcsc.

SwoIk",.",nl. ",.h P""''''6


b.o'k.b... nchi"li~l«llr
fton'l'pI' •• ndtorminsmuch
~~':~k~::t~ ~:,.tn.
lI'O'Ierwllhundul"em.>'lIin•.
gr""nL'h.}'~lI"wfl"w,'""
"n~"'"

~~l~!,:::,~~~~:~ lookJ.) L Bol.


l<>w.frHlyt... nrh;na .... l\lb;I..,"'K5<mlDng;p"'p... no..~
5ou.hAtric.(Or.n~r.... 5t.o"1

89
~:I~,~:::~~:~~:[nt~cfum L Bol
~1<.~:k.!,::~'&h<"-"""~~""'_7cm~~ndlG.-1S_1lucl.

N.om'b!a/SouthAfryaUittlr"'~mMJu.o"'ndJ

~~~~:Y':~~~~~~~:'E L BoI ?~~,t ;,r,~~~i~ longipet~l~ (8.lker) Mel


~t..~=t~b"'~""'''''1-2<'111'''''sand'"''''lt'''''' i~...mon. 5kms to 1 '" IonjI. ,..--tlC-I.,,.,.. ,"8 .... ,..", ~ cmlunA; )"'I~"" nu...... Wltl1
~"r'<'~I"f"*<
SoulhAMa(Ooa1'Ql<'r...... Sc. ....)

90
Dinteranthus inexpectatus Dintl'r Dinteranthuswilmotianus L.Bol.
.tl5r"S.,,,"ITtll,!,,tfAI ...;S~M".VANTHU.'ACEAH
S/('m~, ~"'\'.~"""' "",n.l"h .... ,.'" 10 1,.." long _",ol J

;;'':i;::h tf~",t""""1 Sf,",," ,J"t" ~"tI.'" n",,'~'"


<Tn ".,ol~ borne in ~,,';:~tf:'h!t;~~~~1h~~cnn~~~:!"'ilholarkolots, 5-6= long an<!
SoulhAfn oa(C.pcPI'<W;""")

91
~=~:::l:~,:h~ntipes (L'Her.) Engl. DolichosseineriHarms
Aflo"·l"rmg<tl"m. GiJn""btcrr.m'·JncJ"dc..,.ith34acn.il."'ms'o,n'in)(l>u,h,roonJ.-.l
f.,,',
1• .,·"';VKlleffln"·."'oornt'tn>ml....

92
Dol'5ten i~f()(' tida ("u~ k ')5ch\Veinf Dorsteniasp.

Senn.." 1;,,,. I> ; ~", J""~"'I""~ """,U 1>."",1 ~~ud,·, 2 3cm in d ... "",,,,,;
::::-";:l<m I"",~ "-,,h ~,"".lh· m",);,n,; ~' .. ni"h.y.. lk>w. <1.« ....... )"""1 in·

93
~::Ec::~~h}!~~ ~:~~~~:~~ (L Bo1.) Walgatc Dudleyabrilionii D.A./ohans.
S,.,,,,,h,,,,4-6omlongkanng4-8smuocll,grey-s,,,,,,,I.,.vel<l-5<ml,,,,S; UnbJ".""hrd.<l"""""i'h'~'",of41.1-1201~.,,"... ;wh'Mh'J:'f"\... I.,,,-
pink 110"''''''' M..,,",o2fJern Iongand 8cm ... kle; ""1~y,,I1"'" fl"""",,,"
I"UaJinl~",pn.. '''.''L.BoI \.1e"<iro (Ba,..C.hfomia)
N.mibl.,Sou,hAfrlca(Capel'To";,,c.,1

94
~~~k:~,:;::!foli a !l.1ker
Sio.1n<o/-cw5-.nsk'Ol •• -6<m Io n~,rod 1-1.5= ,n d;am"' .. "·,,h I~to.m.ol,'~ 51""" ... "'""~'''''; I".v.. ,{)-20"", InnS "'''h ""'r~'M'.l"n""2 mon InnS;
In'''''
""''''''''·bro.... n 10 bnlwn flo",.·", wlth l·ell""~~rown ,,'horl
~;.~~j~ n""'~,,,.
~l hAlnt.

95
~~:::~~~~,,~~,if~A~>~~J Schwallt~ Echev('ri~camicolorE.Morrell

"hrub ',,60em h'~h "'Ih br.nd,,-.l 'oom" );'w-~"...-n k ...."" 1-2 em long
,,·,'hd.r.. erd"'-,;r'n.IIo, ..·.......
r...,rnibi". <;"ulh Ain.:. (C.p<'P",,;"""1

%
~~:ff::.~;binor~ v~r.c~run,ul~td 11011 ~!~~:~~~::~cotri(h JA ['tlTPUS
~bI<Ilo I' on h;,.h; "*"",,,.nJ "'.'~.. nn...-ml WI.h ..-II1k'II.....; 1,00'-"1<'1
!e;~~~::.:2~:ni::':~;;: ~:~~ ;.~';...~u~~.=~ 10nnlonj;ond2Snn ... i.k-.redflo,.·"",
=:.."'' ' ' ' .. w''., \k-um(Pu<bIo)

97
Ethvfrilll ' PerlfvonNllmberg'

~' .. "'IS~ind"_""hpon~tin8'..JIe.o''8.''hybridui£.gil>l#fbo LoI'li:~clu ... p>ui"""""" ....... nn",d .. m<"I""!t.... k,l>I,,,,,,,,,,,,,"'''hlUftooi


'oaf ....,.u.... ~I .. ,.,,"'_. h.>;fUlbp
Gonl~"'<>ngln

98
~:,~:.S:[brigida (Robinson &: Seaton) Rose
..."..-.....
~...........w.soI, ""~""'hi'~ IN_ ... 25 ..... m,.~nd 10 ....

:-S..,., ."""1n.
_"'IIh ..... "'..lI',,"'in~lulml.ll~"-.,jR.tw..~

99
~~~P~:K!~:~~ns R.A Dyer &. I. Ven!
I.JnJe.bnonclll'd.""'"P'''Sandmonng.S.tollknglood_w;thltun,.
C!:!";'~::~~~~~~~~"IT';-
KcnYll.. T~1IUlIl.>

100
Eriospermumdregei
EndL

Tuber5-lOcmwidowi.h
,""""";bo.. lh>av ... I'I?d",!'d,
1 ormo",wdlJc'Vdo!,,-'\l,
h.airy.ndw;lhp.,., lId,·c';n"
I.-.l""""nA""""","ce;wh;.e10

:~~!":;;,:~

Euphorb i~ abdelkuri
B.11f.
EIJPHQU'ALrIlE
C .. y.,em< t<>2 m high ""h
5-tull-angk-dbrand..e.and
pron';"""'lulx-n:ks;rud;·
m..... '''y.cadurou.Ie.''·... ,
ydlo"·,I"'",,,,,ou>ia,,,,,,

~!{~~::~~~=:
SOll 'hY'""' ..... (SoroI,..1

CiTESArr· 1I

EuphorbiJ ~crurensis
'J.E.Br

(1l1'5App.1I

101
;~r.~~:~~~i(OmiS S\'ent
M~in<l<'m '-~m "'II. --.:Lory b.-~nchesd.ick. I"""'y ....... ) ~oW<.J, rudi
monIaJy.".od ........ k.,"',~_<m""""l'f'dun"'.'"""
."...
.......m
M.><bSOUC"'- crn";'Appli

102
Eu phorbia brunellii Chiov.

~~ES=~:~~~~m1Jo~3d:i~'~;~=~:~~~
So"",lia CITES"'pp.lI

104
hpllorbi~ '~p-s.linlemariens i5Itluh
~ur.:~:~!~.'!~vigeT. N.E. Br

~i~i~:~::'~::£:~!~F.~~~?~~5~1~ii~';l~·~IUW
Tul>rn>u.ruuI/-.lLo""'in'tomt"/onnt~idr.udn;_10-15<T11,,,"g
~.:=:.,;:;:t::~~~.d~~g In P""'; ru<l, ......... t~ry k.... /""~"'"; in~
,\l,o,I.o~.".r c:tT'fS AI'P II s..·.nil~nd ClTESl\pp II

105
, .. . . . . . . . . EuphorbidcC>ero lescens Euphorbia crt' mersii
;;1 Haw Rauh&Razaf.
Tuberou.root 2-5"",;"d ...
""-~,·,;'h'n.M,,.1(1-l5,,"
luns ", ilh /",,' C.J Ul"""W,,,
"'"p<-'.."-"'c>8cm~"'"

n:n::~:'~.!:O~:'
.,..>Junc'"2·-3cmlong
\I" d"SO',...,,,

CrTES,\pp,1I

Euphorbia
cryptospinos..l
~::I~;~~~~~Af
Tuberous rooI ,,,,heft<1_
2\l-H)I,mlong",,,h 5-IJ""
.pi....",/o'minglong,(ocI""
""p,-,,,;rudi,,,,-,,,I..>ry.
<adumu.I .... ,..,.~mm~
,V"')""p"k"..J,nfk>rNu<l
lthiopi',K."y.,So .... b.o

ClJ'f5App.t1

Euphorbiacyli ndriu
AC. White. R.A.Dyrr.
B.510,1"(>
St ......... h"''Y n,'''''hftow
<}'IH\dn'.lbr'I\("'.. 5<l11~
Ji.m<"I<-f,row",..J"'ith""",,

f,~.~~~T~~t,~
~~:~~~=
niJ,>/tub(r< .....
So,,,h/lfri<,W'p"I'mnnl

CITES Al'p. II

Thlc""ubcroo.src) b",,,,,,,,,~ lOcn, k",S onJ l-"em ;nd;""""~,,;",,,"y


br~N"hos 0;...10 ern I",,~ ""h "I'i .. lly ~rrM"od. g"....,i';)o·Sn.oy I<M"" "'Ih
.....Jdi,hm • .-gm"rt.'dmflc::>re<....",."bome""b,.nrhed pedunc"' .... IOcm long.
I''''nl photogr~p"'-") in 1Mb",.
::;::~"."""'!Jn'tH'I'l"I"f.'Uh"ml;rl"'UtSt... ,,'
CITESi\pp II
106
~l"!!~r.~!~~~i n drifolia MJrnA..lp. & RJuh
PIontw"h ~'h,,;,h >Ubc"ff""-'~ft b, ....dK's . .,.'ms 10-15= IonR " " _ ,,;Ih
~;:~~~k~~~~·~;-~I::'n~~':I1,~=m:~~~:n.
""'" ~ilh. ' hl<k """ kmnmgA cAud ,·,
~"or Cm:SApp.l

EuphorbiaechinusllookJ&Coss.
:~~~~!~,dAeEraryi Guill.
\O>ionJ' moo'SI ...... 1012cmh;~h"' ,''''',M ck."; .. I'''""'Y·S",.,,W<ov''''3-Icm
Iorf; ... 2rm wiJe "'ilh miJul.k m...~ms bvnw.1 ~n,js.,( .I<'rru,; y~llo .... l>h·
p«1'inf\c,,,",,,,,,,,,,,,,,, p,..du""k·1u2<", k,nl\
;,,~~=t~=';:-~~~l~~~~~~"';:;~:;~:&;:"f'~':"k
1Ii<'P""" CITESApp.l V'~W CITES App.11

107
;~t.~::~!~ ::~x \tarloth
;';'::l.<!~~~~~::!~:::'~',::;:"";;::~f~-;:~oh~ iIoo
::;'~1;~:~";'b:"":~'~~""""'·""'·Mm.·""r"" ..... , ""'-':-
/pplo.><t .. , .... ,I'<', .... "'fUt
So.!ulh M ..... (C.po: I'ro, " ..... \ ,'nt',Al'rll
108
,E,"~~~~~:: ,f~~ u i IJ A.C. Wh ill'. lJyt'f & 1:1 Sloo(\('
, - . ..... ,,~ ...... u~" ' ~~ ,nlu"''m IO j , .. m <>uJ, ... lI-lbnn ,tu.~. brA"''''-''
""".mk"'l;.,uJ,,,,,,,,,.,,,,.,.oJ,,,~,,,,k·.""';<w'h .. ..t ...........'\l"'enol"'(
"""'fIo..... <J<,... I".... ,,"'-k--.
'wih·\Ir ... ,C'I"I'r.n;",," ClT'bApp.11

109
;lU!~~~~!~ f~~bos~ (1 Iaw.)Sims Euphorbi~ &r~ndidensHaw.

~Un~:::::d~:::':d=i!.~:''";''2~::t7<i~..!~~~
\laLn 'nrnk/>-IO m hrgh. br.nchrng I_I~ kI form ............,"'= hk ~"'..
b•• nc""'cndin~;n.k.-rulo",· r m r ""11>r.,,,hk.1,,ruJr""'nL1,,,.,",.I""L""
pe~'e" ""'u< h·bra""hPdf"'<1'm,'~l(Irml",,g I...."'" in/ln........."phrun""o/>Or1 r«lu,,,,lcl"''''''''.hull~.'',,,,,,~
r~,%J_gl""""'t. M .rIoth ~, 'hA l rI<~rrra"",· ... 1) {fTI"'~rr.
$ou,h MriGO (Cape Pro""",,) CITES"I'!'-II

110
~t~:~!~f:: llem elii L.fS(h &: Leandri
~."""b;imiJ.,,!oL' mihibu! w;lh,m.U ... I""ves.
J.IoU.POC" CITESApp.1J

111
~~!t,~~~~!~ Ih,~pt~8on~ L.
~~I;:'~~;"~:;~~~"~~~,~J~:b:;'~'~'~~-::~'k~~::; ~~.~':~
brdn<h~?I('"

,.",..... ... "''''''',,'' 11o.'r'!\'-'T


Swlh M,l<. IC.I'" '''''',,,,d Un:Sllpp./I

~I~
bt.nchodp<oJu""ka-'O,m~
1.Lod.o~ ... ,

11 2

Euphorbi~horombt>nsis UI5Ch&lc.lndri

=~::,=::."::::..~~~,!-~~~=

(1'1'5111'\'''
~lU~~~~!:lh,~rwoodii S. Ca rter & lal'ranus

~~I~=i:..OU':~e';~I~:,,:~~~!'C:"':,!~~",":'~=~t..·
;:'::::-.Jpo...wu",k',.......,. {m'~:rr
~,"~,~~f~~!~ ,i~~fll1is ~dr. huttonae A.C. While & Dyer

::~::k'ir::::~~""'" "" . .""" rn."nly,h","'~u,,",,'1>y IML"r I>rMl'~


~Ir""." • .",",
s....'"Mr~" (C 'I~·,''''""c<'l CITESA pp. 1I

113
Euphorbia l~ikipic"sis S. Carter
~LP"OMaIA~1 A.

51~I""br""d,lnllf"'nlb.",>;<vl",Jrk.,t''''''t...,j brd""t....,.5 lU,n •• "'.. . . .


""h.,~.l'ln "' t"l <mlo ...... "td'monIA"·,r.,d,"">U,"'A""":wll",,· ,"fl.",...... ,
borne""",",", p<d~""",
K<'1I1" CTT'E.......I"I'.II

Dw~rf>hrub ,.,20,,,, hi)Ih w"h numerou> b.-~~5 mm "'id.... ui>."'"


~rr~n~,npo,,,,,nocI''''''''' ... .,..<~''urouolN'· ... : 'n~Wl'''''''''''.
:~':::.borrw""ohI>f1pn1u""",.,b<;>""h'p;c.","
UH'S"'ppll

114
Euphorbidledienii A.ll<,rl?,-r

Slwb'o?mh,#,,,.,,,1 '-.'");I"""-"".",, "~f"'''''''''':'l'ln,'''''''2cmlons


"1""~rud ' n,..,, ... n_,_,Jtk"U"""''''' ....,." .... ,·,-11,,,, IIlfl"""",-tK,-oo"",
...,b"'.... h,I''''.,.
SouihM",". '~' f'·r~,,,·,,,.d CIrES App.It

~u!~,~~~:~ :~~ho8ona 1..1111.


r~:~' :::,~';~~~~'.~'~;r~;~'~~~,:~~':.'h~;;~rk,"~~~;::V'"
f"Iu",~-2-1.-m lon~.

\ WJiO-.,...." CrTESApp. 11

liS
Euphorbiamilii DesMouL Euphorbiamillolii Ursch&Lc~ndri

~~E;!::~e~:2;~;5~Em7~=;:~~~~h
Shrubl"20<",h'gh.bro"d ...,)fN",b;i",,;g"-"I,...Jl •• """I04('JJ\ ~ mg"
gm";"g.pic<-;:...dd;sh;nflo"""""'C('w; ' hp<...J""'<y~lhi.,l>.:>moonl",j .
"JI(klolcmlun~
/;uphor/nlls",mdrn.Bojor M.d.~.r Crn'5App_ ll
Modogas<ar Cl1l:SJ\pp. 1I

116
~:!,~~~~:~,n~~irii N.E. lIr, ~LU!:~:~!~:A~humbertii SQitcau
~"",_,t,.,,.to 20cm km);,lnd t 'i mm Ihlc~ ,"i>m~ from b,t..,..,r cauda,
'oJ
I,,\,,~
....., 2 <rn I"ng "lid I mm ,,'id~; InA"",,,,,,,,,,. npo< hps 0/ bran< ...... ~:::~::::,!:t:;~~odk!':~:t~:~o;:i;~T.:=:~a't'
~~.:~~~!~ tuns
""",.pi<Yo
ClTI!S App. It M.d~S""<.r CITES App. II

117
Shnob'110 120C'JI\l<lj:tI.<I...,.t>r.onct....J.to.-~".-to.-.~- ... I' .... !,,' .... ,h .......
~~~'7n:'.:;,":.r.:~ru.h"" ...Y".,...,......,..... ,,-..;.-I ....
M......-ro Ull~"rrn

