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BIRDS OF NORTH AMERICA
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ROCKS AND MINERALS
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SKYGUIDE
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FAMILIES OF BIRDS
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y
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Cy
mLKLK1 V. LLVI
Museum of Comporotive Zoology
Horvord University
CDU
LLKlF K. LLVI
Under the editorship of
HERBERT S. ZIM
Illustrated by
NICHOLAS STREKALOVSKY.
OLUEM FKEbb MEWTOKK
C l
jaws cl osed
HAIRY MYGALOMORPHS (Theraphosi dae) are com
monl y cal l ed tarantul as i n the U. S. Unfortunatel y, thi s
name i s shared with other spi ders. Hai ry Mygal omorphs
are known al so as Bi rd Spi ders, and they may occasi on
al l y catch nestl i ng bi rds, l i zards, or smal l snakes. I n S.
Africa, they are cal l ed Monkey Spi ders. Most are not
poi sonous to man. About 30 species occur in the U. S. ,
most l y i n the Southwest, none i n Europe. The l argest, from
the Amazon Basi n of S. A. , may be 6-9 em (3. 5") l ong,
with a 25 em (1 0") l eg span
Most Hai ry Mygal omorphs l i ve on the ground, but some
dwel l i n trees, others burrow. The eyes are cl osel y
grouped; these spi ders are sensitive to vi brati ons and
hunt at ni ght by touch. Cornered, the spi der may purr
or rear up on the back l egs. The "hai rs" on the abdomen,
easi l y shed or rubbed of by the l egs, are very i rri tati ng
to human ski n. The undersi de of each l eg ti p has a pad
of i ri descent hai rs. Young mal es l ook l i ke femal es, but
after the fnal mol t, emerge sl ender and i ri descent, pal ps
devel oped. Captive femal es have l i ved 20 years and
mol t after maturity; mal es , shorter l i ved , do not molt.
Members of a rel ated fami l y, Barychel i dae (not i l l us
trated), have a di ggi ng rake (p. 23) and make a trap
door to burrow entrance.
20
A. chalcodes
70 mm (2.7
11
)
Arizona
Aphonopelma eutylenum
southern Cal iforni a
These spiders show the diversity
among the Hairy Mygal omorphs
and also their common feature
of hairiness.
Cyrtopholis sp.
50 mm (2
11
)
Puerto Rico
21
TRAPDOOR SPI DERS (Ctenizidae) are mainly tropical,
but numerous species are found in the southern U.S. and
a few in southern Europe. All are about 1 -3 em (0. 3- 1 . 2")
long. Using the spiny rake on the margins- of their jaws,
trapdoor spiders dig tubelike burrows. The tube, includ
ing the opening, is lined completely with silk. To make
the trapdoor, the spider cuts around the rim of the open
ing, leaving one side attached for the hinge. The top
of the lid is camoufaged with debris, and additional silk
is added under the lid to make it ft tightly. The lid may
be held shut by the spider. When the spider feels the
vibration of passing prey, it rushes out, captures the prey,
and takes it down the tube. Except to capture prey, the
female seldom leaves her tube; males wander in search
of mates. Spiders in the small family Migidae (not illus
trated), mainly Australian and South African, are similar
but lack digging rakes.
CYCLOCOSMIA are found i n
southeastern U. S. and southeast
ern Chi na. The spider makes a
fal se bottom for its t ube wi th the
h ardened, squared-of end of its
abdomen and cl oses the top wi th
a si l ken l i d.
22 MYGALOMORPHS
MYRMEKIAPHILA of several spe
cies are found in southeastern
U.S. The bu rrow, often l ocated i n
an ant nest, has a si de branch
cl osed by a second door. The out
si de daor to the burrow i s cov
ered by a si l ken l i d.
BOTHRIOCYRTUM found in Cal i
forni a, i s the most commonl y col
lected trapdoor spi der. Ummidia
(not i l l ustrated) has its thi rd ti bi a
saddl e-shaped . Several Ummidia
are found in the southeastern
states, where they di g al most
horizontal tubes i nto banks. The
simi l ar Nemesia i s found i n south
ern Europe.
FOLDI NG-DOOR SPIDERS, Antro
dioetus, occur from the Gulf
coast to Al aska. Tube dwel lers,
they close their tubes by drawi ng
i n the ri m. Because the anal tu
bercle i s some di stance from the
spi nnerets and there are hard
ened pl ates (sclerites) on the back
of the abdomen, Antrodiaetus is
i n its own fami l y-Antrodi aeti dae.
Antrodiaetus burrow
MYGALOMORPHS 23
FUNNELWEB MYGALOMORPHS (Di pl uri dae) are easi l y
recogni zed by thei r l ong spi nnerets, whi ch may be more
than half the l ength of the abdomen. The spi ders are up
to 50 mm (2") i n si ze and most have onl y four spi nnerets.
Funnel web Mygal omorphs catch i nsects by entangl i ng
them i n a sheet of si l k. The spi der hi des i n a tube i n one
corner of the sheet. The tube may be among the roots
at the base of a tree or in crevices in rocks or wood.
Funnel web Mygal omorphs are mai nl y tropi cal , but about
ten speci es are found i n the U. S. and a few i n Spai n.
Atrax i s t he poi sonous Funnel web Spi der of Austral i a.
The N. A. Microhexura i s onl y 3 mm (0. 1 ") l ong. Because
it has six spi nnerets, Hexura is someti mes pl aced in a di f
ferent fami l y, the Meci cobothri i dae.
`
E uagrus
spi nnerets
Hexura f ulva
12 mm (0. 5
1
)
U.S. Pacifc coast
PURSEWEB SPIDERS (Atypidoe) ore about 1 0 to 30 mm
(0. 4- 1 . 1 ") long. The coxa of each polp is widened to form
on endite, and these, as in all true spiders, sere as
mouthparts. Sphodros, found from Kansas and Texas as
for north as Wisconsin and New England, digs a hole at
the bose of a tree and constructs a silken tube camoufaged
with debris. The spider stays hidden inside the tube, which
may extend 1 5 em (6") up the side of the tree. If on insect
lands on the tube, the spider bites through it with its huge
fangs and pulls in the insect. The remains ore thrown out
through the hole before it is patched up. In the norhern
states, moles ore found after June rains when they won
der in search of females. The European Atypus constructs
a small tube that resembles a half-buried root.
underside of Sphodros
0 European
Atypus catchi ng a fy
J
. ~ M
T R U E S PI DERS
w
Suborder labidognatha
cl osed
Most common spiders belong to this suborder, found
even in the Arctic. Their jaws (chelicerae) are attached
below the head and open sideways, sometimes obliquely.
With few exceptions, all have two lungs.
carapace
carapace
Nops sp.
' 6 mm (0.2
11
)
lesser Anti l les
26
anal tubercl e
31
Carapaces
Pholcus
Spermophora
Physocyclus
u
Leplonela
Ochyrocera
DADDY-LONG-LEGS SPI DERS ( Pholci
dae, not to be confused wi th Phal angi
i dae, p. 132) have unusual l y t hi n, l ong,
sl ender l egs wi th fl exi bl e ends . Most
speci es are whi ti sh or gray. A few have
si x eyes i n two groups of three. Others
have ei ght eyes, the front center pai r
s ma l l . The eyes ar e a l ways c l os e
together. Many of these spi ders hang
upsi de down i n a l oose web i n dark
corners of houses or cel l ars. Others li ve
under stones i n the dry areas of temper
ate and subtropi cal regi ons . Mal es and
femal es are commonl y found together.
Mal es have l arge, si mpl e pal ps; the
femal es l ack an epi gynum but have a
swol l en area on the undersi de of the
abdomen . The femal e carri es the round
egg sac i n her j aws . Of more than
300 speci es of Phol ci ds, about 40
speci es are found i n North Ameri ca
north of Mexi co; a few occur i n north
ern Europe and many in the Medi terra
nean regi on.
There are several other fami l i es of
rare, smal l (l-3 mm) ( 0. 05-0. l ) , l ong
l egged spi ders . These are mostl y cave
spi ders-the Leptoneti dae of the Medi
t er r a nea n r eg i on , Japa n , and ver y
rarel y Ameri ca; and t he Ochyrocerati
dae, whi ch i ncl udes about 20 tropi cal
and subtropi cal speci es found i n Amer
i ca, Afri ca, and Asi a . Carapaces of
representati ve genera.re at l eft .
32 DADDY-LONG-LEGS SPI DERS
LONG-BODI ED CELLAR SPI DER
hangi ng i n i ts web shakes sa
rapi dl y when al armed that both
spider and web bl ur and seem
to disappear. I t is cosmopol itan,
one of the commonest spiders
found in cel l ars.
Pholcus phalangioides
8 mm (0.3
11
)
cosmopol i tan; cel l ars
carryi ng eggs
.
Psilochorus sp.
4 mm (0.2
11
)
southern U. S.
SHORT-BODIED CELLAR SPI DER
Spermophora meridionalis
2 mm (0. 1
11
)
, eastern U.S. ; dark pl aces
Y
Pholcus face
DADDY:LONG-LEGS SPI DERS 33
UROCTEIDS (Urocteidae) of about a dozen s
p
ecies are
found only in the Old World. They live under stones and
in rock crevices, where they make a dense, fat silk tube
u
p
to 5 em [2) wide or a series of sheets above and
below the s
p
ider. Insects crossing the threads become
entangled and are s
p
un into the web as the s
p
ider, its
s
p
innerets
p
ointing in, runs around the insect. The
p
rey
is then cut loose and carried back to the center of the
retreat. The egg sac is also
p
laced between the layers
of silk in the retreat. large urocteids resemble oecobiids
(
p
. 1 15) in sha
p
e of head and s
p
innerets but lack the
transverse
p
late, or cribellum, in front of the s
p
innerets,
with which oecobiids s
p
in their distinctive webs.
carapace
face
34 UROCTEIDS
spi nnerets
HERSI LI I DS (Hersi l i i dae) form a fami l y
of 75 tropi cal and subtropi cal speci es.
They are 1 0- 1 8 mm (0. 4-0. 7") l ong,
wi th di sti nctivel y l ong spi nnerets . Hersi
l i i ds posi ti on themsel ves head-down on
bark or stone wal l s . When an i nsect
approaches , the spi der j umps over i t,
spreadi ng si l k and then rapi dl y ci rcl es,
spi nnerets toward the prey, fasteni ng i t
down . After the prey i s compl etel y
wrapped, i t i s bi tten and eaten. Mem
bers of onl y one genus, Tama, are found
i n southern Texas .
carapace
w
face
COBWEB WEAVERS, or Combfooted Spi ders (Theri di
idae), with more than 2,000 species, make up one of the
large fami lies of common spi ders. More than 230 species
occur i n N. A. north of Mexi co, fewer i n Europe; many
species are tropical and cosmopol i tan. The American
House Spider (p. 40) and the Wi dows (p. 42) are members
of this fami ly.
