Professional Documents
Culture Documents
Guttation.
Submitted to:
Submitted by:
Irfan manzoor.
1. Transpiration 3
3. Significance of transpiration 6
4. Stomata 7
7. Guttation 11
8. Bibliography 12
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Transpiration
The process of transpiration in plants is the phenomenon of loss of water in the
form of vapour from its living tissues of aerial parts of the plant body. Basically
process of transpiration in plants is evaporation. There are various factors that
influence the rate of transpiration like environmental factors like light, CO2
concentration, temperature, humidity of air, wind velocity and internal factors like
root shoot ratio, leaf orientation, leaf area. The process usually occurs in the day.
It is the loss of water vapor from parts of plants (similar to sweating), especially in
leaves. Leaf surfaces are dotted with openings called, collectively, stomata, and in
most plants they are more numerous on the undersides of the foliage. The stoma
are bordered by guard cells that open and close the pore. Leaf transpiration occurs
through stomata, and can be thought of as a necessary "cost" associated with the
opening of the stomata to allow the diffusion of carbon dioxide gas from the air for
photosynthesis. Transpiration also cools plants and enables mass flow of mineral
nutrients and water from roots to shoots.
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photosynthesis in which the stomata are closed during the day and open at night
when transpiration will be lower.
Types of transpiration
Plants show three types of transpiration as follows:
In xerophytes the cuticle is comparatively thick and the extent of water transpired
is significantly less.
ii. Lenticular transpiration : Lenticels are small slit like aerating opening in the
bark of woody stems and fruits through which negligible amount of water vapour
is lost (0.1 % of total loss).
Lenticular transpiration
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Stomatal transpiration : The loss of water jn the form of vapours through
the stomata is called stomatal transpiration. The maximum amount (90 - 95%) of
absorbed water is transpired through stomatal transpiration. Stomatal transpiration
is commonly found in the leaves and stems of young plants.
The mesophyll is the ground tissue of the leaf enclosed on either side by the
epidermal layer. The leaf in general has loosely arranged spongy parenchyma cells.
The air spaces of parenchyma communicate with the outside atmosphere by means
of minute pores called stomata present in the epidermis. The mesophyll cells of
leaf get water from the xylem of the leaf vein by osmotic diffusion, and become
turgid and saturated with water. When sunlight falls on the leaf the water
evaporates from their moist walls into the intercellular spaces of mesophyll, which
become saturated with water vapour. This water vapour then diffuses through the
stomata into outer atmosphere which is unsaturated. More water then is taken into
the intercellular spaces from the xylem through the mesophyll cells. Thus, the
process of transpiration through stomata continues. If stomata are closed, the water
evaporated from the mesophyll cells saturate the entire intercellular spaces with
water vapour, which will diffuse out when stomata open.
Stomatal transpiration
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Significance of transpiration
• The process of transpiration in plants probably plays a vital role in absorption
of water and ascent of sap in tall plants. The passive absorption of water
through roots as well as ascent of sap is due to transpiration pull according to
Cohesion-tension theory of Dixon and Jolly.
• Water and mineral absorbed by plants from soil are transported upwards in
the transpiration stream through xylem ducts.
• The process of transpiration in plants prevents the heating of plant leaves
during excessive day temperature.
• The process of transpiration in plants is rightly known as a necessary evil. It
is evil as huge amount of water (about 97%) absorbed by plants is lost
through this process while it is necessary as it is helpful in many
physiological processes.
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Stomata
A typical stomata is a microscopic and usually consists of a slit like opening which
is surrounded by a pair of bean (kidney) shaped guard cells. The two guard cells
and stomatal pore constitute stomatal apparatus. A stomatal pore is generally
elliptical in surface view. The dimension of stomatal pore varies from species to
species, but on an average, it measures about 20 jtm long and about 10-20 |j.m
wide when fully open. Guard cells generally have thick and non-elastic walls
towards the pore and thin and elastic walls on opposite side. The guard cells are
living cells with cytoplasm and a central vacuole. The cytoplasm has a nucleus and
many chloroplasts. The guard cells are surrounded by large epidermal cells called
subsidiary cells. In turgid condition, stomata open and in flaccid condition they
close.
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Opening and closing of stomata
The small pores on the epidermis of leaves are called stomata. Each stoma or
stomata pore is surrounded by two guard cells. In dicot plants guard cells are
kidney shaped or bean shaped. In monocot stomata, guard cells are dumb bell
shaped .The inner wall of guard cells are thick and non-elastic. The outer wall is
thin and elastic.The adjoining cell walls of two guard cells around the pore are free
and not attached with each other and this help them to stretch laterally during
stomatal opening
The epidermal cells surrounding the guard cells are called subsidiary cells. The
stomata pore, guard cells and the subsidiary cells are together called stomata
apparatus. Each guard cell contains a single nucleus and numerous
chloroplasts.The guard cell chloroplast possess pigments of both photosystem I and
II and thus perform photophosphorylation.They lack RUBISCO enzyme and
NADP+ linked triose phosphate dehydrogenase enzymes.So carbon reduction
cycle is absent Starch is synthesized in guard cell chloroplast by sugars transported
from adjacent mesophyll cells and they are characterised by accumulation of starch
during night(in dark) and their degradation during the day(in light). Mesophyll
cells accumulate starch during the day and decrease during the night.This property
helps in the opening and closing of stomata.
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Mechanism of Opening and Closing of Stomata
There are two main theories about the mechanism of stomatal opening and closing
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Guttation
When leaves lose water as a liquid phase through special cells
called hydathodes it is referred to as guttation. The main cause of
guttation in plants is root pressure. During night when root
pressure is high sometimes then due to this pressure watery
drops ooze out with the assistance of hydathodes.
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Guttatation
Guttation can only take place when there is more water in the soil
than in the roots of the plant. The water potential in the roots has
to be lower than the water potential in the soil (the amount of
water in the roots is lower than the amount of water in the soil).
This contributes to root pressure. Root pressure supplies the
means by which guttation takes place. Root pressure occurs at
night when stomata are closed and transpiration has stopped or
during the day if transpiration is slow. If water potential is
balanced between roots and soil and there is no transpiration,
there will be no guttation.
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defoliation and fruit-drop, seemingly due to excessive and drastic
loss of nutrients in a short time.
Bibliography
1. Goatley, James L.; Lewis, Ralph W. (March 1966).
"Composition of Guttation Fluid from Rye, Wheat, and Barley
Seedlings". Plant Physiology .
3. Wikipedia
4. Google etc.
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Thank you
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