Euphorbii obeu HoolJ Euphorbiiopuntioidrs1\E.8r

~~'i!:7:~:=~,:r.7"'~,~W~I~·i~:::="=:'
.honl",jy ....... ,"""<I""·r·«··>femo... ~nd ....... n.,...-.......... l>.'""'."'d'ff.....
c.... pL'n ....
SouthAfn.-~((".pt'f'n"',""" CITES "I'!'- II

118
119
~~,~,~~:~~~t~ idl'rnli S M.G. Gilbert

~~~,~uK~:n~.~li~".,~ ;~I~uC::l>;~~:~~7";'~~~~~~~~n~'.J In
.plT.l"ndm'·.......Jw;'~""..... I;~~I\''''''I~'';nr~tth<>rnt>'''''lmrt
l"-.Jund",;cy.'h"'Kn."•."""uI>IJ~.l"'II"w in."d~
Elh,UP", Som.>hll CITFSApr II

120
Euphorbia p$eudO(~(tu5
A.Berger

121
EuphorbiasquarrosaH"w.

rhl<k . ..,ht.,...."... "coud~."'1h",,"'<'I"OU.J>fO'Il.>t.O •• ""'SlN""~I.;;rm

~:,g,::';'~rf.,~~~~~..~:~,::~"<:"..!~~U:"I;:',,~~~ ';:~::;::::~~~~:',..'
E"!""',b""",,.,millol>llLom.
<;ou.h AI';c. ((Ap"''r~,·,n< ~l (ri I:';; Apt>. II
123
Euphorbi.l51I'nocl.ld.lB.J.ill.

Shrub to I m lugh "· 'Ih ~t. ....n...""br.""ha ",lh broo<"n o.p;~ ydlo¥. in

\,.od.ol""""'
br.""".pias.
1k>n:ot.""""burnc>I .......
ClTESAppll

124
~!~~:~!~:~:;r~mJ R.A. Dyer
lIody:llcrnh'gh a nd1 5nnthl<kbunf'd,n""I"" tht.pmo!;upto ;o br.n_
~30crnkll\garj..., f" ",',-"udc";b"'n<h.o>wjlhtub,....:k>bo.""jnsp.< jrs'"
~.,.,Tu:~l~t:~~%I~;:;;;.'dim<nt'ry. <,dunn., 10>.,.,..;in!""""'c"", ""
Soo lhAfri<orr'.ns'.)~!1 U1'6.\pp.1I

125
;~f.~~~~!~~~fueri IA>nlS
Sh",b." 15rnk'~h,fo.."clo.~
>1<·n»"·,tl,',,,,,,);uL>,".'
("U'hion'''''"nng'r'''·;~''''''

t~~f:!~~~~:~
Ion); I"d"nd".ri<ing 'n""
"em.I'''',....
\'Ad,.g.w ...

Euphorbia "iros~Willd

Lu'g<· dunlf'S lo2mk,.h


~;.::::h~;~'~f;.rl.=
S"'Y>f',n.,,, l<m ..... ~. n.o.I'
~nfl~~~~=~~';;..
pt."d unde.'b'.... h·f"'.,.
Angol •• '1.m ...". S.~"hA...,.

CITES App. II

~~f.,~~r"~!~f:~ldii LeitCh
~r!i~t~~::~'r~·;,;~ ~=
~~~n~ :~,~:;~.t?2cm
l<>nl:;,." r.f"'''~ ...''iofl,; ....
""nc~bt" ... on lOan Ioo~
pcd uncl".ns.in~"""'-

EE~~f.~'PW.

ClTESAf'l'.II

Euphorbia valida r-..E. Ik

s..h'.',".);Iubo>o.·Iu.~",,~ •• cdl¥',,·n"cm;'o13cmindiMn ........ ndk,1()<m

~;~;~m.~:~~~::IIIo;<br~;h.:r.~';.'7;~~:~~~~a~I,';.;~'r~~;: ~~n~.
Sou,hAIri<A(Al"'"l'n".",,.l"" C1TESApp.11

126
~:::.:'~~~: 1io,,:.~,:-v"'~-4 ern k,ng "'lth wh,t"h ""'T);m•• ~d ~ mMl;"..1

':iouthAirialK'p"I'",,"nC<"

F~ucariaSf'1"ciosa LBol
"">U'"~VA.""~""<..lAl ""'L"."''''''''''''AUAt
~~r~~~~~::::~,2.5 tm kmg w,th c... ,lag"'''u> bord,', Md :l-4o, ,,",'" ~= !~\~<:t2~:,~~,,~~~p.~ ~~,~I:':t;:, \;::,t'~o~~'h~~~.::,g\n'; "'~.., !<'<1h
South Amc. ('-"f""'fi",nC<', So"thAfric.iC.p.PruI'\n«'

127
Feneslrni•• ur.nli.CJ N.E.Br. Fockuedulis{Thunb.}K Schum.
"." ..... ~"'H'''~<'A.
a" .....,I,~I"
..
,""'" IN ...... 2-3 an k>ng w,,11 t.-.n>I,o<,.
~="'d_.J_t'Iow
~1".1..,",,"'~ d .. "'f" <'·~_.·"'lU<m"'d
~7;tbl""l\ ~.~
... m<i ... ;<I,...t>.nl:ortr>ih~I:_I<O,.",......t _
..... tun.... fk-..... l' ...... ~1I"n lIu!:"d",..."""".,"
...~

128
129
FOUQ U I Il RI "

Frere~ ind inD.Jb;clJ G~steri.bifonnisroelJn.

~n'n!.":=~':..~:;'2(D\lh..... ~,w,,_6<m ~~~~,~.~~~",,:,7;'.:t~.:':'..-.do


h,d.. (111'5"1'r1l
SoulhAfri<-~t<:~I"' , _ ; ......J
130
~,~~~Af fl.lSoCOpl.lndAIA B.lkcr Ga~teria verrucosa (Mill.) C.-I. [)u\·al
UUACF.A(
RnIoeIt801l0IV'f)'1V''f"'\.d..nc .......... ,..,.15cm!oo&w2 .... ..-.:to ....1h
fE:.::,,~;;;:=~!i:,~~~~~w!.m"'~;~d nwnrnJU!o""' ... lub••d ••• w' .. ber<-uIo~ .... IJ!l"'"'.-..1no.."ftSV.n
~cId~'hiolOb... '·.riotyofG . .."_C.·lD.".,,l
South AfnGil (Cop" Pru¥i"",)

131
~:f~D~~~N~::r::~~~~. Sr ::r:a"e.~~N~:~OASc~~r(Haw.) N.E. Sr.
Clump"r()1"m;ngp."",oIu~".'I".od"",...,...,,,,"'l<'<Ibyd<'<'Pf..... ,..,;g...,.- Woody root>tock.lorm.ingdumps 6--15 cm in ..r;"m..~er; uooqudl'''c-d.!V''-""
8"""'I ... y .... rov."..,.jwithfiner... ..,"...-ddi;h-violelflow""'" le.,"'2 .... cm.,ng:"'lepmkllowen.
So",hAfri<:.IC'pol'Tovi"",,) C",""" .. ,..,,';riJJ,(tI.w.jN~.Dr
South Alricd (Copc Pn,yil"oC<")

132
~~[~~~:~:}~~I~~~~~~u m (Kensit) L. BoJ ~:E~J~eK~~'<~~~:~~_~~ var. sh~ndi i (N.E. Sr.) Glen
Pl.>nl< forming dump" 20-111 <m in. ,H.lI"'"~r: s"",m,h or S"'Y'g,""n k~,''''' v'r>dywltlll'lalry.y~lk>w·K""norS"'y",hl,... "cs.
6-10cm Ions and ImMd.1 b"",,; P"'~ I" pw~,,b·,,,j n... "'e.,., Gibbon.moluJ..Jii(N£ 1I...)S<IIw.nl""
s"",hAfri<.(DF"'l'm"i"",) SouIIlMric.ICAp"I',,,,·i,,,,,r

133
~:~:~:~Rh;':l~~~r~:~~~~!um N.£. Hr.
!,,!:~';:\::;\;}~":.~~n 1"".... 01 unrqu.1 .".... '''Kcm k"'~ . . i'h t",,,,,I~'
SoothAf";~'(C'I"'I'm";n<Y)

134
C1Qltiphyllum Qligflwpum L. Sol. Cloltiphyllum surrectum {HawJ L. Bol.
M[ 5f". ~ YANn'"MAnA~ M[".,,"w"'I.."H'''AClA~

~~~.14~~~~':d :~~7J~I;:::'~~;.~:;;:.;~:~':"'.Oft.n r<'\l.,;ng...J Ere"". ><m;-cy!;nJrical. t;"-"" Ie.,-"", wi,h tn.a"llular lips; ydlow
Sou, h Afri<I> Il~I"'I'M\"n"')
now~rs.

s.::..,nAlrico (Col'" """,;n<><l

135
~.r:!~~,:~~~m m.cdo\l~Jlji Ale...ander Grtenovi.diplocyd.lVebb
ufi><.. IJ~
=~':.r~~~=:!,,~~~~~"'d-
s.mil.J •• "CMU""bu......
(:~"""Y l.und.
\k>. .....

136
Haworthia a r~(h n oidN (L.) c.-J. Duva l Haworthia attenuata variegMcd form

Sk-m""" ...-,... Io~cm in d","""er "~Ih ••" .."all.",,,,, 15mm long ",In Arult,vatt"d form w,,",·.n.g..t<.'d ..... ,-....
wh'l~ m'''I\"",I,,", .. ; i"fkorc><," "'~ 30 <m ,.n, white flow" .. wilh S""" ,·d, .. Soo",hAIricAIC'p"I'Tovi...,..)
HmJ}oorrhUlgiS.. I''''"nn.;H . ....'''mH .....
SottihAfric.,C'p"'I'"winn>'

137
H~worthiachloracanthal law"
~~!~~E~~ia cooperi fuker
~~~,,';::~c~ta~ ~~~:~~~1~ ~:~~w,~::" ...,,-.s6-8cm long; c .... .,.!"""
ro6ctt ... ~ em in di~rn....er. p.>"'w",n 1",,..,,,,3 em Icn~ with ~mm
~~'"ci~,,"l b.,,, ...; inflor~"",,,,,~oIIl em ,,11; pmk"'h"w~".l1o"....... ,,"i'h
Soulh Afnc. (C.""I'r<,,",....,.,)
Suu!hAlnc&(C.""Pro,"m<e1

138
H aworthi~ limifoli~ Marloth ~,a,::.~~ i~ mirabilissubsp. mU[ldul~ (Smith) M.l:!. Sayer

~';.::~~'~~~:!:fiS~:~~7~$"HC~~~it~r.~·
EE~s§~g£~~;E§i:;it~~~E~~;~~
"""'I';nl7<"IJby\\'Ull.>y<'t"(19<12)
Sou ln Africa (Cal"'i'n""",,)

South Af,jC<l (T""",'"I)


140
~~d~::o;:.'=~;;:!~'~hd:=~~'~~~~:"I~
~,:.t1OWft11W1thll ..'.'n'''''''''""*,,bI,.t''''!.'!I'''''''I'<'<io'''t'''nthor

, _ ...""ml...... tLtl .....