Cobweb Weaver s ar e us ua l l y sedent ary, hangi ng
upsi de-down i n the center of an irregul ar cobweb or hi di ng
in a crevice at the edge of the web. Some make a smal l
web beneath l eaves, stones, or l oose bark. The sticky
outsi de threads entangl e an i nsect that hi ts them and may
pul l i t into the web as they contract. Usi ng a tiny comb of
bri st
i
es (setae) at the end of the fourth l eg, the spi der
throws si l k over the capti ve, then bi tes and sucks i t dry .
. Cobweb Weavers have few or no teeth and do not chew
prey, as do spi ders of rel ated groups.
Most spi ders in thi s fami l y l ack strong setae (hai rs) on
l egs. Many have a spherical abdomen, al most al l have
ei ght eyes, and al l have three cl aws on each l eg (p. 1 3).
As i n other web spi nners, the mal e has poor vi si on and
courts the femal e by pl ucki ng threads of her web.
W
face
Teridion
carapace
THERIDI ON, with several hundred
species, is the second l argest
spider genus and incl udes many
of the commonest small spiders
from the Arctic to the tropics. All
hang upside down in an irregul ar
web. As in most spiders, identif
cation of species depends on fea
tures of the pal pi or epigynum, a
matter for the special ist.
. sisyphium
-, ( 4 mm (0.2
"
)
Europe; low p
COBWEB WEAVERS 37
Theridu/a emertoni
-
<
2 mm (
0
.
1
11)
eastern U.S.
under leaves
Spintharus lavidus
eastern U.S. Ia Bol ivia;
vegetation
38 COBWEB WEAVERS
Dipoena nigra
3 mm (0. 1
1
1
)
N.A.; ant feeder,
on ground
..
Co/eosoma Roridanum
2 mm (0. 1
11
)
cosmotropical ; l itter
ARGYRODES are ti ny, often si l
ver-col ored spi ders that l ive i n
t he webs of other spi ders. They
feed on thei r host's smal l prey or
join them in a l arge meal . Some
have an el ongated worm-shaped
abdomen and can wiggle its ti p.
Mal es have bumps (turrets) on
their heads. About 15 speci es are
found north of Mexico, several i n
the Mediterranean regi on, many
i n the tropics.
Thwaitesia afnis
5 mm (0.2
11
)
New World tropics;
vegetation
COBWEB WEAVERS 39
AMERI CAN HOUSE SPIDER
40 COBWEB WEAVERS
ACHAEARANEA is a very l arge,
world-wide genus recognized by
the patches and streaks often
present on the sides of the ab
domen. Some femal es have a
hump on the abdomen. Mal es are
smal l er than femal es.
TI DARREN female l ives i n a
curled leaf hangi ng i n the web,
and the tiny males hang nearby.
The mal e amputates one of hi s
enormous pal pi before hi s l ast
mol t. Two common speci es occur
from southern U.S. to S.A.
ENOPLOGNATHA are al l dark
colored (except E. ovalo) and
have a l eafike pattern on the
abdomen. Some l ive i n curl edup
l eaves, some under l ogs, others i n
l eaf l itter. Several species occur
in N. A. and Europe.
STEATODA usual l y are
brown with a white l i ne around
the front of the abdomen. S.
borealis sits i n a crevice near
the web. S. hes
p
era, of western
N.A., and S. bi
p
unctata, of
Europe, are si mi l ar.
$. borealis
.
c 8 mm (0.3") A
eastern N. A.; tree trunks,
bui l di ngs
S. erigoniformis
C 3 mm (0. 1
"
)
cosmotropi cal ; under stones
41
WIDOWS (Latrodectus) are the best known and largest
of the Cobweb Weavers. Several speci es are found i n
the U. S. , one i n southern Europe, and others i n the Near
East and S. A. Al l ar e poi sonous ( pp. 1 6- 1 7) . Femal es are
about 1 2- 1 6 mm (0.5-0.6") l ong; mal es much smal l er and
wi th l onger l egs. Adul t mal es wander i n search of femal es
but do not feed or bite; femal es rarel y l eave thei r web.
Strands of si l k are very strong.
BLACK WI DOW (L. moctons) is
found in worm sout heastern U. S.
and West I n di es as for North as
New York. I t is common in trash,
out houses, and du mps. The spi
der hongs in the web, wh ich is
under objects. I n North Ameri ca
many spi der bi tes ore from Bl ock
Widows. Si milar species ore found
i n t he western slates, Mexi co, and
other ports of the world. The
western North American species
is L. hesperus. Bth L. moctons
and L. hesperus hove on hour
gloss mark on the underside. The
abdomen of t he Molmignatte (L.
tredecimguttotus), the northern
Mediterranean species, is marked
with a seri es of red spots. The
young of al l species ore bri ghtl y
colored with stri pes and spots.
The egg soc is brown and popery.
. tredecimguttatus
BROWN WIDOW (L. geometricusJ
is cosmotropical , i ntroduced in
Fl orida. Usual ly they are brown
to gray, some bl ack; they are
found on ar near bui ldi ng
Though poi sonous it i s less l i kely
to bite or i njects less venom
than other Wi dows. The egg sac
is tufted, fufy.
RED WIDOW (L. bishopi) i s found _
onl y in pal mettos in the sandy
scrub- pi ne of central and south
ern Fl orida. The
white, smooth.
NORTHERN WIDOW (L. varia/us)
occurs from northern Fl ori da to
southern Canada; common in Brit
ish Col umbi a. General l y i t is
found in undi sturbed woods, in
stumps, or in stone wal l s. The
egg sac i s brown, paper-l i ke.
The "hourgl ass" i s disti nct but
usual l y broken.
L. moctans
and ather species
DWARF SPI DERS AND SHEET-WEB WEAVERS ( Li nyphi
i dae) form one of the l argest fami l i es of spi ders, wi th more
than 3, 500 speci es. They are among the l east known of al l
spi ders, however. Li ke other web spi nners, they have three
cl aws on each foot, and most speci es bui l d a domeshaped
or fl at web. Al l have l egs wi th strong setae, a row of teeth, .
on each si de of the fang groove of the j aws, and a col ul us
( p. 9) .
DWARF SPI DERS (subfami l y Mi cryphanti nae) of probably
several hundred speci es occur in N . A. north of Mexi co.
They are common in Europe, the Arct i c, and on hi gh
mountai ns; t he tropi cs are bel i eved to have fewer speci es.
Most Dwarf Spi ders are l ess t han 2 mm ( 0. 1 ") l ong, some
l ess than 1 mm (0. 05") l ong. Most make smal l sheet webs .
. The mal es of many Dwarf Spi ders have unusual turrets,
bul ges, or depressi ons i n the head regi on . The abdomen i s
usual l y spheri cal and, i n some, i t i s covered wi th a hard,
shi ny pl ate.
Some speci es are abundant i n l eaf l i tter and can be
col l ecfd in l arge numbers wi th a Tul l gren funnel ( p. 1 8) .
Many can be found under stones; others can be col l ected
by sweepi ng vegetati on with a net . As many as 11, 000
spi ders per acre (0. 4 hectare) have been found in eastern
U . S . , over 21/4 mi l l i on per acre (6 mi l l i on per hectare) i n a
grassy area in Engl and. These fi gures i ncl ude al l the di ffer
ent speci es of spi ders, but the Dwarf Spi ders make up two
t hi rds of the total .
T
he great number of i nsects consumed dai l y by such
l arge popul ati ons of l i ttl e predators i s di ffi cul t to esti mate.
How wi l l thi s natural food chai n be affected by the sprayi ng
of i nsecti ci des over the countrysi de? I t wi l l favor the i nsects,
because the spi ders l ay fewer eggs and each generati on
takes l onger to mature.
44 DWARF SPIDERS
face
side views of ( Dwarf Spider carapaces
Erigane web
50 mm (2
11
) di am. i n depression
SHEETWEB WEAVERS (subfami l y li nyphi i nae) are
mostl y l arger than Dwarf Spi ders (p. 44) and usual l y
have a pattern on t he abdomen. I n contrast t o Cobweb
Weavers, the abdomen is usual l y l onger than wi de, the
l egs may have strong setae, the jaws many teeth. The
mal e and femal e often hang upsi de down under the
same web and run rapi dl y when di sturbed. The webs,
whi ch have a few sti cky threads, are found between
branches of trees or bushes and i n high grass, often in
great abundance. I f an i nsect gets entangl ed, the spi der
bi tes from bel ow, pul l s the i nsect through the sheet and
wraps it up. The web al so protects the spi der from pred
ators from above, and someti mes a second web forms
protecti on from bel ow. Onl y a few of the many species
are widespread and abundant.
web 1 0 em (4
1
)
PLATFORM SPIDER
web 15 em (6
1
)
Frontine/lo pyromitelo
eastern and central
Canada and U. S.
47
^
-
da coccinea
Drapetisca soialis
Europe; others in N. A.
48 SHEETWEB WEAVERS
! 4 mm (0. 2")
so(h,a!rn U.S.; grass
Stemonyphantes blauveltae
a 4.5 mm (0.2")
N. America, debri s
Balthyphantes pallidus
< 3 mm (O. l ")
N. America;
i n webs in
ground l i tter
SHEETWEB WEAVERS 49
PIRATE SPIDERS (Mi meti dae) i nvade webs of other
spi ders. The sl ow-movi ng Pirate Spi der bi tes the web
owner, whi ch is qui ckl y paral yzed and sucked dry through
the l egs, one after another. Some si t wi th outstretched
l egs under l eaves wai ti ng for passi ng spi ders. One spe
ci es has been observed to pl uck the prey's web l i ke a
courti ng mal e to gai n entrance.
Pi rate Spi ders are recognized by the row of strong
curved setae on the front margi ns of the l ower segments
of the frst pai r of l egs. The eggs are left in a stal ked
sac suspended from a twig or rock. Thi s smal l fami l y i n
cl udes about a dozen speci es north of Mexi co.
50
Mimetus notius
6 mm (0. 2
11
)
southeastern U. S.
NESTI CI DS ( Nesti ci dae), often cal l ed Cave Spi ders, are
pal e deni zens of moi st caves and cel l ars. They make an
i rregul ar cobweb. The fourth l eg has a c omb on i t s l ast
segment ( p. 36). Between the front pai r of spi nnerets is a
col ul us ( p. 9) . Thi s structure, i ts functi on not known, i s
found al so i n some Cobweb Weavers, but al l of these ar e
dark spi ders . The Femal e Nesti ci d spi der carri es her egg
sac attached to her spi nnerets .
Thi rty-one speci es of Nesti ci ds occur north of Mexi co.
