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Key points:
Producers, or autotrophs, make their own organic molecules. Consumers, or
heterotrophs, get organic molecules by eating other organisms.
Food webs consist of many interconnected food chains and are more realistic
representation of consumption relationships in ecosystems.
Introduction
Organisms of different species can interact in many ways. They can compete,
or they can be symbionts—longterm partners with a close association. Or, of
course, they can do what we so often see in nature programs: one of them can
eat the other—chomp! That is, they can form one of the links in a food chain.
In ecology, a food chain is a series of organisms that eat one another so that
energy and nutrients flow from one to the next. For example, if you had a
hamburger for lunch, you might be part of a food chain that looks like this:
grass \rightarrow→right arrow cow \rightarrow→right arrow human. But
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what if you had lettuce on your hamburger? In that case, you're also part of a
food chain that looks like this: lettuce \rightarrow→right arrow human.
In this article, we'll take a closer look at food chains and food webs to see
how they represent the flow of energy and nutrients through ecosystems.
Autotrophs are the foundation of every ecosystem on the planet. That may
sound dramatic, but it's no exaggeration! Autotrophs form the base of food
chains and food webs, and the energy they capture from light or chemicals
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sustains all the other organisms in the community. When we're talking about
their role in food chains, we can call autotrophs producers.
Food chains
Now, we can take a look at how energy and nutrients move through a
ecological community. Let's start by considering just a few who-eats-who
relationships by looking at a food chain.
At the base of the food chain lie the primary producers. The primary
producers are autotrophs and are most often photosynthetic organisms such
as plants, algae, or cyanobacteria.
The organisms that eat the primary producers are called primary consumers.
Primary consumers are usually herbivores, plant-eaters, though they may be
algae eaters or bacteria eaters.
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The organisms that eat the primary consumers are called secondary
consumers. Secondary consumers are generally meat-eaters—carnivores.
The organisms that eat the secondary consumers are called tertiary
consumers. These are carnivore-eating carnivores, like eagles or big fish.
We can see examples of these levels in the diagram below. The green algae
are primary producers that get eaten by mollusks—the primary consumers.
The mollusks then become lunch for the slimy sculpin fish, a secondary
consumer, which is itself eaten by a larger fish, the Chinook salmon—a
tertiary consumer.
Picture -1
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In this illustration, the bottom trophic level is green algae, which is the
primary producer. The primary consumers are mollusks, or snails. The
secondary consumers are small fish called slimy sculpin. The tertiary and
apex consumer is Chinook salmon
Each of the categories above is called a trophic level, and it reflects how
many transfers of energy and nutrients—how many consumption steps—
separate an organism from the food chain's original energy source, such as
light. As we’ll explore further below, assigning organisms to trophic levels
isn't always clear-cut. For instance, humans are omnivores that can eat both
plants and animals.
Decomposers
One other group of consumers deserves mention, although it does not always
appear in drawings of food chains. This group consists of decomposers,
organisms that break down dead organic material and wastes.
Fungi and bacteria are the key decomposers in many ecosystems; they use the
chemical energy in dead matter and wastes to fuel their metabolic processes.
Other decomposers are detritivores—detritus eaters or debris eaters. These
are usually multicellular animals such as earthworms, crabs, slugs, or
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vultures. They not only feed on dead organic matter but often fragment it as
well, making it more available for bacterial or fungal decomposers.
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Examples of decomposers: left, fungi growing on a log; right, an earthworm.
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Decomposers as a group play a critical role in keeping ecosystems healthy.
When they break down dead material and wastes, they release nutrients that
can be recycled and used as building blocks by primary producers.
Food webs
Food chains give us a clear-cut picture of who eats whom. However, some
problems come up when we try and use them to describe whole ecological
communities.
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The bottom level of the illustration shows primary producers, which include
diatoms, green algae, blue-green algae, flagellates, and rotifers. The next level
includes the primary consumers that eat primary producers. These include
calanoids, waterfleas, cyclopoids, rotifers and amphipods. The shrimp also eat
primary producers. Primary consumers are in turn eaten by secondary
consumers, which are typically small fish. The small fish are eaten by larger
fish, the tertiary consumers. The yellow perch, a secondary consumer, eats
small fish within its own trophic level. All fish are eaten by the sea lamprey.
Thus, the food web is complex with interwoven layers.
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In food webs, arrows point from an organism that is eaten to the organism
that eats it. As the food web above shows, some species can eat organisms
from more than one trophic level. For example, opossum shrimp eat both
primary producers and primary consumers.
Bonus question: This food web contains the food chain we saw earlier in the
article—green algae \rightarrow→right arrow mollusks \rightarrow→right
arrow slimy sculpin \rightarrow→right arrow salmon. Can you find it?
For example, in the meadow ecosystem shown below, there is a grazing food
web of plants and animals that provides inputs for a detrital food web of
bacteria, fungi, and detritovores. The detrital web is shown in simplified form
in the brown band across the bottom of the diagram. In reality, it would
consist of various species linked by specific feeding interactions—that is,
connected by arrows, as in the grazing food web aboveground. Detrital food
webs can contribute energy to grazing food webs, as when a robin eats an
earthworm.
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The bottom level of the illustration shows decomposers, which include fungi,
mold, earthworms, and bacteria in the soil. The next level above decomposers
shows the producers: plants. The level above the producers shows the primary
consumers that eat the producers. Some examples are squirrels, mice, seed-
eating birds, and beetles. Primary consumers are in turn eaten by secondary
consumers, such as robins, centipedes, spiders, and toads. The tertiary
consumers such as foxes, owls, and snakes eat secondary and primary
consumers. All of the consumers and producers eventually become
nourishment for the decomposers.
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would predict that the primary consumers store only 2,000 kcal/m^22start
superscript, 2, end superscript/year of energy in their own bodies, making
energy available to their predators—secondary consumers—at a lower rate.
This pattern of fractional transfer limits the length of food chains; after a
certain number of trophic levels—generally three to six, there is too little
energy flow to support a population at a higher level.
At each level, energy is lost directly as heat or in the form of waste and dead
matter that go to the decomposers. Eventually, the decomposers metabolize
the waste and dead matter, releasing their energy as heat also.
Image credit: modified from Ecological pyramid by CK-12 Foundation, CC BY-NC 3.0
Why does so much energy exit the food web between one trophic level and
the next? Here are a few of the main reasons for inefficient energy
transfer^{1,2}1,2start superscript, 1, comma, 2, end superscript:
Some of the organic molecules an organism eats cannot be digested and leave
the body as feces, poop, rather than being used.
Not all of the individual organisms in a trophic level will get eaten by
organisms in the next level up. Some instead die without being eaten.
The feces and uneaten, dead organisms become food for decomposers, who
metabolize them and convert their energy to heat through cellular respiration.
So, none of the energy actually disappears—it all winds up as heat in the end.
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