SoolhAh"·.Ic..I"'P",,",,~)

:!,~,7:~~~i~ p~ rksiana roclln ~:,7:E~~i~ rddul . (JaC"q.) H~w.


""""" .. 2J.mind.. '"'-"'-"" n um"ru....ur~!lJ'<""."""""ng)".o.' ... 1-2<rn C.....",_.~ ........ I<.,,,JI'<"l'III<a"""'8<m"""and2<mw .... 4'1».o<
~wilhthinlUb.r< ...... ~25<mt.ill; ... bil~n.:"."... ... ~hgn."'" ~.. h''''.''o'''''nflonooononno:tOnn .. IJ:''"hM"'-"'' i'h...,jd....._ n
..
_I Afri<~ (C~"" ,_,,,,,,,) Sooth MI1<1I (Co"" '·n" ......)
141
Haworthiatruncatd Schonland
.:n-rt.'rulK".'~.&"",nlea\""'2-J(mlong ..;'htrA"-.I""""',,,;ndow;lnllo.....
om<.25<m,.Il;whlt~IIu ...'~rs"'ilhSn.",n "<in>
Sou lh AfricA (Co""l'n,,,,ilK"d

Haworthi~ starkian~ Pocl1n ~~J:~£~~ i. turgid~ l In",

~~'~d'~ r:..:.~ ~~;",;:;t~';f=",~~j:;~~~c~~;~~t;~~~~,~


5remte.""""t'""oIScm,ndi.m"'",",uptnl0g_nl• .,·.. 7cmlongorul
2cmwide;inno""""', ...."l5<m ... II ... hllen,,'"',crswilhb,QWnishwill,
S.,,,th Ain<a (c..I"'I"n.,,·mre) Sou'hAin<a'ClIl"'i'ro.-..>Cl")

142
Hereroacarinans(l law .)L.BoI Her('roamu; rii LBol.
Ml~~M~~V"NTH[MACEAt

~n~~: ;:~I~~~::~Iws.nd ,(en.,; u"'"-'lU,! >i",-.J.g",y-g"-,,n ICd'·"" ~~;;''::..~~.:;: ~'gh; br., ......... ","h 4-Il"""'-<yhn,lnr.I I......... 4-7cm Ion",
SoulhAfrk,{C.I"'[>ru,;,wd S..ulh Afn... (C.p"I>mvi......,j

143
Hood idwrrori (Hook.)Dt-cn~. Hoyaimperial is Undl.

~('mSwithl5-25.l"gk.... ~cmhl~hdoo3-4<nli"d\o\m"L"d\\dbraru;b
;nsIrornN'""t"IDrmI...&(·clump!l;l>"~.,,orpinkfiow .... 11dn'rhoto-
~rd l'bod
in ,,",!>lLIt.
AnsoW.N.m1bi.>
H u('mi~.Isper.l NE.Hr

~;:;::::=~.!; 1tI~1o.>d!ol""'" 111- IScmru"h_lcdh. '"",,101111-"-.1 s...m~'I_Sm\o"'l!..,..J"-''''mllu<k.:ud'''''''',;rwnlN'.... 2'''m'''''K.. ~.


b..,..-n;,.b·!:""" .... cllo.;..o>hn.,...,..... ",th""......:.."•.-purp... l' ......
\!.oIo"'I.Mu<.. mbl<l""

145
~:~~,i:[)~:~~.. rp .. "AT, S( hweinfurth ii A,C. Whit" & B, 510.1n"

~~~~I\n;:~,'O~~&11'd " •.",.7_12 <m hIgh wl.h ''''''h.o 1 ,m lung; m.I


SouJi"'.bi.o,Y"""""

~:~:~J:::~~I. E.A. Druce


~~~"'u.~'!t:~~~~~!~~~"....po~~~~~,
c;."",hAfrlCillc..p<r"""""l

'46
ipomocaboiusiana
5chim:

C.ude'w20cm;n,t"mM~t;
Lhin,ll'ffi>w,tk"mpkor

~~":,~~;6.:'J'~
\-l.Id.g.O<".r

~~~~~~a,,~~~~~:i B.lker
~!~d:,~.~,~ ~~.::~r
hhfonn,gr>1'<1'lt ..,...:,t..."
pin~lopurpknow~r>.
BI" .........,N.mibi.

~
I
••I Ipomoea marmoula
Britten & Rendle
T"b...'rCIUSroot;<lem<
2O-JOcrn hiSh; "'rg<' •• il,·.,....
Sf..,n, p;>1""'h·I ••""",ml~'

6:~~~;::A

lbt-rvillu50norae (S.1V3b;on)GTt't'ne

1.a<;;•• ,,,,,,,,o'l..,g l olx""'<ilu J,· '''· i 'h >.-'' ~r'I<limbi''l\brand,,-.. oV<'r 3m


~;~:.od"rousg""""le.'·''':6,.,...n.\.•'II'',,"fiOW<''''.

14'
lpomoe3 Sp.

Tn.,go.'"u<I,."..om'nd"dc."
sp«;"';ofinlc",.ttoCdUdj.ci,
fOrm~nlh,,<ias!S•• nd ... \,..,..1
fi""~pt.'<;n""",h.\·~Mn
imporlcdfromilicwild
Id"""~CA'i"" .. d,i';""II,.nd

:'::~~d~~~~~I~w
~;,,~~~ ~~::SC.P"-"

Jalrophamu[tirida L.

148
JOVIBAR BA

Jov ibarba allionii (Jurd.&FourrJWebb


Ad __ upofl~eflowen;

149
JOV IBA.RB A

~~:~~~~~Ea::natia (e. Koch) Opiz


Globul.>rl'Q5CtlCS'<>2cn'indianl<.>wr; bri5 iol sn,-"·I,,,vc.. nushc'<lwllhn.'<I.'
al"";<>ff..... ""lton"""t~l.t.ms<l_t0p"rentl'Q5Ct"'.p""'Y"lIown"""'~
Austri.l. IC.rin'hia. soo,hcn. Tyrul. Styri.l.). Italy (Cam .. )

!~:~~~~~~:i:a (I..) Opiz J Ulladinteria deserticola (MariothlSchwantes


).i[5[.\tB~Y~n l l[M~CEA[

~';!~~.~~~~f=r~~;!,hf;~~~~;-:,~willc~~,~id;;~,~1:=1':,; =~:~;J~';.;'::t!~"'fl~.~ndi'h.g"'Y-g ....... k>mol,5-2rm!ong. w;,h


lIu,tr",/l'Jly (~Skl.l AIl's), Carpathian., HunS"ty. 8olk.mo (nor1hw",d

150
Kalan choe fedtschen koi Raym.-H.lmet & E. P. Perrier

~~hW':'::::t~b~:';'~:'il~~:1jO:,1~tb'~~?~':h~;'\\
Maddll"><ac

lSI
~"~~nu(~~:tkewensis Dyer :~~:s~~~~:~illotii i(aym.-Hamet & E.!', Perrier
Ora"dK'<l,h.liry shrubJOcrn high; l~d,· .. l-6crnlo"8.t>d 2-4"" wjd~
~~~1~='Ag~=~"ii'#==~,~~1u';~"~~I~:e~,'~<l. m'·......,j ""th haj""rmk<><yellowfl"w .....
Mo-..d·'Sd«M
C>rd",,<>rigin

152
KabnchlX' J(~piger~ WetII'. :.~:~:C,~~!~yrsirol i. liar\".
Shrublt>"O'mhlgh..I'fU'.-.-Ie.o''''' -l-·S<mloH>A-...-..ln.. ......... o.--If""fv<.l..... "'lmh,&h..I'fU,""""Ie.o,·""1~15'""'~.n.l ~-6un
,""""<"Io."/IO,,_·,,,""I!I "...... "·1thr«lol ......... 'lI"",...,!k>wn.....,,..
~II$<>lII.s.",lh'« .......... Is..mtr.1 5.Ju!h Alnc-.. (l-..I"" ,............... Tt'n,,-,ul

'53
Kalallchoezimbabw ensisRl'lldle
~~~~:~~:~~~,f:.~~:~mus (L. 1lo1.) L. Bo1.
~~~~~r.I~~~:,.s:~";;~~~~'7,!~;":n~~!~ lung and 9 mm Ihic~ "'ilk
Sout h Afn,,"IC,p"l'ro" , ~)

154
Lithop§auc~mpiaeL. Bo!
\if~E\iaMYM"'UfMM'F Af

~n:;;~!~h~~)d:;~~~~:.~!~;~,;~!:.~wn in colour
Suulh I\!rk". (COI"'Pr""iru;d

155
Lilhops aucampiae var. klrelemanii (de IXJer) D.T. Cole
.. ',f- ""MY ~,,"TlU., ... CF~F
A noSito K...."Y·bro"·n ,-.ri<:ly wilhdo,~linco
South Ai"". (Copof'roymc,,)

Lithops bromfieldii L.BoI.


M~'lM.W\~"ttn "~l~H

Budy 1.5-3 "n in di,om..~ ..~; ''''''''Iuol ._w."<.l, 11"-'1. to brown lruV1.'S with ~"'l'
~"'-'"m.t,l\i ... ;YOII"... flo"... rs.A ..·.nJblos?'<'...
Soulh Ainca (Ca",· 1'"",,,,,,,,)

Lithops aucampiae \'Jr. fluminalis D.T. Cole Lithops bromfieldii var.in sulari s(L. Bul.) B. Fl'arn
.. l>E."~M\·""'-If.lBMC'-M "f'L""VA",,"~"A(" _AE

A '."k~y ,,"·tth I"k~'''yn'''rgi'' .. This ..··ri<:lrdi(f(''''(roml''''tJ'P<'.?'<;."inh.>~;nl\m''''....,upl''''.urf~


s...ulhAfrica(C~poP""·lf1<") wtthLo,"&"dal.
U,h0p''''$I''.''''L.IJo'
s....HhAf,""(C.""I'""in<"(')

156
:~~~~~~~~~~~~~!:~fBo!
Body 2 3 <m in di."",-~er. ~""'I~~I ;ih>d, bIUi'1I""" lo pinki<h I~.,,,,, wilh
"l"''l''''''''''&''''';y>,llowflOl.·.... A ".ri.b~',pt'ci ...
SoulhAfrk. rCal"'l'rm'incd

157
~~~?!:~~~~~~(;v::~ l~ctinl'~ O.T. Cole Lithopsh~ lli i dO;' IJoer
".q"BRY"""'f"",'f .. r
~~uPl"" ... rf.ott.""'n\'b!~>p<>b"ndbloo.l''''.Jrull"''' Body I >;..2<m ... d ........... ckq>fi_,..,unoqu.ol...-..l.S""Y "'p'''booo<o...
.... ''''W"h''..... ~.W1''''-... and'''''dob;'''hlkllv .......
South Afn<.. (C ...... I'rm·' ...... )

.;s
lithopsha11ii 'Crl'{'n Soapstone'
",;:s;:."Bn"NTH~ .. "n.. E

ACUltlV.,wtlhg"",,,,h·ydlo,," lcd",,,,"

::~~~~~~hAa~~~~~VAh:!~'. ~~~~?~:AO:~~fr~:~::~icul~le'
A <ulli,"arwilhwh"ish"Il'"CYIc."""," Aculli,"or<,imi!.ulovarrloo.,..;

159
~~~~~~~ ~Ao:.~~r~ ::;:,~~rgil1~b (Nd) D.T. CoIl' ~~~~~~~!~~~"l~~z~~ephdld'
IOO<~'" ,·.ri<"l,· ,,·il~,j.,~ !\,"'fl 0''''-.;1 "'((""flS
A pjn ~ A 1""1~.mil~y.!\,,","rul"'·."""hd.'k .. m.'rk'"&'
l"hup. """X"""
Nd
SaI"h Afnc. (C.p" l'n, ..nw)

160
~~~!~~:f~~::f;!~lIeri (.'J.E. Br) Fearn
5mo11rr_thrryl"'''r-wllhron,·''''ul'r'.,. ... ~·
~/0<1"""~£o.
<;"uthAI""" IC.I'"rn..., ......1

.
~~~~~!: a~~:,~::,I:~: 5ubsp. bella (!\.E. Br.) Of. Colo!
l!.J,Jy2-.l<n\lnd;.,,"'1c"(,d"''''1'I'-.... t.~-Il''...... '''',''''''lth''''''~'"'uPi''"'"
... ,I.,C<'.nd ol, .. "1\n' .... dol ..
/j,",,,,/orIJ."I.E.B,
"'~m;ht.a

lilhop:s lu.r.ismonl.iu (Thnler &. Schw,lnl('<;) N.F Br. ~~I"~:':,k,,.i~;,~:~~:~: , ubsp.l'bl'rl.inzii 'WilputunsiJ'
..,'.·.... ~""Ttl," ..(nr
em.-..Ofbn:m1\"""tuo<hR'bo.Jyl"-..J'""''''du .......... wuh"'~ ..1"1''"'"
:.::;:.... hllfon.-.... ...... " ... n.1bi.......,-.
~~~l~:.~::~;,..~:.,li:,:;:~.rtac."'th""""mIou...nd

16l
~~~:'::"~~I~~~:~:~: var. aiaisensis (de Boer) D,T. Colc
~;'!~:.1;:':=~;::'k~~;;t~h\:;;~~

Lithopsk.arasmontana vu.lcrichca ndCDinlC'f&Schwantcs) ~~~:~~~l!~i~!~~:s Albinigold'


D,T.Colc
MESE ... B.Y""',IE",,CEAE Similar to 'Albini",' but ,,·uh ydlow I'k>"',.....

::~~~n:~~h':~;;;~~rf:'~:d:r;:.:t~~~~~,,""
~=bi.olnidlto"'iDint.".&. Schwantes) Dint.".&. SCh . . an'' '
162
Lilhops mannorala'Framesii'
M~"M~n"'rII[""~LU

~J;;R:~~~~~i.'j':;,'Wik.~i.:;..eq;~."t, pol<: gr~} I~a'.", wIth d""etgrey moT.