Rel ati ves i ncl ude the mi nute Symphytognathi dae ( not i l l us
trated) , the mi nute Archaei dae (Archaea, bel ow) of Afri ca,
and Mecysmaucheni dae of South Ameri ca .
Nesticus cellulanus
' 5 mm (0.2
11
)
N. Hemi sphere
Archaea has
distinctive carapace
and chel icerae. The
chel icerae are used to
spear spi ders. Mecysmauchenius
segmentatus
' 6 mm (0.211
)
Chi l e
NESTICIDS 51
ORB-WEAVERS (Aranei dae, often cal l ed Argi opi dae)
form a fami l y of some 3, 500 speci es found i n al l parts
of the worl d. About 180 speci es occur north of Mex
i co. Al most al l of these spi ders spi n an orb-web, an
engi neeri ng feat practi ced al so by the Ul obori ds ( p. 114)
and by the rel ated Ray Spi ders and Tetragnathi ds ( pp.
70-71 ) .
Orb-weavers have poor vi si on. They l ocate prey by
feel i ng the vi brati on and tensi on of t he threads i n t hei r
web, t hen qui ckl y turn the capti ve wi t h thei r l egs whi l e thei r
fourth l egs pul l out si l k and wrap t he vi ct i m. The prey
i s bi tten before bei ng carri ed to the center of the web
or to the spi der's retreat i n a corner, where i t i s eaten.
Anythi ng i nedi bl e i s cut out of t he web and dropped t o
t he ground.
In the fal l , femal e Orb-weavers of many speci es produce
egg sacs contai ni ng several hundred eggs (p. 14) , then
di e. I n some speci es the eggs hatch soon; i n others not unti l
the fol l owi ng spri ng. The l arge number of eggs produced
suggests that these spi derl i ngs face greater hazards than
do the young of scorpi ons, pseudoscorpi ons, and spi ders
cared for by thei r mothers. Orb-weaver spi der l i ngs make
a perfect orb-web, but as the spi derl i ngs mature, thei r
webs become more speci al i zed and characteri sti c of the
speci es.
Orb-webs are a favori te obj ect of research i n i nsti nc
ti ve behavi or. Strands are pul l ed back whi l e the spi der i s
worki ng to l earn how the spi der compensates for the
change. Or, as the spi der bui l ds a web i n a frame,
t he frame i s turned to determi ne t he i nfl uence of grav
i ty on the posi ti on of the web. Changes i n the web
bui l di ng pattern as the spi der matures are studi ed,
too. Smal l doses of some drugs gi ven to the spi der
on a f l y wi l l a l so c au s e t h e s pi de r to c h a n g e t h e
pattern of i ts web.
52 ORB-WEAVERS
carapace
SI LVER ARGIOPE
Argiope argenfafa
face
grasshopper caught i n web
is rapi dl y turned with frst
legs whi l e fourth l egs wrap
it i n si l k sheet drawn
from spi nnerets
Prey-handl i ng
By Typical
Orb-Weavers
face carapace
carries wrapped prey; wil l
hang it up or feed on it.
ORB-WEAVERS 53
ORB-WEBS are bui l t by many spe
cies at night. Fi rst a bri dge is
made. The spi der may wal k from
one bri dgehead ta another,
carryi ng along a l i ne, but usual l y
the spi der sits with its abdomen
i n the ai r and l ets the wi nd pul l
aut a si l k thread ( 1 ) . I f the
thread touches, a bridge is es
tabl i shed (2); if nat, it may be
pul led back and eaten.
When a bri dge is establ ished,
the spider may rei nforce it by
wal ki ng back and forth, l aying
down more si l k. Then the spider
drops on a thread ( 3) it has
fastened at the center of a strand
i n the bri dge. I t secures the verti
cal thread and returns to the
fork, the hub of the fnal web (4).
A radi us thread, attached at
the hub, is carried up to the
bri dge and across a short dis
lance (5) before it is tightened
and fastened (6). More radi i are
formed by the same procedure
54 ORB-WEAVERS
(7-9), and the hub may be
strengthened with additional
threads before t he spi der starts
a temporary spi ral ( 1 0) . Once
around, a leg touches the previ
ous turn, thus measuring the
distance between rounds. When
the temporary spiral is com
pl eted, the spider reverses di rec
lion, rolls up the ol d and puts
down new more numerous and
closer spaced spi ral s of sticky si l k
( 1 1 ) The compl ete spiral path
back to the hub is retraced ( 1 2) .
Most rebui l d t he radi i and spi
ral s each day ar ni ght; some re
move the web duri ng the day. A
central decoration characteristic
of the species may be added, or
a si l ken retreat i n rol l ed up
l eaves may be made at one si de
of the web, with a di rect l i ne to
the hub that transmits vi brations
of i nsects caught i n the web.
There are many vari ati ons from
this si mpl ifed descri ption.
ORB-WEAVERS 55
A. nordmanni
2 15 mm (0.6")
N. Hemisphere
56
ORB-WEAVERS
ARANEUS is the l argest genus of
spiders, with over 1 ,0 species
found in mast porta af the worl d.
Many speci es make a retreat In
a rol l ed-up leaf near the web.
Web vibrations are transmitted to
the retreat from a signal l ine at
tached to the hub.
SHAMROCK SPIDEl
A. trfllv
wldetpread
In N. A.; maGc
may be white
ORB-WEAVERS 57
Araneus cavaticus
eastern U. S. and Canada
FURROW SPI DER
Nuctenea cornufa
wi despread in
N. Hemisphere; on bui l di ngs
Nuctenea patagiata
' 1 1 mm (0.51)
N. Hemi sphere
58 ORB-WEAVERS
LA nCE SPI DER
Araneus thaddeus
' 8 mm (0.3")
eastern U.S
.
Araneus pegnia
east, U. S. , Mexi co
Y
< 6 mm
(
0.
2
1
)
ORB-WEAVERS 59
8 mm (0.3
"
)
. merianae
Europe
60 ORB-WEAVERS
Zygiella africa
8 mm (0.3"}
Europe, Brit. Col umbia
and eastern Canada to New Engl and
Zygiella x-notata
8 mm (0.31}
Europe, U.S. Pacifc
Atlantic coasts
ORB-WEAVERS 61
*
62 ORB-WEAVERS
Mangora gibberosa
eastern U.S., Canada; grass
CYCLOSA hang in the hub, hi d
den by debris i n a vertical l i ne.
Egg sac i s i n center of web. The
spider's abdomen extends be
yond spi nnerets.
C. conica
5 mm (0.2")
N. A. , Europe;
woodl ands
3 mm (0. 1 1)
#
C. turbinata
N. A.
ORB-WEAVERS 63
64 ORB-WEAVERS
Cyrtophora citricola
<
1
5 mm (0.6")
Mediterranean;
hos web l i ke Mecynogea
Aculepeira sp. 3
< 20 mm (0.8
"
)
western U.S., mountai n
meadows; A. ceropegia
widespread in Eurasia
NEPHI LA, found in southern U.S.
and i n the tropics, makes o huge
web, 1 m (39
11
) or more i n di om
eter. The strong webs, matted
and twisted, are used by South
Sea I sl anders for various kinds of
bags and fsh nets. Young Neph
ita make a compl ete web; adults
bui l d only the bottom portion,
l eavi ng the top irregul ar. Neph
i/a has conspicuous tufts of hai r
on the l egs. Femal es vary i n size.
65
BOLAS SPI DER
Mastophora bisaccata
southeastern U.S.
Bol as spi ders make
no web, but attract mal e moths
GASTERACANTHA, tropical spid
ers with a hard abdomen armed
with spi nes, hang i n mi ddl e of
web, adorned by white tufts
and often hi gh i n a tree.
by imitati ng the pheromone
(perfume) of femal e moths.
underside
ORB-WEAVERS
69
RAY SPI DER
Theridiosoma
gemmosum
3 mm (0. 1 1)
Eurasia, N.A.
shaded woods
RAY SPI DERS (Theri di osomati dae) form a smal l fami l y of
ti ny spi ders rel ated to the other Orb-weavers { p. 52) . The
smal l web, onl y 1 0 em (4") i n di ameter, l acks a hub but has
several radi i ti ed together near the center. The spi der hol ds
up the web i n the center by a ti ght thread so that i t forms
an umbrel l a. I f a fl y gets caught, the thread i s rel eased,
causi ng the web to spri ng back and entangl e the catch . The
spi der has a gl obul ar abdomen; the sternum i s short and
square behi nd. The egg sac i s suspended on a stal k. One
speci es i s wi despread; about 1 20 are known from the
tropi cs.
face
Tetragnatha sp.
70 ORB-WEAVERS
TETRAGNAT HI DS (Tetragnat hi dae)
make an orb-web, usual l y at an angl e
between verti cal and hori zontal . The
orb usual l y has 1 2 to 20 radi i and
wi del y s pac ed s pi ra l s . The s pi de r
hangs i n the center or cl i ngs to a stal k
somewhere near the web. Unl i ke other
Or b- weaver s , however, the fema l e
Tetragnathi ds, except Leucauge, l ack
an epi gynum { p. 9) . Fewer than two
dozen speci es of Tetragnat hi ds occur
north of Mexi co.
ORCHARD SPI DERS (Leucauge)
are common i n wooded oreas of
eastern U.S. The spi ders hang in
center of horizontal orb. About
1 70 species are tropical .
LONG-JAWED ORB-WEAVERS,
Tefragnafha, at rest may cl i ng
l engthwise al ong a twig or grass
bl ade, hol di ng on with the short
third pair of l egs. The long pai rs
of l egs are extended. More than
a dozen speci es are common i n
meadows near water throughout
N. A. and Europe. There are
more than 250 species i n al l
parts of the worl d.
Tetragnatha pal/escens
THI CKJAWED SPI DER
Pachygnatha sp.
6 mm (0.2
11
)
eastern U.S.
THI CK-AWED SPI DERS (Pachy.
gnafha) are found under debri s
or i n dense vegetation near wa
ter. Young make smal l orb-web
on ground; adults make no web.
eastern U.S. to Central America
ORB-WEAVERS 71
spinnerets
I
spinneret
face
Agelenopsis Tegenaria
FUNNEL WEAVERS (Agelenidae) are seen most easily in
late summer when morning dew makes their webs in lawns
conspicuous. The spider hides at the narrow end of a funnel
that spreads out across the grass. On feeling the vibration
of an insect crossing the web, the spider dashes out, bites
the insect,
ace
spi nnerets
GNAPHOSI DS ( Gnaphos i dae) ar e us ua l l y u n i f or ml y
bl ack, a few brown, some wi th marki ngs . The l ong abdo
men i s s l i ghtl y fl attened, and the front spi nnerets are cyl i n
dri cal and separated . The l egs have onl y two cl aws ( p.
13) . Al l are nocturnal hunters and i n dayti me are found
under stones or l oose bark, often i n a s i l ken sac . The egg
sac may be a shi ny pi nk or whi te papery di sc attached
ti ghtl y to the undersi de of a stone; i n some speci es i t i s a
whi te sac guarded by the femal e . Most Gnaphosi dae have
posteri or medi an eyes oval at an angl e (see above) and the
endi tes ( p. 9) concave and sl i ghtl y constri cted i n thei r
mi ddl e. Many of the 2, 000 speci es of Gnaphosi ds are
found i n temperate regi ons, probabl y 250 speci es i n N . A.
and fewer i n Europe.
Gnaphasa muscorum
N. Hemi sphere
86
Zelotes subterraneus
N. Hemi sphere
' 9 mm
(0.4
"
)
Drassoces lapicosus
' 14 mm (0.61)
Europe
GNAPHOSIDS 87
L
@
l
l
spi nnerets
SAC SPI DERS ( Ci ubi oni dae) resembl e Gnaphosi ds but
wi th l ess fl attened abdomen, l onger l egs, and wi th cl osel y
spaced, coni cal , front spi nnerets . Many are l i ght l y col
ored. They make a resti ng tube i n a rol l ed l eaf or under
bark or stones . More than 200 of about 1 , 500 speci es
occur i n N. A. Anyphaeni dae are s i mi l ar, but wi th s l i ts of
respi ratory tubes openi ng mi dway on the undersi de of the
abdomen. Mai nl y neotropi cal , thi s fami l y has 36 speci es
north of Mexi co.
photo Potzsch
Clubiona abbotii
5 mm
(0. 2
11
)
N. A.
ANT MI MI CS are abundant i n
t he genera Micorio and Coslion
eiro. They often l ive with the
ants they mi mic, but the advan
tage of mi micry to the spider i s
not understood. The abdomen
may be constricted or covered
with scal es; the gait i s antl i ke,
and the frst legs are held l i ke
antennae. Micaria, mai nl y a
north temperate genus, prefers
dry areas. Costianeira, mai nly of
New Worl d tropics, hos a
grooved thorax and usual l y i s
l arger and more bri ghtl y col ored
than Micorio. Micario is now
often consi dered a Gnaphosid.
Chelracanlhlum mlldel
juv. 1 0 mm (0.4")
Medi terranean; N. A. in
bui l di ngs. First l eg l onger
t han l ast; many speci es poi sonous
Phrurotimpus borealis
northeastern N. A.
l itter
SAC SPIDERS 89
Senocu/id
carapace
Prodidomid
carapace
face
Homalonychid
carapace
.
9 0
C L
O
face
SENOCULIDS (Senocul i dae) i n
cl ude only about two dozen species
i n the Ameri can tropics. They have
three claws and are rel ated to
Nursery Web Spi ders (p. 78) but dif
fer in arrangement of eyes. Senoc
ul i ds hunt on pl ants. The femal e
guards her egg sac.
PRODIDOMIDS (Prodi domi dae) i n
cl ude 60 species with two cl aws.
Related to Gnaphosids (p. 86) but
difer i n eye arrangement and i n
havi ng l ong, spread chel i cerae.
They are found under stones i n dry
areas in southern N. A. and i n
southern Europe.
HOMALONYCHIDS (Homal onychi
dae) are found only i n Mexi co and
i n southwestern U. S. The l egs of
these spi ders have two cl aws, may
be held strai ght.
WANDERI NG SPI DERS (Cteni dae) i ncl ude about 550 spe
ci es of 5-40 mm ( 0. 3- 1. 5") subtropi cal and tropi cal spi ders
wi th two or three l eg cl aws . Eye arrangement i s di agnosti c .
These spi ders someti mes travel as stowaways i n bananas .
lora ( p. 14) hunts duri ng the day on ground l i ke a wol f
spi der; others l i ve on fol i age.
9 1
GI ANT CRAB SPI DERS {Sparassi dae) are mostl y tropi cal
spi ders that hol d thei r two-cl awed l egs crabl i ke. larger
than Crab Spi ders (p. 94), they have teeth on thei r j aws .
About 850 speci es.
HUNTSMAN SPI DER (Heteropoda
venatoria), found around the
worl d i n tropical regi ons, is wel
comed i n houses because it eats
cockroaches. The spiders hide i n
crevices duri ng day and came
out i n eveni ng. The femal es carry
thei r egg sacs with the jaws. They
Huntsman Spi der,
or Banana Spi der
Heteropoda
venatoria
9 2
are found i n the southern U.S.
and are commonl y i mported with
bananas. A si mi l ar huntsman,
Olios, occurs i n the southwestern
states and i n southern Europe.
The front mi ddl e eyes of Olios
are as l arge or l arger than the
l ateral eyes.
face
PLATORI D CRAB SPI DERS ( Pi atori dae) of about a dozen
speci es occur i n Asi a and i n tropi cal Ameri ca. They are
characteri zed by fl atness, l ong mi ddl e spi nnerets, wi del y
spaced front spi nnerets, and two l eg cl aws .
w
face
carapace
,
^
.
/
/ ' /
:'! f
/
spi nnerets
SELENOPI D CRAB SPI DERS (Sel enopi dae) i ncl ude 200
speci es of l arge tropi cal spi ders easi l y recogni zed by thei r
fl atness and by thei r eye arrangement-si x i n a si ngl e row.
Li ke other crab spi ders, they have two l eg cl aws . Common
i n houses and under bark or rocks; i f di sturbed, they dash
si deways i nto crevi ces .
carapace
face
-
i l l
.
. .
_
M1sumena
carapace car0pace
Philoramus
CRAB SPI DERS {Thomi si dae, Phi l odromi dae) hol d thei r
l egs crabl i ke, out at the si des, and can wal k forward,
backward, or si deways . The l argest are tropi cal speci es
1 2-20 mm ( 0. 5-0. 8") i n body l ength . Many have horns or
ornaments on the head or abdomen, and some mi mi c bi rd
droppi ngs. Mal es are smal l er than femal es and have much
l onger l egs . Crab Spi ders wai t i n ambush for passi ng
i nsects; some hol d thei r front l egs outstretched i n readi
ness. Thei r vi si on f or movements i s good. Thei r j aws are
smal l , and after prey is bi tten, it is hel d above the spi der
and sucked dry. Those that si t on fl owers apparentl y have
a toxi n potent to bees, fl i es, and other i nsects much l arger
than themsel ves . They do not use si l k to capture prey, but
i n courtshi p, the mal e may wrap hi s prospecti ve mate
l oosel y i n s i l k. Femal es of most speci es guard the egg sac
but di e before the eggs hatch. Of about 2, 000 speci es,
more than 200 occur i n N. A. , fewer i n Europe.
FLOWER SPI DERS (Misumena,
Misumenops) sit on fowers, and
can change col or slowly.
Misumena vatia
10 mm (0.4
"
)
N. Hemi sphere
bi ti ng bee
Stephanopis sp.
6 mm (0. 2")
Chi l e
M. asperatus
5 mm (0.2
1
)
N.A.; vegetati on
CRAB SPIDERS 95
X. cristatus
< 7 mm (0.3
11
)
Europe
Tmarus angulatus
< 7 mm (0.3
11
)
N. A.
96 CRAB SPI DERS
X. emertoni
5 mm (0.2
11
)
N. A.
XYSTI CUS of more thon 230 spe
cies occur i n all parts of the
worl d, mainl y i n the Northern
Hemi sphere. Most are found
under bark ar on the ground.
Dul l brownish, they resembl e
thei r background. Mal es may l ook
quite di ferent from femal es.
CORI ARACHNE has several spe
cies occurri ng i n N. A. Extreme
fatness adapts them to hi di ng in
narrow crevices i n bark.
Tibe/lus oblongus
9 mm (0.4
"
)
N. Hemi sphere
PHI LODROMI D CRAB SPI DERS
TI BELLUS, l i ke l ong-jawed spi ders
( p. 7 1 ), stretch al ong twi gs and
grasses. They usual l y are col
l ected i n grass by sweepi ng with
a net. Mal es and femal es are
simil ar i n appearance#
THANATUS l ive on vegetation
and bork i n N. A. and Europe.
A captive T. formicin us had a
food preference for moths.
PHI LODROMUS are attive
Spi ders that cl i mb on
pl ants, or sometimes on
i n houses. Egg sacs are
to l eaves or bark. Many
more than 1 70 species
N. A. and i n Europe.
CRAB SPIDERS 97
face carapace
Sa!ticus
J UMPI NG SPI DERS ( Sal ti ci dae) are among our most
attracti ve spi ders . Most have bri ght col ors, often wi th
i ri descent scal es. A l arge fami l y of more than 5, 000 spe
ci es, J umpi ng Spi ders are most abundant i n the tropi cs,
but about 300 speci es occur north of Mexi co and many i n
Europe. Al l are smal l , most l ess than 1 5 mm ( 0. 7") l ong .
Jumpi ng Spi ders are active duri ng the day and l i ke
sunshi ne. They wal k wi th an i rregul ar gai t and l eap on
thei r prey, someti mes j umpi ng many ti mes thei r own
l ength . Most of the j umpi ng power is suppl i ed by the fourth
pai r of l egs, though they are onl y sl i ght l y modi fi ed for
j umpi ng . Before j umpi ng, the spi der secures a si l k thread
on whi ch i t can cl i mb back i n case i t mi sses i ts mark. At
ni ght or when it is cool , the spi ders stay in l i ttl e cocoons or
i n crevi ces .
Jumpi ng Spi ders' eyes are among the best i n i nverte
brate ani mal s . Two of the ei ght eyes are l arge. Many
speci es can recogni ze prey or other spi ders at a di stance
of 1 0-20 em (4-8"). They can al so change the color of thei r
eyes .
When a mal e J umpi ng Spi der fi nds a femal e, he stops,
waves hi s bri ghtl y col ored fi rst l egs, wags hi s abdomen ,
and hops . If the femal e is of the same speci es, she si gnal s
wi t h her l egs . The fi rst l egs ( and someti mes t hi rd) may be
col orful and enl arged. After mati ng, the femal e constructs
a s i l k cocoon for her eggs and guards i t .
98 J UMPING SPIDERS
ZEBRA SPI DER
Salticus scenicus
N. Hemi sphere;
bui l di ngs
6 mm (0.2
1
)
Pe/enes viridipes
5 mm (0.2
1
)
eastern U.S.
J UMPING SPIDERS 99
1 2 mm (0.5
"
)
100 J UMPING SPIDERS
P. clarus
N.A. ; common
on pl ants
1 02 J UMPI NG SPI DERS
PHI DI PPUS is common i n N.A.;
none i n Europe. Large, heavy
bodied and conspi cuous, th.ese
spi ders are found on vegetation,
stones, and someti mes i nside
houses. I n captivity, they are
active and have good appetites.