5outkAtn.:a(Cop<l'rml""")

lilhops lesliei var. venteri (Nel) de Boer & Boom l ithopsmeyeri L.BoI.
"1:S~"~Kr"N!f<"M"n", "1i>.",""r,,~rf<l" " UH
A pologrcy '·a';,~y ,,·ul> fine n""ki,,1f' ~t!;.-~iC:!';~~~~I~~=;d"wg',"I.pal~greyka'·"'hnSed wllh
LIlIo.'f"t<'Illmtl.<"i
Sou'hAiri("'IC~pcr.m·;"",) 5ou,hAfriollCap<l'rv,inu:)

163
lithop5 pseUdotnmCiteliol subsp. dendritiu (N<'IJ D.T. Cole
.. 1:l>< .... I .. NTt<Ut .. C( ...
nu:. ...b5p.'<......... """""'P"c""",..,..koi'fi".,.d.rl 1Iom<o...........lonF;O;
,..... 1Iow=
~::-bIo ..... nf... 'k1

164
Lilhopssc hwanlesii Dinlcr
M[SEM8RYANTU[MACH..,[

~~i~",:,,;n,,!t~:~~::;r:~fk,~rz:'~ ~~~j~\;~;:~:' ~d ..h·b,,,wn wilh

~~~~~: ;:~:~~~~ ;~intcr & SchwlI ntes) N, E. Sr. Lithopssc hwantesii 'Triebn~ri'
,"W "."VA"'HHMUAI
8ody2"';,5cmind~,mctor;rouoo""d. upp"" u '{.K'erom ·""'R"'Yorc"'.m 8u<Iy )-4cm j"d;"m~~~'; "'"'II"", tbou'e, uPP'" .urla.x 11.II.g,,')I ur pale
~:::''''''';'h~''''''()'kO/lin"",yelto'''flo'''M"< ~:,:~:;th broken .... tWOl"k of dn"""",,, lin ..; yollowII",,·....

165
:~~~~~~!:~:~~l::<:~:,~. Br ~~~~~~~::;:::.~:l~:~~e!
Body 1.5 crn ind"'mctOI. p"' ... g"'y,,, .....,j ,,·,thoWk1"rg ...."n.. hd<>t>""· ... Body2_)'nl;nd .. ''''.. c"r.dc~1'[i»u''';g'''y'''ilhfi,....Jyfi .........-dd."Iu."''
mglhelop;yo-IIo ... llo,,"«S. "'ind""·"',·.,...,Jwilhlwo""nd"'.. ~... llowflowon.Av.,.;.,ble,pt"<1 ...
/j'h,)psl"...d"S<h .. on ...... 5outhAfrica(C~l"'rrovi"",,)
/IIamibi •. 50 ulhM"""(DopcPN'o;",,,,)

166
167
M~rremiJlub~ros.I Rcndl<,

=~'j",;"';:~~~)~;~lk'::~'\'~:"I!~~=IlOn(""'Y
Trop;c..lMn<oo

168
169
Monarlenium guenlh eri p~~ MonJdeniuminl'enuslum N.E.Br

Tub"rrul.t".tl'1r,.1$-2!i<mUO'm<>rchighwllhl-3>pi"''',gr,,-,,,k,,,,., Tu!x-rou:.,ool;fl<"lohy.<jlinJ,,,,.ISlcm.108(leml on.o\;darkl\"""'..uc<u"-,,,1


l-1cmlon~criopcd.tap"~;K""'ni$lH"hi!Oinn_~ 1<l.w... 4 em Ion~ ,,'iln 1"'" ~""'" ,'e;n" w"i~'"1"nk inA"",..,,,......,
Kerl)" K,''')~

170
~=~:I~~~~' rhizo phorum P.R.O. Bally Mon. dcniurn schu bei (Pax) N.E.&-.

~~~~~:~:a~~~~n:'~
Kft'If'
...... -
~~~'2d.o":\'!;=~~,C:~~~!~dO;:~-';
T"",-,nlA,Zunbob...,·

I7l
MllnadeniumstapelillidesPax ~!~~s:~~hce:~~nJgens is I'rae,.;cr
En'<''''·m•. wkkha'eJecumb<''' .. hc",,'''''''I'''"15(Jnw"!;.w'th'ub... n;I .... t....>w,>f.>n.."J;,>g.lll'\lb;K"'b"'... ~n."",al~m.. ',·lca'·''S2.S'''"lung ..·' ' k
AIT.,ng<.'d in spt .... I.; g''''''' ".,·""l-S.m!ong""",,,on,<:mwin'<:l"rt.ofyru,ng gro<Wod urp;!'ts,,'I,",.. ;y~lInwisk ..<:,......, A""·('t,,
'~'n"pll'k,nflu~ C"'''''Y 1•1,,",hrr''''e';fd
Kenya,T.",.ol"'A

In
~!~~:~~ce:A~ubuas§icaulis (Kuntze ) I'raeger ~~~:~~~ia echinala MiG.
G""",>lcm.I· 2cmIQtlg.den".". ""'fiI-.,ytul<lricalJ.,,,,,.. I ern IOlll'-h.:uryflo.> ...
oring .....".;rroll<>w<-.. ~~t:;'.rcll':,';'.;~~~:'<!?d~ \:'.~~::::::!~:;,,'!.!",~'j,:~'~I~~~~"" ],,<,w>;
Co"",), Isl.mds (Gom<",. La Palm.:>. T~'l<'fiM Mal.:ly.;"

173
NA M AQ lI AI\T lI lIS

Namibi~ ponderos~ (Dintcr) Dintcr &: Schwantcs NoJinarecurvata Lem


"'E5,,,,.",',,'tHF".\(tM
~!::~ d~~:;'~;'!j~~~'tj~~.':~:;:O~~,\~18 rnm w;d~ furm clump'
NOlechidnopsis
tcssellata {Pi!1ans)
la\'r~nos&B1Kk

"".n.... 6-lo lO-an~I<d.I"m.'


11) -20cm~ ish;lubctdcs
tipp«i""ilh""lt,gf<'<"l\.'pi,.,.,..

l:~P~tb':;~7.i~~IC\~~W
c.."'I1" .... t."...!L!a PiILo""
50uth Africa (C~I"'I'rovi""")

Nycleranthussp.
MI:5[MBR\ANTlllMACEAE
~;..=llf:i'~~""'nof.n
5oulhAfrk.(Copcl'rovinwl

Nyctl'T~nthus
viridiflorus (Aitonj
Schwantes
Mr5E\t8RVA>ITIiEMAC£AE

r~r~~~~?t~.::·
2-4<mlong.pcrstsling,,"
:;:n·t,,:::}t"'dryin~,,,1i'

rr.~.:::~~:n~·6~ridjtl"'"'
50uth Africa (C'p"I'rovince)

~~~:~~~~~~I~~:~EU:[L. Bol.
u.. ".,,- 1-2"", Icn~grry tod.,. s"",n lo,w", 1-2 em long with d",,'~'o
marsu>'; .. h\t~flo"""",,,
Namib;.o!SouthAlrica(u ll lpNA"""l".Lond)

17S
Oper(uliur)'~dt(uyi llrl'lTier

~r."",", lO-IO",,~h.b<""'h'~r:;..,.J .......~"''''''l\ ..'....' .•n...llp'''...


.... '''''.hnld .. ''''Adrv..:.oo< ...
"0<1."",-",.<

~~~:~~~n~~~,~,Y,~~~':~~ull\'ri 1..1I'is
Ob..,"'.... Ib<oJ" ... Z<n,I""S""d.1'mind"','...1l~.sn'''''"Vl·11o''.,h.Ir.....
I""·,ll.Iol...., 1 2,mlon~"','h'"tn-.lllC<'n',,·'"d"ws,ru<T'lI<hnnw' ....
Sou,h A,nc. «_'.!". 1"""'''''''1

. . ._ _ . . .""'. ...._ _ Ophth.. lmophyllum


lalum Tisch
.... ,,"U\ .. ~Wf ... MI· .. 1
Cylmdno;.albod~IOH<III

1!!;i~~ '~'lC~~~i:I Iur:;!o.nd2.". ........ "r:;~1


t~ ~;~~~~~\ont.
South Alrio.. \C."'· I',,,, ,,,,,·)

~~~~::I\~~f,~,)~I~~~~~in'tri Schwantes
Soh ...., .... Ioo:~IoncI"'.lb(oJ .... 2-l="""'-r:;..............",h,.. ·~"""'h"~-.l •."h
~..;,;:I<.n'tur.nI"·'nd" .....;J'ln~n.c..."'"'-

\7'
~r~.~~~~~~r:.}'~~~~~lfongum (\ .E. Ik) Ti!>Ch.
~b~~'~;;ht~~.~~.:~I~~;'~~~:'~~~'~;I~'~f~t;~~\':,~i\.'.~'." ' ....,11"" i,h
r>.:.m i bio/S.",,~ Afr.... ,l.ot'I,·\J.•"''''lu.,I.lIIJf

r-=:::::;;;;;O.IIi:J!iiiI ~;~~~~:t!Y~~I~,Y~~,::~,
OIi"··~r"",,boJi""2·2-S<m
I""g.,nd l.'icrn"·..:I~"';lh
m,"utcpap'IL1~;'~'TI>I"'"-"r"
Iol","-hil"n..,,~~
t\am,b'd/5ou11t AfTko (L,IIl,'
.....,-"....... t\., .... qu.bndl

Ophthalmophyllum
praesectum (N.E.ijr.)
SchwJnte.
'''~F''""YA''''''''M'FA'
?t~~n~~~~t~ ~:u~~~
\"j",lo,,·.;form1Mg<m.n
rlu"' I,,;I""''':!-;mmlon!l'
I-',,,~,,h ,,.,1.1 n",w",
~o~"'tt.",,,,,,p,",,,,,"'"m
"'~'lhAfn~A(t.1'" 1'1"0\',""')

177
?~~~_:,:~~:~:!a (L) Haw Orbeopsiscaudala (Y E.lJr.)l..t>ach

A<lo.... upoflhc/lo""'" 1-I'<"CI'I" 'egr...-n""m'b-10nn l""g:"T.'ghl" pon)'''-"'-~hrnor.'h.lnl<m


l..,ng.)"n"wIlUWCf"> WHh nu,,,,,,,,",, pUlpl. dot>
G.-,.llu"'.~''''t.r-..r.tI".C /'''''8,",i/,_, Obenn.
M"'.,,..i. '1,mJl'.,Dmb,bwo

178
~r~o>~~~~!:~:rubesccns A. Berger
!..";i~"'c~~:::~~~ ;~1';',.".:r.: ~~.h...'i,,:;,~nn~r,,~~r;;."1;"'''' I........ "f Z MI."",.
Clu .... (nortltl. J""'m.K,,"'"

179
?~~~<~:A~~pensis L.H. Bailey ~~~<~:AJ:pidocaul i s5chJtr.
DWMf ,h",b '0 l~ em. , .• 11; ,hk~ b, • ...,h .. «"....,.d ",'h~A"; I .~j"", .. k>.t
F.::'~ ~1&:k:S~~:'j~''::.; ~'~J~:~,:::-~::"rt1~;:~:>~~;\~=Z,I::~.~'1 t>.a_;&',""nl.O\·"'3-<>rml""~y.llu"· n""c",,,
";"u,hAIri'A((;Apo'l'r",·;nco) ".m;!>i./<;nu,hAIri... (I .,ttlpNam'q",lJnd)

180
181
~:~!;J:~!~~ oviferum JA Purpus ~~[!:.i:,m barunii var, windsorii Pichon

PruoA,ak', wh,te.t.,,,,,, 1(}-15cm long: whlt~ .... " .. 2-1cm long 000 2-3 em Cl~ .terns to IOcm l"diOII><'t<'l'; thick, cy~ooric.l b,~n<he<; short 'pi_;
widc, n.'<IfI""..,... 1"',...... t ..nd<nfbrnnrl'k><;r<!<I/1""· ......
M,ld......""" CITI'.5Arp I

182
w 'Ie",m
(;.00 .... In.u.m..i,,,
",lhcyllndnull>nn<M t.. 70,m (K"""" Lol<.'tal
.pmod~'2m);"J""",""m","",g;d.rI'l;"""''''''''''2 1<m ~'''''thh''''
"'hl~h.o,,,,,,,, ... ,,.......... rI,,,,,.}~I"""'~
\bd.og;o."''' CITESAppU

~:!;~~[m deul)'i I'oiss


Short. ......... fd.lubrrouo<.""""w.,<mhixh.tudd""lybo~rochi .... 'nLhiA
_"'"<~j-6",,Ionf;.nd4-5nn,,-,d~.rT.nt;"'l'n ....... ,.... I""""'''.''''
onIhoolt~Lh.o;"" ;lI"""'."b·"·hil~llu_

I ~.r t~"M' 1

183
1'~ch ypodium s u((ulenturn OC.

('Aud"" t"',
<n> m d ... m....... !(t""'m); ,,, ..l,...~,,,,,",t i" !> ••
~""'d".J,t"""2[k,11<onl"ns; J..",·,... 4,·"'I,,n~,'nJl<m,,·,deb<'mt'.lonS
b,'.".
li ..... ~\.

",-,,..;hoot,, >p'n,-~ I 2 em Jon); u>lI •• lIy ill p.m>.l'in~ n""'",,,


s.,uthAf,ic"K"I,,'P""'HK'" CITES IIpp. II

184
Pel~rgonium
cotyledonis{UL'Ht?r
Thick • ...,n,tollfcmhiSk:
ruu"d,,J,g ......lIl~..'.,
~~U~~~ :;:~k:~~~, ~~:::.
:~i:; ii~~-..I!,::Y h.o,r; bduw;
5.,,", H"I~n"

pel~rgonium
crithmifolium
GG.Smith
Su<nJl,lll,br.ncht..l ...,,,,,.,,,
5(Jcmhij\hwilhuh."ul."