One captive Phidippus ate more
than 40 fruit fies in succession.
P. apacheanus
9 mm (0.
.11)
Rocky Mts.
P. johnsoni
displ ayi ng at
Rocky Mts. and westward;
stones
P. regius
! 1 8 mm (0.7'
1
)
southeastern U. S.
13 mm (0.5
11
)
JUMPI NG SPI DERS 1 03
Synemosyna formica
< 5 mm (0.2")
eastern U.S.
Synemosyna americana
< 6 mm (0. 2")
Central Ameri ca
J umpi ng Spi ders That Mi mi c Ants
carapace
Panama
Lyssomanes, because they
eyes i n four rows, are `"""
pri mitive j umpi ng spi ders.
1 05
CRI BELLATE SPI DERS have a cri bel l um, a fattened
sievel i ke pl ate i n front of the spi nnerets. Through i t the
cal ami strum, a row of curved setae on the next to l ast
segment of the fourth l eg, pul l s si l k. The hackl ed threads
produced are covered wi th fne wool that entangl es the
prey. Every struggl i ng movement of the prey onl y i n
creases the entangl ement.
I t was thought that fami l ies of cri bel l ate spi ders are a
separate, di sti nct group. Now the opi ni on is that the pres
ence of the cri bel l um i s an ancestral condi ti on and the
structure has been l ost several ti mes. For i nstance, the
Oecobi i dae (p. 1 1 5) are si mi l ar to the Uroctei dae ( p. 34)
though smal l er and owners of a cri bel l um. The cri bel l um
i s sti l l a good character f or i denti fi cati on.
Hypochifus
cal amistrum
L L
.
face
ACANT HOCTE NI DS ( Acant hocten i dae) r esembl e t he
Wanderi ng Spi ders ( p. 91 ) but have a cri bel l um. They have
two l eg cl aws. Thei r hackl ed webs are found near thei r
hi di ng pl aces under l oose bar k. About 25 speci es ar e
known f r om t he Ameri can tropi cs.
ZOROPSI DS (Zoropsi dae) have two or three l eg cl aws;
most cri bel l ates have three. They di ffer from Gnaphosi ds
( p. 86) by the cri bel l um and by scapul ae under the l ast two
leg segments . Two dozen speci es occur around the Medi
terranean , i n Mexi co, and i n Central Ameri ca .
OGRE-FACED SPIDERS (Dinopidae) usually have two
enormous posterior median eyes; the remaining six eyes
are small. During daytime they hide in shrubs, and at
night, with a few parallel hackled threads, the spider
makes a rectangular web supported by silk lines, which
it holds with its front legs. When an insect approaches,
the web is spread and thrown. Cosmotropical in distribu
tion, about 50 species occur in the American tropics,
one in southeastern United States. Menneus, of Australia,
has small
p
osterior median eyes.
OGRE-FACED SPI DER
Dinopis spinosa
< 20 mm (O. B")
southeastern U.S.
ULOBORI DS ( Ui obori dae) have a cri bel l um but make orb
webs, though the spi ders are not si mi l ar i n appearance to
Orb-weavers ( p. 52) . Ul obori ds l ack poi son gl ands. About
200 speci es occur i n al l parts of the worl d . Fi fteen speci es
are found i n N . A. , fewer i n Europe, but they are most
abundant i n the tropi cs .
1 1 4 CRIBELLATES
ULOBORUS orb is often horizon
tal . I nstead of sticky thread, as
i n the web of Orbweavers, the
spiral is made of hackl ed threads.
Species of U/oborus are found i n
the U.S. and i n Europe; some
species are soci al . Rel ated genera
are found i n the tropics.
U. g/omosus
' 7 mm (0.3")
N.A.
HYPTI OTES web is tri angul ar. The
spi der attaches itsel f to a twig
by si l k from its spi nnerets and
hol ds the web wi th i ts frst l egs.
The spi der i tsel f bri dges a gap i n
the threads. When an i nsect gets
caught, the spi der pul l s the web
taut, then l ets it go sl ack agai n.
Thi s further entangl es the prey.
The web is made by constructi ng
a bri dge, a vertical l i ne, and two
more radi i . Scafol di ng threads
run out from the center, and
hackl ed threads are put down
separately i n each sector. H.
cavatus i s found i n eastern U. S.,
southern Canada. H. paradoxus
i s commonest European species.
TRIANGLE SPI DER
H. cavatus
4 mm (0.211
)
N. A.
OECOBI I DS (Oecobi i dae) have a
cri bel l um and are ti ny; otherwise
t h e y r es e mb l e t h e Ol d Wor l d
Uroctei ds ( p. 34) . Oecobi i ds make
smal l flat webs over crevices i n walls
and on vari ous leaves. They feed
on ants, and some are social . They
have a l arge, hai ry anal tubercl e.
Most speci es are tropical and sub
tropi cal; a few i nhabit houses i n the
North.
CRI BELLATES 1 1 5
S P I DE R R E L ATI VE S
A number of groups (orders) are bel i eved to share a com
mon ancestry wi th spi ders ( pp. 6-7) . Al l have j aws (chel i
cerae) , pedi pal ps (someti mes pi ncer l i ke), and four pai rs of
wal ki ng l egs. They di ffer in adaptati ons of the appen
dages . I n some, the abdomen i s segmented and j oi ned
broadl y to cephal othorax.
WHI PSCORPI ONS (order Uropygi ; about 85 speci es)
occur i n some parts of southern U. S . , but are most abun
dant i n Central and South Ameri ca, Asi a, and t he East
I ndi es . They have no sti ng but can pi nch. Al l are nocturnal ;
they have poor vi si on, but are sensi ti ve t o vi brati ons.
VI NEGARONES, or Vi negoroons
(Thelyphonidae), ai m a vi negar
scented mist from a gl ond at the
base of the tail when disturbed.
The spray contai ns acetic acid
and a sol vent that attacks the
exoskeleton of i nsects. Other
tropical species smell of formic
VI NEGARONE
1 1 6
acid or of chl ori ne. Vi negarones
burrow i n sand or under l ogs and
carry their prey i nto the burrow.
Female carries 20-35 eggs i n a
membranous sac under her abdo
men. The young ri de on the
mother until thei r frst mol t, then
become i ndependent.
l argest whipscorpion
known; mal es and
femal es are si mi l ar
i n appearance
8 em (3
"
)
T rithyreus pentapeltis
6 mm (0.2
1
) ..
southern Cal iforni a
SCHI ZOMI DS (order Schi zomi da, someti mes cal l ed Schi
zopel t i di a) ore smal l , none more than 6 mm ( 0. 2") l ong.
They l ack eyes, and the short tai l has onl y three segments .
About 80 speci es l i ve i n the tropi cs and subtropi cs under
stones and i n l eaf l i tter.
TAI LL ESS WH I PSCORPI ONS, or Wh i ps pi der s ( or der
Ambl ypygi ) , have a wi de head and thorax, t o whi ch the
abdomen i s attached by a stal k. The fi rst pai r of l egs i s
l ong and whi pl i ke. Ambl ypygi ds hi de under bark or stones,
and i f the stone i s turned, scurry si deways to the other
si de. I n the U . S. , they ore found i n the warm South , where
they may come i nto houses . Others are found i n tropi cal
Ameri ca, Afri ca, and Asi a . Femal e carri es eggs and young
f or fou r t o si x days . About 70 speci es ore known i n two
fami l i es, Charonti dae and Tarantul i dae.
WI NDSCORPI ONS ( or der Sol i f ugae) ar e s omet i mes
cal l ed Sun Scorpi ons because most l i ve i n deserts or dr y
areas. From 1 t o 5 em (0. 3-2") l ong, most are yel l owi sh or
brown . They have enormous j aws, and thei r l egl i ke pedi
pal ps are used wi t h t he fi rst pai r of l egs as feel ers. They
wal k on onl y si x l egs, as do Whi pscorpi ons ( p. 1 1 6) and
Tai l l ess Whi pscorpi ons ( p. 1 1 7) , and run swi ftl y-"l i ke the
wi nd . " Sensi ng prey, they may stop suddenl y, then start
agai n. Some can cl i mb trees .
Wi ndscorpi ons are voraci ous feeders . Wi th thei r l arge
pi ncer l i ke j aws they can ki l l even smal l vertebrates and
may someti mes feed on smal l l i zards. They can bi te but do
not have poi son gl ands. I t i s bel i eved that some use thei r
eyes f or hunti ng, but most use touch onl y. The mal l et
shaped organs on the l ast pai r of l egs are probabl y sen
sory. At rest, Sol i fuges may burrow or si t under stones .
Most are nocturnal , but some are acti ve dur i ng the day.
Mal es are general l y smal l er and have l onger l egs than the
femal es. Femal es bury thei r eggs and may guard them.
Adul ts l i ve l ess than a year.
Wi ndscorpi ons of the Ol d Worl d bel ong to several fam
i l i es. They are abundant i n Afri ca and the Near East;
absent from Austral i a . Si x speci es are found i n southern
Europe. Al most 1 20 speci es occur i n N . A. north of Mexi co,
900 worl dwi de. They are found as far north as southwest
ern Canada but are most abundant i n Ari zona and the
Great Basi n area . One speci es occurs i n Fl ori da, the onl y
one known from eastern U. S . Wi ndscorpi ons of N. A.
bel ong t o two easi l y recogni zed fami l i es, but t he speci es
are di ffi cul t to i denti fy. I n the fami l y Ammotrechi dae, the
fi rst pai r of l egs have no cl aws, and the front edge of the
head i s rounded or poi nted . I n the fami l y Eremobati dae,
the fi rst pai r of l egs have one or two cl aws, and the front
edge of the head i s strai ght .
1 1 8 WI NDSCORPI ONS
mal l et
shaped
organ
underside of
wi ndscorpion
Ammotrechella stimpsoni
20 mm (0.8
11
)
Fl ori da, West I ndies
Sol pugi dae, eastern Africa
E. durangonus
28 mm ( 1 . 1
11
)
Texas to Cal ifornia,
northern Mexico
E. pal/ipes
26 mm ( 1
11
)
N. Dakota to Arizona
DSCORPIONS 1 1 9
PSEUDOSCORPI ONS (order Pseudoscorpi ones) are com
mon everywhere but are rarel y seen because of thei r
secretive habi ts. They are smal l , most of them l ess than
5 mm (0.2") l ong. Al most 200 speci es are known from
N.A. Most species l ive i n l eaf l i tter, moss, manure, under
l oose bark, or under stones. Many can be col l ected from
l itter with a Tul l gren funnel (p. 1 8) .