~~7:!~~~~~£E~

185
~~~~~~~~ :~~ lobatum (BurmJ.) L'HCr.
PI'I\I20-J(l~"'hi;;h;L\lg"lub...,.., ... \=-dwilhbru"'n""k;>hull'(CI\"';
3- Iobod. hal,),.!;""'" I~.\"n to 300» in ,j0.1ml'l"; ,I",~ p",pl~ A""......
SoYlh Afrk. (Cn!", Pro.i"",)

186
f>eperomiadsperul~ HulchiSOll& Rnuh

5u=I""'_"e""k'IO"",h~hw ; th > h ort;nt,·m<><l<.. ;I""·"'I-8cm1ongand


9 m"'wid~.".,,g<odind,.,...,rows.I"'Mlucont.bove.green-greybetov.·;
~:~;",.grt1'flOW<'fS

187
Piaranthus framcsii Pilians

:~~~i!r"':~h:~=~;t!·t~t:;:'t,7.~j~~';Zi~'~"~;:. \- I.S<m in
South Afnc. <Cap"P"",mcd

188
~~~~~II:::c~:~l iduS CA. Liickh.
;;:'~f.:"",gl'*"-t"obl"ng">i>!;<:un:ly .nsl<d .."n"2-3t"mkmg;),,Ilow

Snuth Afrioo {C>I""f'r<l\""",)

~~~!~::lhuS amboinirus (Lourtcig) Spreng Pleiospiloscompactussubsp.canus (Haw.l


II.E,K. Hartmann & Ul-de
;:;!t7~i~=T~:~~..:r 1~~:fg~;..:;~~:t!';;~~i:;-~~:h~::: ~!= 4= 5<crnk><;..clump-fOfminl\""""t""of4-8~"'Y·g""''II,pu'l'I .... 'ng<>dI~Av .. t''
TropK'olAf""" 9rrnlong:)"nowfiowc ...
P"""",!",~",,,". (II,"",)LBoI .
s.,uth Afnr-. (C.po P"",,,,,,,,)

189
~~~~~r.i!~~,~~1r~~·datuS(Haw')Sch",alltes 1'lumeriarubraL.
C...,.1\""'" or b'""mi,b tcaVM ·\-9cm Ion)\. unito..l., b.lsc/or 1 em.nd ","h
/la, upr"'- ",<I""" ron'·~. b;ock and pmm,...-nl dols; , ... lIow flow ...
Sou'hAIrkA(l·~p"I''''''"i~)

190
Portulaca pilosa L. Pterodiscusaurantiacus Welw

IInnl .... h.p"'t ... "de'''':l(!<mi" di.mo:t'''Wl,h"",· ... ].hICkbfilnche>; ... ".
Eit~:~~~~: 'k~{,',~~~t:~!~kto~~~~~~~~7~~~::~f~:~a ~~:; blui>h'S""n I,""'" 10-15cm loog with .i~ u"c '"''1\'''5. ycllow

\1~'rro. USA (<.Outlw"!1

191
~~:e,~~~a.~~h a mJlvifnlia Engl ~~~~~~~~:;;:rburkei N.E. Hr.
~::~~:;:,~~f~~~::; In cl,.'In"~r prod»nng '''-,nong. !P.,"- ",'''': g""" (;rry<,,,M, 1I-15nn in di~",," .. r. ",u,h·I>r.lnd...,,; <WI\< IS,1n 1001:0 ~""Y
s(,"'nlc~'·~.,.3,.,.,lung;purpk'f1u"'''''''
llo4,w.,,,,,. "'omibh. s"",h Af,i<> (Ir.lM".l.,I,

192
~~~~::~~~~~~~~:::;:Ai'~ii Schwantes
Shrub2O-JOcmhij\h;Slcms w i,h.ho"in'crnodco,hllobi.,l,polcbluish'<>
!;~'~ ~"'.... \_~<'" kmg; yellow Am.........
Soo,hllfn",(Col"'l'rov;"",,)

193
Rosubriachr}'!hInth~ (Boiss.lTakhl. Rus(hia nassa (L Bo1.) Schwant es
R.....~t..,. 2.5 om in d .. m<:tc'f; gn..., ... twit)' ka"'" 2-3 en, Ion~ wIth ruundod ~~:~''1lh"re<luTp'''''t''t •• tem.':8Laurou.gr.,.n"'''-''1_2cmlons:wh,te
'po>';y.llowfIoWffS
Turlu.')'(,;outhan.J..,uthw.."t) 5.>uthArrica(Cap"Pruvina)

194
Ruschia sMmenlos.! (Haw.) Schw,m les

;~r~:;~r~!rl;'::t~:~~:~~~C:~1d~t~'~~"~"f.:~;.~:t~d11; :i~·;
Sou,hA(",,(C'p"I'rovinccl

Rus{hianlhusfa({alus LBol.
MbH,~n~'1ltn_,,~nAl

Shrub .. ilhlub<""'."-'Ob; >l"'''>''''',.....'\!", th r~'''''I''.lIfdryl!'..'''';u ..,'<Iu.r1


~~l<blu.,..~"""'""w.... tllr> l,'ll""3--Scmlong..bo""'inp";"'y"llow

195
196
198
!kh"·.ntesi~ roedebuschii Dinler ~::~:J:~udrlor~ Saker
."ls('un"'rnl\'!M. l"~

~:~£t~,::_~k::h~~~~~~·'mdS"ilh SonoUhuJ .... """"""'6.ndfurm,"lI LorgedW'tl.... poitKn."' .......... ""hduk


;;:;"~.:;:~n.,.."'"

199
S~dumiClYL Sedum ~1I~ntoides Ro!.e

l\"mrrtIUO_f"""lIl.., .. oIon/"tlrung ...... <I~/lll'.:<"' .. <k..J.rrud· ...


ycllu".SI'«"Ir.,..,. f-2mm ""'G-r"""'·tl<>oo......
I<.n"""", .... "..•. lurop.·

200
SedumhinloniiR.T.Clau5t'n

51<-"" 5-10 em Ion~ ,po''''''' bt..",h<.", d~'"d~ )w'ry k-.l,."" 1.5-2 em 1<>"5
~rT.,n~'.J in , __U"" ,,·hi'~ now,....
\In.ru

201
~::~~t~~~~~rOllense R.I'. Murray
Stm,l.rt<>S."'<J""'b"\""'!ht.'gt"I .... _;rell''',·n''w~ .....
C.n.ry IsI,nd. (L1nz.>rote)

SedumlanceolJtum Torr. Sedum middendorffianum (Maxim.) A.G. Hori~.

~~~:~It~:~~:,~t::::~f\:'..~~~~~r"j'~:'~~~:~'~ b~i!"l! ~,,"r,::.:~.~~:~~~J~;:~;.,~~r't,b1~t::'J.7;i;h:fu'~·b~~~


wj,hydluwn",,·crs. ~uins)".. lIowll"w ....
~~;;,<ho>I""" IIntton; s. <I'''''I",',Iu'N v.r . • u""'p",um Frod ~ s. ",/lolp","m KorN.. MMlIo1i •• Sibcri.

202
~~S~~l~::~~ A. Gray
\tu,h b.. nd..._-.d"~om."..",.""..",.I .... I)·~bn""'And moIl~g~tnod"":glAUC'
~~~,~,:,""n 1"""... 1-1.5= Iung.nJ 4 mm ,,'Hl~ ,,,,..,,ge",, ,n """-1".~; n,,",cI'>

USA (V"}pn,at"Alabamo)

203
SM\.'lI>h",b2Scmh,~ I~nu n "''''''''.M">b'''''''''' \gr,,,'''I>a><·,l;J90lgn.'-'''
l",v .. "-;,hrt'd1i1',1-20»Inns·wrou ndin-,iQn<ro wdru.lonl\.,,,m.;

r~.:l~:~~:~~:n in Me>iro

204
Scdum s.artor;anum Boi"s,

=:~:~~:;l:';~2,~~~~~~;n ,,, ~r"~-grwn ~~" .. In.ngul~r


G"-""","'rm<'1'Y"~o< l,,,,a(""bi.)

~::~~~!~eE~~ldii 'Foliis Medio-m;egatis' SedumspuriumM.Bieb.

r~~~al~~<~ ;~?cmcm~,'::~~p~~~<1o':~~~:~::I~~~t:.i::,~.~~~;. ~~';;' ~~J~~~~,":nfl~~=~~;~dl!'r~!:~~,~~~~~r::.,!~~8


,.ri<w'..... form ,,...,)....11,,,,' &./um<ili.,..,S"..."" S.<"".~c>t"'" i<u<k; 5. trnM"", 8",,,,,.; S. d""r,,,,~,,u,,,
Joron Donn,S.I.,;'-umlloiso.
CIS (Arm,,,,,.. A,... rbo,dd, .n, C"><"'r'g,aj

205
=~:~:~~ ~t1anlicum O.M. Ball
Small .hrub wilh ,;en,..1 br~n<hes ansing from 10..-"" parts. <Ii OMIl<; blui,h·
~~rui""",,Ie.'·"'3<mlo~S;Y"1Iowflow,..... ;~~~';~tflo!!~~dl~it~h:~~~~~~~I!~~';"~
".~

206
Sempervj\lum cantabricum J.A. Hu ber ~::r~I:~~~~ cilio5um vaT. galicium A.C. Sm.

~~!i:drl:(::.~,.~:,'~:::"~~;~k~=~ . ''''to4<mlonganJ
Ruo.o.'IIo." 10 2.5<nt indiarn<:lcrw ilh long.lOlons;J; ....,..ish-s."""'Ic....,. .... ilh
ohortholi,",flower'""",,,lO"<."J"J\~igh;}"plJnwA"""''',
Spotn Europe (Cali<i,. M.o<:odonia)

'"7
;:~~~el~~i:~~ kindingeri Adamovic Scmpcr\'il'um monl~num I'dT. bum~lii W<'tts!. ex I [,,}'ek

1~~': i~~~~~~~f;:I%';'~~: ~:~r~r.~~~~~tlb:~SI.nd~l.r, half} =;'i.:!~~I:~~:~~~~~i'.:.r:"h.~~7 I~a"., wIth ddrk "P" ,,,,Ie!
Europ"IM.><c'<ionia) Fr.I""'/It"ty/~" "'l'"'lu 'h"''"'t Atl"'I'}"n..~,,~... )

208
;::~~r:~:~~ purnilurn M. Bieb Sl'mpl'rvivum tl'(toru lll var, italicum (Ricci) Zonn

F"'dy ..,sf",~'iI\S ms<-~t"" I 2em;n diamc"'" ~r""n. ~1'ndul,r.II.';ry ""' ...... Koso1t""l-iIcmind;'m",..-;nunwrou.II..iry);"",,,I~...... wi'horw"hO\1'
~~"(~~:::;'l~In ..p!1'T'''' flo",,,,, ,,·;tll!,,'" ma'll'n." A '·.n.lbl,,"f'<'<1"'. b",wnllp".nd .."thcihat~rna'll"",;p"'kn"'",".... ""th ""truw"'rut~""''ll;ns
PLlIl'poorogr.phcdinh.lblt.l.
Smlp<'l1""'" Iful,,"m R"",i
IMly(Ahru>.>n.L",io)

209
~::~~e:~:~~ wulfenii Hoppe Sempervivum lectorum 'BiAnco di DAniele' Zonn
c."'>!,.l,,(;v.~
"d......~up ...r'l ... "".... ~'" R""""""5-<I=lnd ... mel"'T;!;"1'-sre<"',halry"",,~';fl,,w""',,"hlto
l'aly(\'.I I ~III"";lim"odd",nbu""n)

210
~~~~T:~ iculatus (L.f.) S\:h. Bip.

i:~,~~::2~~:~~".!r&1~~~~~~ ;jj;'~hl~!.~h g~y m•.,klngs;


Suulh Atrk;t{Capc 1'""",,,,)

211
~necioh~worthii (SI<i.)Steud.
co"I'o5flI\~
SlorublOlI'l""h""; ..1"'"'Y1l......,1f,o_5C111Ionj!.,...• • lnJPll"~1o..0<alI&'

~~Iinn.
5nuthAfrI<".((.·~"" I 'r<w; .......)

212
~~:~~'.:ndulu 5 (Forssk.) Sch. Sip.

~~:;'':I;'''';~=io>o~~~:::.':.~~anthcn~
n."",,..nd~"<k>f.;o.UOC
BI\iop<.I.K""~~<"""""',u.Som.>"'. y .........

213
:::'~T~~mpc.rvivus (DC) Sch. Bip

~~~~TfZ~~J.~$al~~·~~J:~i;,:rn:~~;~~7~~:~
~~~~=C""IJC
214
~~~~~~~:C~~lbrrtii Kcr<ludrrn ~!~:.r:,!!-~~af }'unnAnl'nsi~ (Franch.) A Berger
(·yI,"'t"..~I,nllbo.t.<i,,,,b'''S'''''''''1u3rnloo5' ......ltuJ .......... ).Iuk'' . "~~(IO;IO).l>I_t....Vftionnmg~ ......"",k'a_",,""'"
C.t:;;~~~i~~~~_ 2O<m~ir6i~nm~.n'I""t.s-n w"lt<hnnwlu"'N,,<; .. I\'h>hflo,.-""' ... rthn:dt;p.o.
erun...(Y ....... n)
M.>dJIt-~r
Stapelia grandinora Masson

G"-'''n.4-.n~''',j",-'''>1<>30..,,, "-,lh"... ,IlI,...1h.nddc"""·lycu>',,n.<i w'lh,",,~


ruir<;",rli""",,,ry. "",hO({)<O, 1Nv<!<. PU'1'Ii"IO bIJd-pur-pl.-n"....,,,,, ,,-ilh
('h'I~"",~,n.''''''J'''fr''m",,,," ..fyo,,ng'IL"fJl'
sr·I"'''·m/oj~".M.l,"""" . S_/Lr'·''WJ, .. \lE.8,
Lc;o(oo,SoulhMJK"i>(C.""P,m''''''')
~~r:~,i=;:~~~:r17.iae N.E. Br
Il"'Y,wngl«l"""".8-IO<mh\gh,,"\lh~·"".I'm.1IlI"'lh;rud\m""t.>'Y.

:;~~A~;:~:~:~~~;bLtck o. d"k brown "o,,"er>

217
~~~~::~!~~:,demYi Chou;.; Stomalium ermininllm (l1.1W.) 5chw'}nll'S

Eftct.. ... , .. S-."Jdo..J ...,.,.. 10 <m rup.. ~n~"''' wI,h ~It ....... n~ """h nl'!""'d
",.h ''''''" ... ~ mm lonK' d.or~ 1"'' '. . 'u ..... ,... /1"...."'" 2-2 ~ <In h'ng ~~3:1~i."!rt~,· r..~~IoJth,,!,'f,d~~~ ~'.:~~~'7';',j~':.~~"-"h
M.>d0V>C.'
Sou,h M""o (C.I'" I'ru' ""'d

218
Talinum pJniculalum Uacq-l Gacrln.

Tub.-nou, roo! w'lh~"'J. r,n~


loyd""'· no.......
-.I",,,tu I m h,sl\;g ...."n 1e"'·,,,,!Oem lon~ ",J

CI·"lr.I • .-.lN""h""",n,~(,,",,,lhl·mSt.>t..,,)

219
~~!~:~~~~.~::~!~(Schwantes) H.E.K. Hartmann & lic-'de ..
~!:v;r~: ~[f~~tjit$ii IlA Dyer
Tn."&,,IA'·"""""'tl'"'Y IN, ... J..-4<m ionllwilhdulo._l"Jband.-,_ Ilotkj;twfttobn)wn,6-tolJ..onpN_8-IOonhigh""",'f\'\lwdh""",JI
lov..... owf""'bum.on ..... , <lumf".S-'Ocm;nd~ ........ I-2poil'f04IN"""
frompckohoot.)-.:Ik>wfk>w<>n =:"-::~': )~='~~:;.fli..~~.~=r;~ ItIln>m>U5 "'"
PIt;.p;Jotpn-,,,,," (M.rluIh) Sckw.nteo Swth Alrico (T ......, ..... I)
Swlh Afnao (C.I""'mY;nno)

210
~:~:~,~~~~"f~~~~:;~i~h
Clump-f"mlin~ """"",or ':Hi I"'ir<nf "',"..-;;~""'nww,,.,,'<l
wn ... ~''''''n
infull.un)I~""",2cmlon6''''h 0.., uPI"""",rfao.·.mundod k",,1 tok,,,"~,
su,fOtt.,.dlube,,,,L,'eh?;ydl,,,,flo"l'N.
m,hMn<a(C.p<" I'mv,.....,)

Tit~rt0l'sis hugo·schlt'<:hteri (Tisch) Dinlt'J" & Sch"'~llt ('S Trades(antia siliamontana Mal1Jdn
\'[S[MBn ... "fll[" ... n ... [

~~:k!f~'~,~"~,;~~~~~~::"",:,;:-~~:~,~~~t~udb.~II:::~~~~: 0' ;:~·f~~~~~~:t7~:~~~·fl,:~.~m Iong.nd CO"~ ,,,In wn,'~


yellow Oo,,-..'f5 Bd,,~, Gu~t<"""t". M~>KO

22 1
Trichociulon {Javum l\.E. Ilr. Trichodiadema bulbosum (Haw.) Schwantes
Mr5E MeRVANTtlfMACfAE

~~:;~~f'i5;:~'~~u~;~:%~L;!~!~n~~ :':,~,U; and T"b<>fou. """" wi!~ ".ms!O 20 em high; ~f1'y·g,.,..n J.... v.. ~ "'''' Jong and
3 "'''' thock with "",,,,.1 ... h i t~ bn.tlto<at tlPO; rY<l f1u".~rS.
"""'h Afri<. (C'l"'i'ro""""i
Sw'hAfdOl(Dj><"PYu'i'''''·)

222
Trichodi.,dem~ den sum (I [al\'.) Schl\'imtC"i
"''Innu ... """,, ... c .... ,
Tuborouo~ .. 1t".han_'iV"<nl<.''Ql.s-2<m\on5u.d4mmtlUd
wilkl!l-2'i ..·ki"'''"'' ...... AI''I'''-(.nn'''''n.,..,''''"5.
Soutk Ai""... (Col'" Pro,;"",)

Tri{hodi~dem~ pel'rsii L Ilot. Tromotriche revolut~ (M<lssun) Hal\',


\ll!>l"UV ... \II!£"IAl..lAl
=~=,lO<Tnhigh;5' ..... It....... lnnlong ..,'h~...... lbrownbri5tlfo<; ~-.s;::ww,.;!,~v!"JI<'5O!I""'JoJ<m~p"<pI<fk>w<-nW;"''''''''rvro
5",uhAln..~(C.I"' I '",ml(") S'oprl""""0'4 \.1_
Soulh Atnat (C.P<'l'rovin<t')

223
~!~:~oc~f:arsonii (Schonland) Toelken ~!~=~Aoc~:~hal'fl'riantls (Dinter) Toelken
s ....... ""...,...d ,,~'h Srryish p""lin~ ",rk;g"'}'~ togreyioh·bmwn Je,o~ . .; "ubo..,..."'"r:hr.I'oC"N!d .... """"25<m;sr-n- bm....nl@Av.. S-2IJ<miong;
l"'wb"'''nflow .... yell""'-S"""'fIo,,·~ ...
CDtyWonI~I"'"'!""""' .. I""'lIn.;C "...,.."'IJISc-honL>r>d ~~:-::"'~I!III)in'.".
N~[J\ibi.a,SOUthMri<a(Cap"Provint1')

224
Ty lecodontornlos um
Toelken
s..,m<toZSannighand2an
,nd .. m"'",",w,er.:d,.,th
dar~bro"·ntowhitcborl...

f:..7~ct~\!':,i:.~n
round<d api""";y<:lIowish."
S.-..fl""·......
Sou,h Africo (CopePrtwi"c<»

Tylecodonwallichii
(1 Inrv.) Toclken

Mu<h"br.""h.:d ..~m'"l.5m
higltoo,..".<d with IMmtWIU
ofoldl~.,"";borl<~!!ng
m>m''''mbo..,;gn-y"S'''''n
~~]2cmlong;y<:Uow

CDlylNo"o-..lIichiill.rv
SouIhMrka(CapeProv;"",,1

Uncarinadecaryi
Humbert

;;:~~3!:li:'!::i;"'1h
1In.-yish-<><]""b.J.rl<;lrianguW
le.'.... ~<m""'g..1>""'"
uppn.urf.rero,"tn."dWlIh
small hairo,lowc<,uff"",
Wry;)"'!!o"'!!"".......
M.d"l!-'"""r

225
VK'WoflheNdmibDe.en wit h Wrl",il..-hio ~n<l thc.UI""".fomily

Welwilsch iamirabilisH ook.f Xerosicyos perrieri Humbcrl

Climb"'I\.(cn .. , "oody dli>.u<, (hId.. h~hl !In" ", k""'co2 em k"'!I~"d 18 m m


wide; whili.h flm'......