Pseudoscorpi ons can wal k backwards as wel l as for
ward. They do not have a long tai l and sti nger as do
scorpi ons, but most have poison gl ands i n thei r pi ncers.
The poison i s used sol el y to capture smal l i nsects, thei r
prey. Pseudoscorpi ons are not l arge enough to bi te. They
have si l k gl ands openi ng on thei r jaws, and use si l k to
make chambers for overwi nteri ng, mol ti ng, or broodi ng.
Many species l ack eyes and, judgi ng from the l ong hairs
on the pi ncers, their mai n sense is touch. Pseudoscorpi ons
someti mes attach themselves to fl ies or to beetl es; they
are carried as hi tchhi kers, not as parasites.
In courti ng behavi or, the mal e waves hi s pi ncers, vi
brates his abdomen, or taps hi s l egs. The femal e responds
and the two ani mal s, thei r pi ncers l ocked, pul l each other
back and forth. Eventual l y the mal e deposits on the
ground a stalked capsul e contai ni ng spermatozoa. As
the femal e i s pul l ed over the capsul e, she pi cks it up with
the l i ps of an openi ng on the undersi de of her abdomen.
The eggs, i n a l i ttl e sac, stay attached to the femal e' s ab
domen, and when the young hatch, they remai n i n the
sac and feed on a mi l kl i ke secretion from the mother's
ovaries. Usual l y there are fewer than to dozen young,
but there may be more than one brood a year.
After l eavi ng the egg sac, the young ri de on the
mother for a short ti me. I t may be one to several years
before they become adults; duri ng this period, the young
mol t three ti mes.
1 20 PSEUDOSCORPI ONS
Lamprochernes minor
3 mm (0. 1 ")
and ' in courtshi p
Chitrella sp.
3 mm (0. 1 ")
N. A. ;
suborder Di pl osphyroni da,
al l tarsi divi ded
HOUSE PSEUDOSCORPION, one
of the l argest, is found i n houses
all over the worl d. Probably they
feed mai nl y on cl othes moths and
carpet beetl e l arvae, book-lice,
and other smal l i nsects and mites;
they are reported to l i ke bed
bugs. In search of moisture they
often become stranded i n si nks or
bath tubs, unabl e to cl i mb out
over the smooth surface.
Apochthonius sp.
2 mm (0. 1
11
)
N.A. ;
suborder
Heterosphyroni da,
tarsi of l ast 2 l egs
divided
cancroides
4 mm (0.2") cosmopol itan;
suborder Monosphyronida,
none of the tarsi divided
PSEUDOSCORPIONS 1 21
SCORPIONS (order Scorpi ones) have pi ncers and a
l ong tai l wi th a sti nger at its ti p. The pecti nes, a pai r of
combl i ke structures underneath the l ast l egs, are sense
organs of touch. Scorpi ons have two eyes i n t he center
of the head and usual l y two to fve al ong the margi n on
each si de. They do not see wel l , and depend on touch,
usi ng the l ong setae on thei r pi ncers. When runni ng,
they hol d thei r pi ncers outstretched. Mal es have broader
pi ncers and l onger tai l s than do femal es.
Most scorpi ons l i ve i n warm, dry cl i mates. But one
speci es i s found as far north as Al berta; i n Europe, Eu
scor
p
ius germanus i s found to al titudes of 1 ,800 m (6,000
ft.) in the southern Al ps; others are found near snow i n
the southern Andes. Speci es are restricted t o smal l areas,
and only one i s found worl d-wi de i n tropi cs. Scorpi ons
feed at ni ght on i nsects and spiders that are caught wi th
the pi ncers and someti mes stung. Scorpi ons that remai n
under stones or bark duri ng the day carry thei r tai l s to
one si de; burrowers hol d thei r tai l s up.
I n courtshi p the mal e hol ds the femal e by the pi ncers
or jaws and l eads her back and forth. Eventual l y he de
posits a stal ked package of spermatozoa (a spermato
phore) and pul l s the femal e over i t. She picks up part
of the capsul e wi an orgn on her abdomen. The young
' . DI PLOCENTRI DAE contai ns tropi-
cal scorpi ons from the West I n-
e
di es and southwestern N. A. The
' -
l arge Nebo is found in eastern
_. .. ,. ,
s
p
srnum
ri de on the mother's back unt i l they shed thei r ski ns for the
fi rst ti me, then become i ndependent and l i ve a sol i tary l i fe
that may l ast several years .
Scorpi ons sti ng i n sel f-defense. Most
s
ti ngs are not seri
ous , but dangerous scorpi ons occur i n Nor t h Afr i ca, S. A. ,
and Mexi co. I n the U . S . , Centruroides scul
p
turatus of Ar i
zona has poi son t hat affects t he nerves, causi ng severe
pai n . Speci es of Centruroides i n Mexi co have caused
deaths . Scorpi on anti veni ns are avai l abl e.
Of over 1 , 200 speci es known, 20 t o 30 occur i n t he U . S .
Scorpi ons are best col l ected a t ni ght when they are acti ve,
wi th a bl ack l i ght , whi ch makes them fl uorescent i n the
darkness .
SCORPI ONI DAE contai ns Ol d
Worl d and Austral i an scorpi ons.
Some are gi ants. Onl y one, Opis
thocanthus, occurs i n the West
I ndi es and Central Ameri ca. The
last two l eg segments have onl y
one spur on the outsi de, as i n
Di pl ocentridae, but there is no
tubercl e u nder t he sti nger.
Pandinus sp.
to 17 em (7
"
)
Africa; rocky areas
SCORPIONS 1 23
CHACTI DAE, represented here by
Euscorpius, i s a fami l y world-wide
i n di stri buti on. The sternum i s
al most square or wi der than l ong.
There may be either two spurs
between the last to l eg seg
ments or one-and if one, i t is
on the i nsi de. They moy have two
eyes on each side of the head or
none. Superstitiona, a three
striped scorpi on about 4 em
( 1 .611) l ong, i s found in south
western U.S.
Euscorpius carpathicus
S cm (2
11
)
southern Europe, very common
Sothriurus banariensis
5.5 em (2.2
11
)
southern S.A.
A
BOTHRI URI DAE contai ns mostly
South American scorpi ons, a few
from Austral i a. The ster num con
sists of two transverse pl ates,
much wi der than l ong, someti mes
barel y vi si bl e.
VAEJOVI DAE scorpi ons have a
broad sternum, l i ke Chactids, but
have three to fve eyes on each
side of the head. Some species
occur i n the Old World, and
many are American. .
Northern Voejovis
Paruroctonus boreus
5 em (2")
western U. S. to Albrta
VAEJOVI D SCORPI ONS
MORDANT UROCTONUS
Uroctonus mordax
6 em (2.4
1
)
Oregon, Cal iforni a
STRI PEDTAI LED VAEJOVI S
V. spinigerus
5 to 8 em (2-3")
southwestern U.S.
SWOLLEN-STINGER ANUROCTONUS
Anuroctonus p#aeodactylus
6 em (2.41)
Utah to Cal ifornia
YELLOW VAEJOVIS
V. faus
4 Cm (1 .5")
suhwesern U.S.
GIANT HAIRY HADRURUS
SCORPIONS 1 25
underside of Buthus
EUROPEAN BUTHUS
B. occitanus
5 cm (2
1
1
)
southern Europe;
sti ng pai nful
1 26
BUTHI DAE, wi th 600 speci es, has
some representati ves on oi l conti
nents except Antarcti ca; i t i s the
l argest fami l y of scorpi ons. The
sternum i s tri angul ar, poi nted in
front and often l onger than i t is
wi de. Many speci es have a tubercl e
that i s l ocated under the sti nger.
The pi ncers have a graceful , sl en
der hand. Some speci es are poi
sonous to man-Tityus i n Brazi l ,
Androctonus i n North Afri ca, and
Centruroides i n Mexi co.
.
SPOTTED I SOMETRUS
lsometrus macu/atus
5-7 Cm (2-2.7
11
)
cosmotropical
:
HASETH'S CENTRUROI DES
C. hasethi
5 Cm (2
11
)
with young
Lesser Anti l l es
HENTZ'S CENTRUROI DES
C. he0tzi
Fl ori da; under bark
SCULPTURED CENTRUROI DES
Ce0truroides seu/pturatus
6-7 Cm (2.4-2.8
11
)
Ari zona; poi sonous
SCORPI ONS 1 27
HARVESTMEN (order Opi l i ones) are commonl y cal l ed
Daddy-l ong-l egs, but many have short l egs . They hol d
thei r short, round bodi es near the ground. I f caught by a
l eg, the l eg may break off. The head, thorax, and abdo
men are j oi ned broadl y, and i n the mi ddl e of the head are
two eyes, usual l y one on each si de of a hi gh turret . In many
speci es, spi nes, tubercl es, or bri stl es arm the head; scent
gl ands open al ong the edge of the carapace. Mal es are
smal l er than femal es and have l onger l egs. Unl i ke other
arachni ds, Harvestmen do not court before mati ng. Wi th
a l ong, stout ovi posi tor, the femal e deposi ts her eggs i n
the ground i n the fal l ; the young hatch i n the fol l owi ng
spri ng. Most speci es l i ve l ess than a year or two. Harvest
men feed mai nl y on l i vi ng i nsects, someti mes on dead
ani mal s or pl ant j ui ces. There are 200 speci es i n N . A. ,
4, 500 to 5, 000 worl dwi de.
MI TE HARVESTMEN (suborder Cyphophthal mi ) are al l
l ess than 3 mm ( 0. 1 ") l ong. Eyes, i f present, are far apart
and i ndi sti nct, and the scent gl ands are on cones . Unl i ke
most mi tes, they have a segmented abdomen. Mi te Har
vestmen i nhabi t humi d l eaf l i tter. Probabl y not rare, but
sel dom col l ected. There are five fami l i es. Fi ve or si x speci es
are recorded for the U . S. Captives l i ve four to si x years .
Siro duricorius
Austria, northern
Yugoslavia;
forest l itter
LANIATORES (suborder) i ncl ude about 2,000 species,
mostl y tropical , but with a few representatives i n south
ern and western U. S. and i n Europe. lani atores have
the basal segments (coxae) of the frst three pai rs of l egs
touchi ng at the mi dl i ne. The smal l fami l y Oncopodi dae
( not i l l ustrated), found onl y i n southeastern Asia, has
the body (except for the l ast segment) covered by a hard
pl ate. The Assami i dae ( not i l l ustrated) carry thei r sl ender
pedi pal pi crossed over. I ncl uded are 300 species from
southeastern Asia, Austral i a, and Africa.
Stygnomma spinilera
4 mm (0.2
1
)
Fl ori da
PHALANGODI DAE hove two
cl aws, vi si bl e with hi gh-powered
hand lens or miscroscope, on the
fourth l eg. There are 20 to 30
species i n N. A.