226
Zygosiryos trip~rtitus Humbert

~~~:;,~: ;.,oi~2'3 ~~e::;~r.~~ ~~~~,~i,::\~"!J~3e~ I,,,,sand


M,dogosc"

227
CONVENTION ON
INTERNATIONAL TRADE IN
ENDANGERED SPECIES
C ITES Secre ta rial, PO Box 456, 1219 Chatelai ne, Ge neva, Swilz(!rl.md

Habitat clt'Struction is thl! major cause of the dt!Cline in wildlife populations, but
the second most important thr<'ilt is the exploi tation uf plants ilnd animals (and
their products) of wild origin. [t is when this exploit.llion leads to intcrn<ltion,ll
trade that C ITES, the Convention on International Trade in Endangered Species
of Wild Filuna and Flora, plays an important role.
CITES contributes to nature conservation by regulating the intcrniltional tTilde
through a system of permits and controls, which are issued and implemented by
the exporting and importing (l)UntriL'S. In July 1993 120 countries were party to
CITES. The word 'trade' refers to all intcmntional movements of specimens of
species listed in its AppendiCes, and it includes plants carried by tourists, large
commercial consignments and material for scientific purposes. However, there
arc certain exemptions for registered scientific institutions. Through this control
system, alll'arties assist in k.....ping Tare and threatened wildlife where it belongs
-in nature reserves and in ilsnatural habitat.
The CiTES controt system also permits trade in specimens from the wild up to
a levellhat is not detrimental to thesurvivill of the species in the wild. The inter-
national trade in animals and plants of wild origin involves a large number of
spt..'Cimens and has great value. The number of wild plants traded internationally
increased gre,ltly after the Second World War when people travelled more freely
and to more distant countries. In addition, the incTe,lsed usc of central heating
alluw~ more and mure pt:'llple 10 keep specimens from tropical regiOns.

II OW DOES CITES WO n K!'


The purpose of CITES can perhaps best be illustrated by quuling fmm the pre-
amble to the Convention.
• Rt.>COgni:.:ing that wild fauna and flora in their many beautiful and varied
forms are an irreplaceable part of the naturill systems uf the earth, which
must be protccted for this illld fur the generations to come;
• Conscious of the evcr-growing value of wild fauna and flora from aL'Sthetic,
scientific, cultural, recreational a nd ccunomic points of view;
• Rl'Cognizing that peoples and states are, and should be, the best protectors of
their own wild fauna and flora

Through CITES, sustainilble trilde in spe.::imens of certilin spe.::ies is regulilted,


recognizing ilt the S<lmc time that countrit.'S have a right to exploit their naturill
resourccs or not to permit such exploitation should they so decide. The degree of
protection afforded to a species by the Convention depends on the Appendix in
which it is listed.
CITES
• Appendix I includes all those sJX>Cies of which trade in spt.'Cimens of wild
origin is jX'nnitted only in exceptional circumst;l1lce$
• Appendix II contains those spt!Cies of which commercial or private trade in
wild sJX>Cimens is permitted as long as it is regulated to ensure that it is sus-
tainable
• Appendix 111 contains the speck'S for which one country has asked other
C[TES sign.ltorics for ao;sistance in their protection.

The inclusion of species in Appendices! ,md II requires a two-thirds majority


dl'Cision of the oountril'S that are party to CITES. The listing in ApJX'ndix III can
be done by an individual country. No cacti or other succulents are listed in
Appendix 111; all cacti Jre included in Appendix I or [I.

WILO - CI{OW\' SI'ECI.\IENS


The regulntion of tradl.' in the species concerned is provided by a system of
permits. Importation for commercial purposes of specimens of species listed in
Appendix [ of wild origin is not allowed. Trade that is not commercial- for sci-
entific purposes, for example - rt.'<juircs that the country of importation issu('S an
import permit before an export permit may be issued by the exportin g country.
Commercial trade in specimens of species listed in Appendix II of wild origin
b allow(>d on the condition that an export permit is issued by the country of
export. [n the case of rL'-export of specimens that have beo:'n imported from
another country, a re--cxport certificate may be issu(>d once it has been provt.>U
that the specimens were legally imported.
These are the basic re<Juirements as incloded in the text of the Convention.
However, e<lch country may take stricter measures if it wishes. For example,
some countries - Pem, BrJ7.ilnnd Mexico - hnve decided not to Jllow the export
of any wild pl<lnts from their territories. This means that they do not issue export
pt'rmits for wild-coJlecti!d plants. l:ly t<lking this me<lsure they ensure that, if
plant., arc taken illegally, Ihey may nol be imported legally inlo another country.
The EuropeJn Community requires thJt nn import permit or certificate be issued
for any importation of CITES specimens into the territory of ils member st,ltes.

AHT IFI CIALLY PBOI'I\GATED PI ,ANTS


A less strict regime applies to artificially propagated plants. If these plants nrc of
species listed in Appendix I they are treated as if they were in Appendix II, so
that only an export licence is rt'quircd. Although an import licence is not
required under CITES, some countries may still demand it as a consequence of
stricterml'<lSures.
Artificially propagated plants of species listed in Appendix Jl are subject to a
number of exemptions from CITES controls. Raska! seedlings, cut flowers,
S('('(\s <lnd po11en are not covered by the Convention. They may <llso be trad("Ci
with a certificate of artificial propagation. A number of Europenn countries usc
the phytosanitary certificate for this purpose.

The above is n brief summary of the work of CITF..5. The Convention is imple-
mented in Illany other ways, and interested readers are di rected to Tile Evolllliml
ofClTES by W. Wijnstekers (see Further Reading)

229
INTERNATIONAL
ORGANIZATION FOR
SUCCULENT
PLANTS STUDY
CODE OF CONDUCT
Sara Oldfield

This Code was first published in 1990 in volume 10 (supp lement to no. 4) of
Pillute GraSSI', the journal of the Ita!i(ln Cactus nnd Succulent Society. and it h,'5
since been printed in severn! specialist journals. It is reprinted with kind permis-
sion of the author, 5.1ra Old field. The original edition of the Code contains much
useful information that, for reasons of space, CaIUlO\ be included here. Copies of
the fuJI Code may be oblainl.>d. from Sara Oldfield or from the lOS or the AlAS

TO Til E COLLECTOH I~' TilE FIELD


Before youcolil.·ctanything;
• DO acquaint yourself with CITES and nalional and stale controls, and find
OU! which species are protected
• 00 obtain all ne<:cssa ry pcnnits, both for collecting and for export and
importtoothercounlries.
• DO notify in tcrcsted IOC.ll org~nizationsofyourintentions.
Then:
• 00 strictly observe restrictions on what may be collected (which speciL'S, how
milny specimens, what kind of material). Where possible. culll'Ct seeds, offsets
ur cuttings, nut the whule plant.
• DO leave mature plants for ~ production. They are nt.'eliL>d to perpetuate
the wild populatiun, and are unlikely to transplant successfully.
• DO collect discreetly; do not lead local people to believe that the plants are
valuable or encourage or pay them (or their ch ildren) to collL>ct for you
• DO make careful field notes, including precise locality, altitude. type of vege-
tation and soil, da te of collection and your own field number. Try to assess
the number of individuals and extent of the populatiun, the amuunt uf seed
settingandthefrequt.'flcyofseed1ings.
• DO note possible threats to the habitat - e.g., through graling, drainage or
cultivation, urban spread ur road widening.
• DO take photographs and/or preserve representative lu::rbarium material.
Submit tlus material, with a copy of your notes, to an appmpriiltc ins titution
ororganilation.
• DO NOT underrate the value of your field observations: carefully rt.'COrded
they will be a usefu! contribution to science and to conservation.

230
if;
• you plan to colle<:t in commercial quantitit'S, don't.
• you plan to sell any of the plant~ you collect to defray the cost of your trip,
don'l.
• you plan to collect for r~arch or study, obtain the agreement (and prefer-
ably thc collabor.ltion) of competent scientific authoritit'S, ~uch as a govern-
ment agency or university dep.1rtment, in the host country.
• you think 'two or three plants won', be mis.wd', rcmember someone else may
be thinking the Silme tomorrow, and the next day, and the next ...

TO T il E 1l' II' OHTEn. I'BI VtHE on C:O I\ l ,\ l i-:UC IAL


• OON'T import wild plants, even if It>gillly permitted, except as iI nucleus for
prop.lg<'ltion ,lnd seed-production.
Then:
• DO check the credentials of suppliers offering wild plants and satisfy yourself
IhattheYiIT~'k-gal'.

• DO observe international and n<ltional export/import rcguhltions.

• 00 sell nursery-raised or prop.lgated ma terial only; do nut advertise ur sell


un-prop.lgated wild plants under any circumst.lnccs, even when legally per-
millcdlodoso.
• 00 try to propagate all rare or documented ma terial and distribute it to TCC-
ogni7oo lOS Reference Collections.
• 00 keep more than one done of rate species, even self-fertile ones, for seed
production.
• 00 kl'Cp careful re<:ords of the origin of all stock, especially any wild culkoc-
tors' number or locality data, and pass on the information to interested pur-
chasers.

TO TilE C IWWEU OU COtLECTO H AT 1I 0 !\1E


• 00 make successful cultivati on your prime objective, not the size of your col-
It'Ction or the rarityofi he plants.
• 00 NOT buy any plant unless you arc surc it was nursery grown; remember
that your choice will influence the seller's market.
• 00 not buy wild-collected plants, even if with the ilim of Silving the 'individ-
ual'. We want to save Ihe species, not the specimen. Only when importers see
their wild-collccted plants rotting because nobody buys them will they stop
the import.llion of wild-collected plimts.
• 00 enjoy the S.1lisfaction of raising (rom s~. Some o( the rate or 'difficult'
species will t(OSt your skill and patience, but reward your success accordingly!
• 00 record when and from whom you got your plants or seeds and ask your
source for any dat.1; colle<:tors' numbers. locality, and so on. All these are just
asvitaltotheseriousenthusjastasthenameoflhel~bc1.

231
• 00 try to propagate r.lre and documented material and distribute it to other
l'llthusiasts. Remember the proverb: To keep iI plant, give it away!

TO T III~ SOC IET Y Ar-..D CL UB


• 00 endorse the pn:<:cpts of this Code as a guide for responsible a.nd COlooen-
tiousbchaviour.
• 00 NOT permit wild plants to be advertised for sale in your publications,
eithL'I" openly or by hints.
• DO publicile nationa.l and international regulations on the export, im"ort~­
tion ilnd sale of wild plilnt~.
• DO sponsor or support national and international measures 10 protect the
habitats of rare and threatenoo species.
• 00 infonn th(' competent authorities of any suspt.'Ct sale of collt.'Ctlod plants. If
you know of people tra\·elling to countries when! 5uccul('nts grow wild with
the intention to collt'(:ting. inform the competent authorities; the best way to
stop habitat exploitation by collectors is to catch th\'"m at the port of ~mtry
with the plants in theirho1Ods.

TO T il E 5 11 0 W COJ\ l j\ IITTEE A\I) JUI) (: r s


• 00 include in the schlodule some classes for plants raised from seed by the
exhibitor.
• 00 NOT )Jl'Tmit spedes prolecloo by CITES Appendix I to be shown in com-
petitive classes, except lIS bl't.odlings or other artifical1y raised propagations.
• 00 make a policy of giving preferenre to well-grown seedlings over field-
collected plants. ChL'Ck that obvious or suspcctoo 'imports' art' properly
rooted and established.

232
CHECKLIST OF ALTERNATIVE NAMES
The following list contains some of the synonyms in common use
This is a partial listi ng.. and several obsolete names th at ilTC seldom used have been omit ted

Gen us Species ... See Genus Species ... See

Abromciticlla pu/viM/a Abromeiliellachiormlilm Aloc abyssilliell A/oc e!lI


albo-duela AJOt's/riala
Ailellill (lIlgrlst/seeM Adelliadigitala barblldensis Aloevem
bue/l11/mumli! Adell/adigilll/ll «hinutll Aloehllmilis
1IlIIllif/om Adl'lriadigilala var.m;/Iillola
cl/ell!Jergai AiQfar;stalll
Adromischus eWlealus Adromiscllllscool'eri garmsanll Aloegarieprl1sis
festioll s Adromischusrooperi hlillburyllnu A/oestrin/II
Iwlesowt'/lsis Ailrolllisc/lIIscooperi Ilumilis Aioelllllllilis
v3r.cmldollei
P'()curvus Adromisc/lIIslrif/oru5
suocomllTessl/s Adromischu5lrif/oru5 indica Aloewra
5ubpelio/aris Adromisc/rllslriflorus III11Zl1!:' Aloe vera
lingua Alot'plicali/is
A!:'llIIirlm bcrlole/iaUlIIII Aermiumlabulaqormc lilrgllaeformis Alot'plimlili,
macroleplllII Aeonirmllabliloqorme IOllgiarislrlla Aloearislala
meyer/leimii Aeolliumglalldil/oslim maclilosu Aloeobscum
pal/iClilala Aloes/riala
Ag'~ Ullglislissima Agavegemilliflom per/oliala AllIell1lmilis
vaL/wlllilis
baker; Agawkanvill;;kii
piela Aloeobscura
cure/ll/fiOOOIlt/ra AgavexylOllacmlllrn
pralifem AllX'brevifolill
c/rihrmlrualla Agaveparryi
pUl/clala Aloevariegata
coelum Agawfrrox
rlrodocincta Aloes/ria/a
cOll5iderallli Agavevic/oriae.reginae
schmid/iaml Aloecooperi
coret/aray Agavekarwillskii
5empt'rviooidl"S Aloeparvllia
t/iscepta/a Agavesc/ridigera
tripe/lila Aluepliculilis
film/1el/losa Agauefilifera
lubcrcrllala Aloehumilis
glatICescl'lls AgaWlltlemUlla var. echirwla
m;lis Agawce/sii vulgaris Aloevem
\'ar. albiCUIIS
"iekel,ii AgaVt'vic/oriae-regillae
Amaryllis corallica Alllmoclwriscomllica
mgra Agavecolimalla
/IIsm/ls Agat'(.colimUlm
Allaculllpseros <lraclrlloides Allacampserosrufesc/!Ils
IMlolli AgaUt'J'Urryi
illtermedia AlIllcampserosfilame/llosa
Sillllldersii Agavepctatorlllll
1!Ot'IIIilzialla A'IIlCllmpst'Toslome/ltosa
scolymllS Agalle l'otatofUm
lodaroi Agavemarmomla
Allredem wnlifolia Boussillgaliltiacordifolia
<'fSI;la Agaveschitligera
wisliulli Axaveparrasalra
AlllegibbaeulIl fiS50idl"5 Gib/Nlerlm[i,scides

233
SU CCUL ENT S- TH E IL LUSTRAT ED DI CT IO NA RY

Genus S pecies ... See Genu s Sp ecies ...See

Argyro:Nfcrmll margaret/llle u.pidarinmargllrelhllc CraSSlila alooides Crassuill/wmispllaericn


all/hufUS Crassu/lll1t'rfOflllll
Beaucarnea recurwla Nolillam:urvalll arc/lai CraSSlIllIl'Ymmidlllis
eOlljmlcla Crassu/irperfofilia
Bryopllyl/um verticil/atum KalallcJlOetubif/orn cotyledoll Crassu/aaroorescells
cylindriC/! Crllssullli1Jlramidiliis
Burst'1ll vdomla Bursemfagaroides jlawuirells Crassu labreuifolia
\lar.e/ollga/a g/ilbrifolia Crllssu/atolllelrlOSiI
Iycapodioides Crassu/amuscosa
Cillibil//us caespilosum Cillibil/U/shookerii IIcolerllla Crnssulaperfomta
pearsollii Cros5ulabrevifolio
urollrlma CD/ldata OrlJeopsisC/!udata perfilata Crnssulaperforota