TRIAENONYCHI DAE have a si n
gl e cl aw (wi th three teeth) on
the fourth l eg. Members of thi s
fami l y are found i n Madagascar,
Austral i a and the Americas, sev
eral north of Mexico.
HARVESTMEN 1 29
1 30 HARVESTMEN
COSMETI DAE ar e a l l Amer i can
speci es and most ar e tropi cal . Al l
have a smal l medi an t hi r d cl aw at
the t i p a/ the fourth l eg. The s mal l
pal pi have segments fl attened and
keel ed .
GONYLEPTIDAE is a fami l y of
about 600 species in the Ameri
can tropics. It i ncl udes the har
vestmen with the l argest fody.
All have a smal l medi an thi rd
cl aw at the ti p of the fourth l eg.
The l arge pal pi are not fattened.
Sadocus po/yacanthus
1 1 mm (0.5
1
)
Chi l e
PALPATORES (suborder) contai ns most temperate zone
Eurasi an and N.A. harvestmen. Of a total of about 900
speci es, 1 50 occur north of Mexi co. Basal leg segments
(coxae) separated by the breastpl ate (sternum).
Tragulus tricarinatus
1 0 mm
(
0.
4
11
)
Europe; u nder stones
NEMASTOMATIDAE have very
short jaws and no cl aws on thei r
pedi pal pi . The frst and fourth
basal l eg segments (coxae) have
margi nal rows of spi nes. The eyes
are on a tubercl e. About fve spe
ci es occur north of Mexico, 50 i n
the worl d.
Cros cus
d
asycnemus
1 . 5 mm ( 0. 06")
eastern Canada, U. S.
CERATOLASMATIDAE have
mi croscopi c scul pturi ng on
TROGULI DAE are sl ow and noctur
nal , wi th the eye tubercl e proj ecti ng
i n front and overhangi ng the j aws.
The European Trogulus, commonl y
encrusted wi th soi l parti cl es, i s a
predator on snai l s . T. tricarinafus
has been establ i shed i n upstate N. Y.
undersi de
undersi de
Nemastoma sp.
4 mm ( 0. 2")
Europe, eastern Canada
HARVESTMEN 1 3 1
l
|
1 32 HARVESTMEN ,
pal p
`
I SCHYROPSALIDI DAE usual l y have
huge j aws, l ack cl aws on the pedi
pal ps, and have shari l egs. About
30 speci es i n fami l y, 8 north of Mex
i co.
PHALANGI I DAE ( Daddy- l ong
l egs) are wi despread i n Europe and
temperate N. A. Each pedi pal p has
a smal l cl aw at i ts t i p, and i n al l
speci es, the l egs are l ong and sti l t
l i ke. Of about 800 speci es, 1 00 to
1 50 occur north of Mexi co. Caddi
dae are a rel ated fami l y wi th l arge
underside
northeastern U.S., Canada
HARVESTMEN 1 33
MI TES (order Acari ) are found throughout the worl d and
i n al l types of habi tats . They surpass al l other spi der l i ke
ani mal s i n numbers . Many are parasi tes on pl ants or ani
mal s, and some transmi t di sease. Others are freel i vi ng
predators . Some are aquati c-mostl y i n fresh water, a few
in the ocean . Among the parasi tes are some extraordi nari
l y speci al i zed speci es, adapted to l i ve onl y i n bi rd feathers
or nostr i l s , under bat wi ngs, i n monkey l ungs, bee tra
cheae, and si mi l arl y restri cted habi tats .
Most mi tes are very smal l , but they can be col l ected i n
abundance from soi l or l i tter by usi ng a Tul l gren funnel ( p.
1 8) . A mi te's body i s fused i nto one pi ece, wi th no separa
ti on between head and abdomen . In parasi tes, the mouth
parts may be speci al i zed as sharp, pi erci ng styl ets . Adul t
mi tes have four pai rs of l egs, but l arval mi tes have onl y
three. Some worml i ke, mi croscopi c gal l mi tes have fewer
l egs. More than 30, 000 speci es are named, probabl y a
fracti on of the total . Many have become pests s i nce the
wi despread use of pesti ci des .
MESOSTI GMATI D MI TES (suborder Mesosti gmata) have
ei ther a si mpl e cl aw on the pedi pal ps, or none. Many have
pl ates on the back and undersi de. Some are free- l i vi ng
predators and can be col l ected wi th a Tul l gren funnel .
Others are parasi tes and l i ve i n the fur of mammal s , on
bi rds, or on i nsects .
1 34
MI TES
Spinturnix sp.
1 mm (0.04")
laelaptid Mite
Laelaps vacua
1 .5 mm (0.06
"
)
on i nsects
TROMBI DI I D MI TES (suborder Trombi di formes) i ncl ude
many pl ant and ani mal parasi tes and some predators.
Water Mites (p. 1 36) are usual l y pl aced i n this group.
CHI GGERS, or Harvestmites
(Trombi cul i dae), form a family of
about 700 species. Adults are
predators on i nsects or i nsect eggs;
l arval stages are parasites. Fewer
than 50 speci es attack humans,
biti ng where clothi ng i s tight and
causi ng severe itchi ng. After
feedi ng, they fal l of. Some peo
pl e are i mmune. In the Ori ent,
chi gger mites carry scrub typhus.
I n the U. S. they are most abun
dant i n the South.
SPI DER MITES, or Red Spi ders
(Tetranychi dae), 0.3-0.8 mm (.01 -
03
11
), are seri ous pests on vari
ous crops. They often i nvade
bui l di ngs i n fal l . Al l have si l k
gl ands openi ng near mouth and
make a loose web among l eaves.
CHIGGER
Trombicula sp.
adult, 3 mm (0. 1 11
)
VELVET MI TES (Trombi di i dae) are
of l ittl e economi c i mportance
There are several thousand spe
cies. Larvae are parasites on i n
sects; l arge adults (to - mm,
0. 2
11
), usual l y velvety red, eat
i nsect eggs.
WATER MITES (Hydrachnel l ae), a
group of several fami l ies, are
related to trombi di i d mites (p.
1 35). Many are colorful -red,
green or yel l ow. Some crawl on
stones, others swi m. The l ang
hai rs on thei r l egs hel p them to
paddl e, but their bodies are
streaml i ned and nearl y smooth.
Most Water Mites are predators,
but some are parasitic on cl ams
or i nsects. Those that parasitize
dragonfies attach themselves to
nymphs ready to leave the water,
then to become ai rborne on the
newly mol ted adul t. They drop
of i nto the water, possibl y
when the i nsect lays its eggs.
SARCOPTID MI TES (suborder Sarcoptiformes) i ncl udes
the Scabi es or Mange Mites, the Cheese Mites found i n
many stored products and causi ng the al l ergy "grocer's
itch," and the Ori batid Mites.
ORIBATID MITES, al so cal l ed
Beetl e Mites or Moss Mites, in
el ude several fami l ies of free
l i vi ng Sarcoptid Mites. With their
hard, shi ny shel l s they may re
sembl e ti ny dark brown or bl ack
beetles. Some can tuck thei r l egs
under hi nged "wi ngs,'' formi ng a
l itle bal l . Most are len than 1
mm l ong. I n some, the shed ski ns
col l ect on the back, maki ng a
hi gh ornament. Most Oribatid
Mites l ive i n soil or l itter, but
those found on decaying wood
are seen most often. They are of
economic i mportance in break
down of l itter and formation of
humus. Some feed on
eggs and transmi t
Phthiracarus sp.
1 mm (0.04
11
)
wal ki ng
MI TES 1 37
wel l fed
WOD TI CK,
Dermacentor, sp.
2 em (0.8
11
)
TI CKS (suborder l xodi des) are the l argest of the mi tes,
as much as 3 em ( 1 .2") l ong after feedi ng. Al l are ex
ternal parasites of repti les, birds, or mammal s. Most drop
of thei r host after feedi ng. They mol t and then wai t on
the tips of l eaves, forel egs outstretched, ready to attach
to any ani mal brushi ng post. Young ti cks, whi ch hove
onl y three pai rs of l egs, mov attock smal l er host ani mal s
than the adul ts. The bi te of some ti cks may cause mi l d
paralysis to man; others transmi t disease. Ti cks attach
themsel ves to the host onl y wi th thei r mouthparts, and
feed on bl ood. I n removi ng a ti ck, toke core not to l eave
mouthparts behi nd.
About 300 speci es of ticks occur around the worl d.
Soft ti cks (Argasi dae) hove a l eathery i ntegument; they
hove no head pl ate, and the head is on the undersi de.
Hard ti cks ( I xodi dae) hove a hard pl ate above the head
and the head i s di rected forward.
4 mm (0.2
11
)
1 38 MI TES
SOFT TI CKS
BLUE BUG
Argas persicus
parasite af bi rds
and bats; often pests
af paultry i n warm, dry
parts of worl d
Ornithodoros sp.
parasites of mammal s,
i ncl udi ng mon; may
transmi t rel apsing
fever i n western U.S.
5 mm (0. 2
11
)
Dermacentor andersoni
western N.A. , Rocky Mts.;
aHacks l arge mammal s, man;
may transmi t
diseases. D. variabilis
i n eastern N.A.
Boophilus sp.
4
mm (0. 2
11
)
stays on host
throughout l ife
HARD TI CKS
LONE STAR TI CK
Amblyomma americanum
4
mm (0. 2
11
)
southcentral U. S. ;
on l arge mammal s, man
BROWN DOG TICK
Rhipicephalus sanguineus
4 mm (0.2
11
)
found in houses,
rarely bites man;
cosmopol i tan
MI TES
1
39
Euloenenia sp.
2 mm (0. 1 1)
without tail
Texas
MI CROWHI PSCORPI ONS (order Pal pi gradi ) are agi l e
arachni ds l ess t hen 2 mm (0. 1 ") l ong. They l i ve under
stones, goi ng deeper i nto the soi l i f i t gets too dry. Of the
50 to 60 speci es known, 3 occur i n the U . S . , i n Texas and
Cal i forni a; other speci es are found i n the Medi terranean
regi on.
RI CI NULE I DS (order Ri ci nul ei ) resembl e ti cks. They move
about sl owl y in l i tter and l eaf mol d of humi d, warm
areas, and may feed on termi tes . They are 1 0- 1 5 mm
( 0. 4-0. 6") l ong, wi t h a heavy cuti cl e. I n front of the head
i s a uni que hi nged hood coveri ng the j aws . Young Ri ci n
ul ei ds have onl y si x l egs . As recentl y as 1 929 fewer t han
1 40
Crptocellus sp.
4 mm (0.21)
Central ond S.A.
40 speci mens of Ri ci nu
l ei ds were known to
have been found, al l
from Afri ca an d S. A.