eodolloides urallulI/aspeciOS/l pelersollinc Cmssrlla pcrfomta
comll/utala CamllulI/alrersperidum teres CraSS/llllbarklyi
subsp.ilesperidulI/
corrugata CaralllllllaSO(:otrana DecabelOIlC hark/y; Tavuresiabarklyi
dicapuac ural/lIl1/atllmeri c/cgalls Tavuresiaallgo/clIsc
subsp.lumcri
data Caralllllllapriogollium
Didierl!ll adseem/ells Alluaudiaadscelrdelrs
mall/illaris Quaqrlamalllillaris
pmegracilis Orbropsiseaudata
Dudlcya lllilralli/ Oudlcyaalbif/ora
uml/ulI/aSO(:otmmr
lessellala Notalridllopsislessellata
Eehl!'Wria discolor £cheuerill/lodu/osa
glmlea Ec/revefUlprmrila
CephillophyllulII e/avifolium Jordalliellac/avifolia var.glauea
obscum Echevcrillagavoides
Ceropesia debilis Cerol'l'gia/iIlMris sangrlsla Ecirl'Ut'riasllbrigida
subsp.debilis
Ilians Ceropegillilic/rotoma
Eciridnopsis clrrysuntlra Eeiridllopsissculdlala
SUbsp./llallif/ara
Cheiridopsis Irill/illri A/oillol1sishill/1ari
soma/e/lsis Ec/ridllopsisdammal1ial1a

C/lidosco/ljs /atrophlll1rens Echillol/rall/nlls pech/le/ii Adeniapecirm!iii

COllophy/ll11r ,,,aesecllllll Opirli11l/moplryllulII


EUl'llorbUl all/allkara Euphorbia pac/rypotiioidcs
lI'aeseclum
caespitosa Errplrorbiaferor
ea/derel/sis £uphorbiacopiapilla
Colyledoll brrchlw/zUllla Ty/ecodonbuchlro/zUlmls
frllctus·pinii Ellpirorbiacaplll·medllsue
data Cotyledollorbicrrlala
glomeralll £uphorbia globosa
fascicrr/aTis Tylecodonpalliculalr15
gynophora Euplrortriaespinosa
leuteosqllamala Tylecrx/OIrpt"arsonii
/remrdill/Ill Ellplrorbiatrigona
mol/is TylecodOlrpalliculatus
lrul/Dlllle EllphorbUli/lcrmis
ab/onga CotyJedDllorbielllala va r.lllIl/ollat'
parriclilaia Tylecodo/lpalliculalus
wamll/illosa Eup/rorhiasqllarrosa
pearsonii Tylecodonpearsonii medrlSile Ellpllorbiaeapll/.wedrISile
pygmea TylecodOlrf1!lgmacus mDqlladare/lsi5 EUI'lrorbiaresillifera
ramosa CotyJedollorbiclllala lIIorilli! Errp/rorbialleplagOlla
reliw/ala Tylecodoureliwlalus rlripsaloides Ellpllorhialirllca/Ii
schaeferimm Ty/cco</o,rschl'e[erialllis SIlIl-SIllvudor Eupilorbiaresimft'rll
lardif/ora TylecodollpaniClilatllS splelldens Ellphorbiamilii
wallichii Tylecodonloolliehii 5u/lIIl11bcllala Euplwr/IUleopiapillil

234
CHECK LI ST OF ALTERNATIVE NA MES

Genus Sp e cies ... See G enu s Species .. .51.'1.'

It:Sselllllll Eupirorbiampul-mellusae L:.>debourm socialis Sdlla socialis


vimillulis Eupl1arbialimmlli
Utlwps bella Lilho/JSknrnsmonlallll
Gasleria ace/sa Gasleria/U5Caprlllclata subsp.bella
lIabueri Lilhopsllookeri
GibbaewlI /lcrviride Gibbtwumgibbosmu var.dablleri
delrdrilica Li/Jwpspseudolrrmcillelw
sill/Ildii GibbacuIII/mbesccll5 subsp.dClldriliell
var.s/llwriii
eli511bethiae Liliropspst'ridalrlllleilid/a
var.elisabel/riae
Hllworlhia IIc.I:'ola Haworlhialreroocca frilleyi Lillropsjuliisubsp·frilleri
eha/willii Haworlhiacoarc/alll iusu/ayis Lilho/JSbromfieldii
fal/ax HaworllriacOIIrclalrr var.inslilaris
frllUll HaworlhillCOIIrclal1l Il'Ticill'lmll LiliropskllrnsmOU/lmll
gIgas Haworliriallrncimoidea var.lerie/lrQ/1il
"'pida Hmvorllriacymbiformis lOCI/lis U/Jro/JSlerricolor
iI,teoro5ftl Hllworlllialrerooce margina/a Lilhops/lookeri
mllrgarilifern Hllworlhiapurni/II "ar.mllrgillllta
mllscu/illll I-/Ilworihillcoorclllill mllrtl,ae Lilhopsschwlmll!Sii
"ar.mllrlhlU!
pay"el Haworllrialrubacca
mcmre//i U/11O/JSbromficidii
pt'll/agOlla ASlrololxr/lelllagOlla VilT.lllcmreili
piclll Hliworlirill emelYlie rugOSll LiJhopsschw(l/Jlt:Sii
pilifem I/Ilworlhiacooperi var.TllgOSll
Illnlll/olin Hmoorl/riacymbiformis Lillrops/lookeri
5t'lnlll Haworllriaorochlloidea var.susmlllnc
tmllsll/cells Whops/rerrt'i
I-Ioyo partmlii Hoyabelfll uau~jlii OiulemllllmsVQuzjlii
wllleri Willlpslesliei
Hllemia tllt'llre;;ii TIiVIlll'siaallgo/ellst' vaT.lIellleri

Idria colzwlllaris FOliquicrilico/lwlllaris Milropilylliwi dissililm COllOphyllumdissilllm


frlllllesii CUlloplryliumfromesii
IlIIilllria muririi Giblxrerllllllebrowllii pisifvrmc Mmrilariapisifvnne
I'TOXIIIIUS COllO/Jllyl/rmlprorimrIS
/lIl1lldirrlcria proxima Drnwphilllsproximlls
NllIllmtlws IlIckhoJfii Aloiuopsis/rlckhoffii
KnlnllcirOt' fllrillllcea KallllICIIOt'5Clll'igem IIInlilerbei A/oiuopsislllnlllerbei
grlllldiflom KnlllllchOt'JIIllrmomlo arpeuii Aloillopsisorpcllii
/nciuntn Kalnllchoeilllegrn sciloolleesii Aloirropsissc/u)(meesii

Kleillia nrc/,ai Serrcciourc/reri No/illn luberCillaln No/illl1recur1llllu


nrlieu/Illll SelleciourliclzilllllS
desflersii Srlllxiodesf/crsii Or~en lIIutabilis Siapelinmutabilis
flligeus Sellecio/lllg"lIs
grlllllii Seueciogft1ntii Qroslnchys knnbocllsis Qrostncl!ysmitllltus
III.'Yiifvlia Selleciok/eillin
//Cudula SClleciopcllduirls I'llcilyphylulII f'nehyphylulllilookeri
pic/icaulis SelleciopicliCilU/is
SIIgillnlll Sellecio5llgillllln Paehyveria schrideckeri Echrucrillxscheidcckeri
scmperuivrl Selleciost'lIIpervivu5
P/eiosl'ilos arc/lai Tallqllallaarcheri
spieulllS(l SmCcillSl'iculosllS
f'leiosl'iloscomp<!clus
slnpeliae{ormis Scrrcciostulli!lillr/ormis subsp. CllrlUs
235
SUCC UL ~N T S- "I'II ~ I L L USTR A T ED OI CT I ONA RV

Gen us Species ...See Gen us Species ...see

Pll'iaspiiascompacll/s uroillei SedlllllSllrtorimlllm


subsp.millor verlolii Sedllmmzopclaillm
pettl/lleulatus Pleiaspilasnelii woodi Sedumspallllllifoliul/J
prismalieus Tallquanaprismalica var.pllrpUrf'1II11
Iricalor Plrias/Jilos/lelii woodwardii $t>dllmaizOQII
Zi!lItaro-tashiroi SedllmlillCare
Pseudo/ilhops splwericl's PstUi/olitllOpsmigiurtinus
SempcroivulIl a/bUill SemllCnlivellllalba
Rlwdiola ScdmnrDSeulll aroemt'llst $t>lIIperoivumlec/omm
gal/dini SClllpenlivulI!
Rocht'll {alcala CraSSllia/a/cala grandifl°rl/III
ilalicum SelliperolVUl/lleCIOru/ll
Ruschia II/biflora Polymilaa/biflora var.italieul/J
disarticu/ata Eber/llnzilldisarticlliata SelliperoivumteclorJ/m
vilr.a/pilUulI
tlYlorum SemperviVrl7llCairnrculII
San::oeau/Oll bumlallllii Sarcocauloucrosslcaulc va r.rakareulII
lorrd Sarcocau/Olilrerrd Scmpervitlllmleclo"Ulr
var·slaucum
Schizobasopsis vo/ubilis Bowit'llvolubilis
Senecio Sellcciehaworlilii
Sedlllll aizoidcs SeduIIJaizotm
a/heulII Sedu/llil/bum Sphllllllallll1ll5 viridiflorlls Nycleran/hllsviridif/orlls
bait/cum SedllllJa/bllllJ
boloniellse Sedmll;:e;rllIIgu/are Slape/ia amblglla SlI1peliagralld;f1om
blm/atii SedllmdllsypilyllulII dUllllllai Caral/ullladummeri
alli/omlcum SeduIIJspatlllllifolir/1IJ europaea Curul/umal!uropaeu
var.purpureum f1avirostris Stopeliagralldif/era
clliare Sedumspurium lIIargarita SlapeliahirSllla
cougestllm SedwlIspl/rium revo/uia Tromolricilerevolula
crellalum SeduIIJspurium slel/aris Siape/iaas/erias
de/llicl/lall/m Sedl/lIJspl/r;um stellata Stapeliaasterias
glanulifemm Sedumdllsyphyllum uariega/a Orbcauariegata
glaucum Sedumdasyplryllum OrbcaverrucoSll
glallcl/m Scdulllliispallicum
hillebrand/Ii Sedumsexaugu/are Slu/I;lia hardyi Orbealltlmslwrdyi
ibcriel/m Sedumstolon;/erulII
/aziCilm Sedumspurilllll Tacitus bel/us Grap/oIJe/ttlumllCllulII
maximowiczlI Sedumaizool1
mile 5edum sexanguiare Teslw/inaria elephanlipes Dioscoreaelepllllntipes
rotundifolium Sedumanacampseros palliclilata Dioscoreasylualica
TU/esCeIlS Sedlllllllicaullse var.paniCli la/a
sedlforme 5edumnieaUIl5e
sexf/dum SedumhispaniCilm Titmwpsis luckhoffi; Aloinopsisiuckho/fii
shasltmse SedulII/aliceolatl/m seli/era A/oillopsisscll/cra
spirale Sedumsexangulare
slellopelalulII 5edumlanceolu/um Urbil1;a ugaooides Ecileveriaagavoides
var.sllbillp;llu/li
suhalpillum Sedlll11lmlceoiatllm We/wi/sclria baillesii We/witsc/liamirabilis
sublile Sedllmlineare
IlIrgidlllll 5ed1lmalbum Yucca gualemalellsis Yuccaeiepiralllipes

236
FU RTHER R EAD I NG
r. t:NEIIA L
Gilbt,ort M.G., 'A r~view of Caml/ulIIll R.Br. and its scgre-
Jacobsen, II., A }/audbook of Sl4ccult'll/ Plau/s (3 volumes),
g~tes', Bmdkya, vol. 8, pp. 1-32., 1990
Blnndford Press, Poole, UK. 1960 (reprinted 1978)
Whit~ A . and Sloane B.L., The Siapeliae. Abbey San Encino
Rnuh. Werner, Tile Wonderful World of SUCCl4/e'lts, Press, ras.ldena. California, USA, 1937
Smithsoni.m rress, Washing ton D.C., USA, 1984

Wijnstekers, W., Th.. EooIu/;OII of CITES, 5ecret.,uia l of the BII OME U ,\CF.,\F.
Conve ntio n on International Trade in Endangered Species
of \Vild Flora and Fauna, LaUs.lnne, SWit7.erland, 1992 (3rd Rauh W., Bromtliads for Home, Ganim and Grtt7lhouse,
edition) Blandfo rd Press, Poole. U K, 1979

Wille rt, 0.1. vo n, Eller, B.M., Werger, M.l.A., nrinckmann,


E. ,md Ihlenfeldt, H .D., Ufe Slratrgif'S of Succl/ lmls ill CUi\l I 'OSIT,\ t-;
Deserts, Clul\brid)l~ Univen;ity Press, C<lmbridge, UK, 1992 !·laHid<lY I'., 'No teworthy Species of Kldllia', Hooker's l colles
PllllrlarulII (Bentham-Moxon Trustees, Royal Botanic
Gardens, Kew, UK), vol. 39. part IV, 1988

Chahinian n.J., n~ Sallseflieria trifascialll Vilrirliel, Trans


Terra Public.ltions, Reseda, California, USA, 1986
Evans R.L , Hlmdbook of Cultit'ilted $cdums, Science Rcvicws
Gentr)' 11.5., AgilVt:S of Continental Northem Amcricll, The Ltd, Northwood, Middlesex. U K, 1983
Ari7.ona Uni versity Press, Tu cson, Ari7.0n.l, USA, 1982
Toclken 11.1{., 'Crassulaccae' . FIOri! of Sollthem Africa, vol.
Matuda c.
and Pina Lujan I., LAs PauillS Mex;cll1ws del 14 (ed. a.A. Leis tner), Bot,lnic Resca rch Institute, Pretoria,
Gellero Colcecion Miscclanea Est(ldo de Mexico,
YUCCIl,
South Africa, 191!5
Toluca, Ml'xico, 1980
W(l lthcr F_. Elhl!Vf!Tia, California Acadcmy of Sciences, $.ln
Ulrich B., 'On the d iscovery of Aglllif! schidigem Lemaire Francisco, California, USA, 1972
and stat us of ('('rlai n taxa of the section Xysmil81fVf' Berger',
Brit ish Carl us & Succl/ lm l/ournal, vol. 10, pp. 61-70, 1992
UIIJI En t: AU: ,\t-:

,\ S(:LEI'I,\ I)'\ C I~ ,\ E Choux P., 'les Didiercacees, Xerophytes de Madagascar'.


Mcmoircsdc/"Acadbnit Malgache, vol. 17, 1934
Bruyns r .v., 'Notes on Ceropcgias of the Cape Province',
8radleyn. voI.3,pp. 1-47, 1985 Rowley, G.D., 'Did iereaccae', Cacti of the Old World, British
Cactus & Succulcnl Society, 1992
Bruyns r .V., ·A revision of the genus Echidnopsis Hook. f.
(Asclepiadaceae)', Brndleyn, vol. 6, pp. 1-48, 1988
Et:I' II Olt lll ,\ CE,\ E
Dyer R.A., Ct'I'opegin, Bmchysldmo Ilnd RiOC7eUxin in
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Nethe rLands, l91!3 Berne, Switzerland, 1961

2.37
Carter S.. 'New Succulent Spiny Euphorbias from Eastern Van Jaar:s ....eld, E. 'The genus Gtlslf!rill: a synoptic review',
Africa', Hooker's fronts Pla/ltllrllm (Ben tham-Moxon Aloe Ooumal of the Succulents Society of South Africa),
Truste<'S, Royal Ootanic C.,rdcns, Kcw, UK), vo l. 39, part vol. 29,pp. 1·32 (special issue), 1992
111, 1982

Carter S. and Smith, A.R., 'Eu phorbiaceae' (pa rt 2), Flora of ,\1 ES D lilit " ,\ NTII EMACfo;,\ 1::
Tropiall Enst Africn (cd. R.M. Polhill). A.A. Balkcma,
Cole D., Lil1wps.: F/oWt'ril1g Stones. Acom Pres.s, Randb urg.
Rotterdam, The Netherlands, 1989
South Africa, 1988

Euphorow Journal, vots. 1-8, pUblishl>d by Strawberry I>ress,


Hammer 5., 'Conop/lyillm, an annotated checklist', Bmdlf'!/'l,
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pp. 53-84;S-z. vol. 9. pp. 105-28, 1988-9 1
White A., Dyer ItA. and Sioo ne B.L, n,e
Srl"ul(lI/
Euplwrbille (Suuthem Africa), Abbey Ga rden Press,
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P<lS<ldcna,Califomia, USA, 1941 Balkcma, Rotterda m, The Netherlands, 1973

FO UQ Ullm IM :I\ Ali


Humphrey R.R., TIll' &xJjurrr alld lis Home, The University Mnthew 3., Tire GeJWs Lewisill, Timber I'fl'tiS, Portland,
of Ari1.ona Press, Tucson, Ariwna, USA, 1974
USA,1989

GEII ,\ NI,\ CE,\E

Van der Walt, J.J.A., I'(/argolliums of Southem A/rica,


Purnell, Cape Town, South Africa, 1977

Van dec Walt, J.J.A. and Vorster, P.J., Pe/orgolliums of


Sou/hem AfriCfl (\'01. 2), JUla &. Co. Ltd. Kenwyn, South
Africa, 1981

Van der Walt, J.J.A. and Vorster P.)., Ptlargoniums of


Sou/lrern AfriCfl (vol. 3), National Botanic Gardcn..<;,
Ki rsten~h, South Africa, 1988

LlU ,\ CE,\E
Bayer M.B., 111( New Haworth!o Hlmdbook, Nationa l Botanic
Gardens of South Africa, Kirstenbosch, South Africa, 1982

Jeppe B., SoI/IIr tlfricillr AIOt's, Purnell, Cape Town, South


Africa,l%9

Pilbrom J., Howarthill IlIId Astroloba: A CO/l((IOr'5 Guilie,


Batsford, London, UK, 1983

Reynolds C.W .• The AIM of Tropical Africa arId Mmlagascor.


The Aloes Book Fund, Mb~b.'ne,SlVa7.iland, 1966

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Rotterdam, The Nethl;'rlands, 1982
CREDITS
Key: t lOp, c centre, b bottom, I left, r right.

All the photogr~phs w~re taken by I'ierfranco and Daniele Costanzo, with the
exception of: Enzo Bisso 33cr; AndrC'n Caltilbrign 120b!; CilTlo Doni l06bl; Luca
Magagnoli 26bt 2611, 3311, 45bl, 9Obr, 9611, 97cl, 194tr, 196d, 20211, 203bl, 20511 nod
227tr; Roberto Mangani 94br, IZler, 138tr, 140c! and 14311; Annilrosa Nicola 5Id,
55bl,67cr, 7Ocr, 70br, 7111, 71bl, 7ltr, 7ler, 7lbr, niT, ncr, 72br, 73tl, 73cl, 73bl.
73cr, 73br, 7411. 74bl. 74tr, 74hr, 75c1, 75bl, 75<:r, 7611, 76d , 76b!, 761r, 76cr, 7711.
77d, 77b1, 17IT, 77br, 7!:!tl, 78bl. ?SIr, 7&r, 78br, 79c1, 9011, 9Itr, 91cr, 132bl, 137tr,
137cr, 138bl, 139c1, 173cr, 176hr, 177b! and 177cr; Pasquale Ruocco 72bl, BOer,
85cr,97cT,119Ir,13&1,145d,165tr, 174cr, 182tr, 19Otrand221 tr;M(lu rizio5.1jcvi\
Ocr, 4511, 46bt 46tr, 47cl, 49br, 52br, 56tl, 58br, 5911, 8911, 89cr, 9411. l03bl, 106tl,
107bl, 107tr, 116c1, 124d, I26br, 129d 129bL 135tl. 143c1,I44bl, 196bl, 197cr,
199cr,2261l,226c1,226bl,227c1 and227b1.
'Ill.e following photogr<lphs were I"ken at the Succulent Collection of the
Municipal of Zurich: 31cr, 36br, 41cl, 4lb!, 4%1, 89br, 90cl, 9Obl, 91cl, 95br, 13lbl,
134bl, 13&r, 155cr, 173tl, 173d, 17Str, 179tr, ISSt!, 189bl, 189cr,194bl,200tr,201tl,
202d, 202tr, 203tr, 204cr, 227tl <lnd 227cr; the following photogr~phs were taken
at the Garden of the National Herbarium of Namibia, Windhoek: 46bl, 46tr, 49br
and I%bl; and the following photographs were taken at the Botanical Garden of
the University of Palermo: 33cr, 48tr, 59tt 89cr, 94tl, 135t1, 143c1 and 227b1

2"
Dr Muurizio Sajeva is based at the
Department of Botany at the University
of Palermo in italy and is president of the
Associazione Italiana Amatori Piante
Succulcntc (the Italian Cactus and
Succulent Society). He is the Italian CITES
Scientific Authority representative for
plants, a member of the International
Organization for Succulent Plants Study
(lOS) and other scientific societies, as wel l
tiS being respo nsiblu for thu Ruscue Centre
for Cacti lind Succulents at the Botan ical
Carden of Palermo.

His co·author is Mariangela Costan7.0,


Secretary of the Associazionc IUlliunn
Amntori Piunte Succulcnte. She is
pnrticulnrly inlerestt.>d in the stud y
of Alpine succulenLS. especially Sedllm,
Semperuiullm and related genera and she
contributes to several horticultural
magazines.

AllJ(lofi'lltrc~1

SUCi'::ule ntJl II: New Illu stra t ed Dictionllrl'


Mauri:.uo&tj"'·RRnd Mll rillnl,:"1,, COoIl.lln~.o
Rock Gard e n Pl ant.s: A Color E ncyclop"dill
1l1l1dauare :\hnco
Caeti: The Illustrated Dictionary
Ken & Kod P,..,~lon. :\t afham

Tirnbt:r I'rt!M_. lnc.


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