Now they are known to
occur wi del y in the
Ameri can tropi cs and i n
Mexi can caves . Onl y
one speci es, found i n
Texas , i s known from
the U. S . The femal e car
ri es the si ngl e egg be
tween the bent pedi
pal ps and the hood.
About 35 speci es are
known .
MYRI AP ODS
Myri apods are the cl asses of arthropods i n whi ch the body
i s made up of numerous s i mi l ar segments .
PAUROPODA, a cl ass cl osel y rel ated to mi l l i pedes, are
soft-bodi ed, l ess than 2 mm (0. l ") l ong, wi th 1 2 segments
and usual l y ni ne pai rs of l egs . The more than 500 descri bed
speci es are found i n the tropi cs and temperate regi ons,
where t hey l i ve i n decayi ng l ogs, forest l i tter, humus, and
under stones. They probabl y feed on fungi and decayi ng
ani mal s . Most are sensi ti ve to l i ght and to dryi ng .
SYMPHYLA, a smal l cl ass of 1 60 known speci es cl osel y
rel ated to centi pedes and mi l l i pedes, are found i n the
tropi cs and temperate regi ons. Symphyl a are 2- 8 mm ( 0. 1 -
0. 3") l ong, wi th 1 5-22 back pl ates, 1 2 pai rs of wal ki ng
l egs, and a pai r of spi nnerets . They l i ve i n moi st , rotti ng
wood or i n soi l and l i tter. Unl i ke centi pedes and mi l l i pedes,
they move al l l egs on one si de together. They feed on
decayi ng vegetati on but may attack l i vi ng pl ants and
become garden pests .
GARDEN CENTI PEDE
Scutigerel/a immaculata
8 mm (0.3
"
)
Europe, N. A., Hawaii;
pest i n gardens
and greenhouses
1 4 1
CENTI PEDES ( cl ass Chi l opoda) have one pai r of l egs on
each segment . The fastest are the ones wi th fewest l egs,
the Scuti geromorphs and Li thobi omorphs . Si nce the pl ates
on thei r back hook together, they can hol d the body
strai ght i n runni ng. The Scol opendromorphs and many
l egged Geophi l omorphs are sl ow and may move wi th
snakel i ke moti ons; al l thei r dorsal trunk pl ates are al i ke
and move freel y. About 2, 500 speci es of centi pedes are
known . The orders of centi pedes are descri bed and i l l us
trated on pp. 1 43- 1 45.
Li ke i nsects an d mi l l i pedes but unl i ke spi ders, centi pedes
have a pai r of antennae. Most are nocturnal . I n dayti me
they h i de i n l i t ter, under l oose bark, stones, l eaves, and
debri s . Some di g i nto the soi l . They avoi d extremel y wet
or dry ni ches . Centi pedes are predators, feedi ng mostl y
on other arthropods. Al l have poi son gl ands openi ng
t hrough thei r j aws , and t he bi te of even a smal l centi pede,
i f i t succeeds i n breaki ng the ski n , can produce pai n.
Scolopendra, the l argest, can gi ve a very pai nful bi te,
but usual l y i s not dangerous. Some centi pedes l ack eyes
and are bl i nd; others do not see wel l . In experi ments i n
whi ch gl ass beads were di pped i n "fl y j ui ce, " some centi
pedes tri ed to bi te the beads . They wou l d not touch gl ass
beads di pped i n water, however, and so i t i s assumed
that centi pedes fi nd thei r prey by smel l and perhaps
by touch.
Some centi pedes can produce si l k, whi ch i s used onl y
duri ng mati ng and when captur i ng prey. When a mal e fi nds
a femal e, they t ouch antennae, and he fol l ows her.
The mal e makes a smal l web i n whi ch he deposi ts a
package of sperm. The sperm is then pi cked u p by the
femal e. The mal e's bui l di ng of a web has been observed
onl y among the Geophi l omorphs, Li thobi omorphs, and
Scol opendromorphs .
1 42 CENTI PEDES
face
compound eye
antenna
maxi l l a 2
Scutigera co/eoptrata
3 em ( 1 . 2
1
)
southern Europe and southern U.S. ;
outdoors. Farther north
in bui l di ngs
SCUTI GEROMORPHA i s an order of mostl y tropi cal
centi pedes. About 1 30 speci es have been descri bed; al l
bel ong t o the fami l y Scuti geri dae. Unl i ke other cen
ti pedes, they have a round head and l arge compound
eyes. They have 1 5 pai rs of l ong l egs. A femal e of Scu
tigera produced 35 eggs over a peri od of days. The
eggs are not guarded. Young centi pedes are born with
onl y seven pai rs of l egs, but the number i ncreases with
every mol t. I n captivity, the common Scutigera found
throughout the U. S. may l i ve more than a year. I t i s the
onl y centi pede common i n houses, where i t runs rapi dl y
al ong wal l s and foors as i t hunts for fi es and other
i nsects. I t i s surpri si ngl y agi l e. I n summer they l i ve outside,
or i n warm cl i mates they l i ve outsi de the year around.
CENTI PEDES 1 43
undersi de of
LI THOBI OMORPHA, or Stone Centi pedes, are al l less
than 4. 5 em ( 1 . 7") l ong. Adults have 1 8 body segments,
1 5 pai rs of legs and 20 to 50 or more segments in the
antennae. As i n Scuti geromorphs, the young have fewer
legs. When di sturbed, they move the l ast pai rs of legs
rapi dly, throwi ng dropl ets of sticky materi al at the poten
ti al predator and sl owi ng it i n the tangl i ng mass. About
1 , 1 00 speci es have been described i n thi s order.
GEOPHI LOMORPHA, or Soi l Centi pedes, are sl ender,
eyel ess centi pedes that have 31 to 1 77 pai rs of l egs and
antennae with 1 4 segments. The l ongest are 1 6- 1 7 em
(6"), but most are l ess than 5 em (2") . The number of
pai rs of legs is always odd, usual l y vari abl e wi thi n the
species, and each l eg can be moved i ndependentl y. Soi l
Centi pedes penetrate as deep as 40 to 70 em ( 1 6-28")
i nto soi l , feedi ng on i nsect l arvae and worms. I f di s
turbed, they coi l up and ej ect from pores on thei r under
si de a secreti on that seems to repel potenti al predators.
A number of species can give of phosphorescent mate
rial . Femal es guard thei r eggs. About 1 ,000 species.
SCOLOPENDROMORPHA have 21 or 23 pai rs of l egs
and 1 7 to 30 segments in thei r antennae. Worl d-wi de
and mai nl y tropi cal , the order i ncl udes the l argest cen
ti pedes. Many are an attractive bl ue-green, ol i ve- green,
or yel l ow. To pi ck one up, i t i s best to use two hands and
forceps as they can i nfi ct a pai nful bite and can al so
pi nch wi th thei r l ast pai r of l egs. They have been ob
served feedi ng on toads and l i zards. These l arge cen
ti pedes di g burrows and al so make chambers i n whi ch
they rest. The femal e often coi l s around her eggs or
young and may peri odi cal l y l i ck the eggs, presumabl y
to cl ean them. About 550 speci es descri bed i n two fam
i l i es: Scol opendri dae and t he bl i nd Cryptopi dae.
Sco/opendra 5g.
1 4 Cm (5. 5
"
)
guardi ng young
S. heros
g 1 5 cm
(611)
Ari zona
CENTI PEDES 1 45
MI LLI PEDES ( cl ass Di pl opoda) are "thousand l eggers , "
di fferi ng from the "hundred l eggers" or centi pedes ( p.
1 42) i n havi ng two pai r s of l egs on most body ri ngs (whi ch
are two fused segments) . They have one pai r of antennae.
Some mi l l i pedes are cyl i ndri cal , adapted for burrowi ng;
others are fl at . Some resembl e woodl i ce ( p. 1 52) . Some
are soft-bodi ed, very smal l , and covered wi th saw-toothed
hai rs (p. 1 47) .
Mi l l i pedes are found under stones, i n moi st soi l and l eaf
l i tter. They avoi d l i ght and feed on vari ous pl ant materi al s,
especi al l y soft decomposi ng pl ant ti ssues . Most mi l l i pedes
have pores, usual l y al ong the si des of thei r body ri ngs,
from whi ch they di scharge strong-smel l i ng secreti ons that
may be repel l ent and poi sonous to other ani mal s . Some
l arge tropi cal mi l l i pedes squi rt thei r secreti ons . Neverthe
l ess, mi l l i pedes are prey of bi rds, toads , and other
ani mal s .
I n most mi l l i pede groups, the l egs on t he seventh r i ng of
the mal e are modi fi ed as copul atory organs (gonopods) .
The shape of these organs i s i mportant i n i denti fyi ng spe
ci es. The femal e l ays her eggs i n the soi l ; some speci es
construct egg capsul es and guard them. J ust-hatched mi l
l i pedes have few r i ngs and three pai rs of l egs, t he number
i ncreasi ng wi th each mol t . Some speci es l i ve onl y one year,
others to seven . More than 1 0, 000 speci es are known ,
about 600 north of Mexi co.
1 46
egg chamber of
Atopetholus
broken open, with young i ntact
Polyxenus lagurus
3 mm (0.
1 11)
Europe, si mi l ar
species i n N. A.
PSELAPHOGNATHA i s a subcl ass of soft mi l l i pedes, al l
l ess than 4 mm (0. 2") l ong, with 1 1 to 1 3 ri ngs. They
have many rows and bundl es of barbed hai rs. Mal es do
not have l egs modi fed for copul ati on. Found i n most
parts of the worl d, there are about 60 speci es of two
fami l i es; Pol yxeni dae has 5 speci es in N. A. They feed
on al gae and l i ve under l oose bark and i n l i tter.
CHI LOGNATHA, the subcl ass i ncl udi ng al l other mi l l i
pedes, have hardened body r i ngs and mal es have l egs
modi fed for copul ati on. There are a number of orders.
GLOMERI DA, Pi l l Mi l l i pedes, are short and wi de, re
sembl i ng woodl i ce (p. 1 52), but have more than seven
pai rs of l egs. The mal e's l ast pair of l egs are modi fed
to hol d the femal e when mati ng. Gl omeri ds can coi l i nto
a bal l when di sturbed, some tropi cal speci es approachi ng
t he si ze of gol f b"l l s. Speci es found i n southern states
and in Cal i forni a are never more than 8 mm (0.3") l ong.
Common i n Europe, where some speci es reach a l ength
of 2 em (0. 8").
PLATYDESMIDA are fat with a tiny head
and from 30 to 1 92 body rings. I n mal es,
the 9th and 1 Oth pairs of l egs are modifed
as copul atory organs (gonopods) . The
usual col or is pi nk. The rough dorsal sur
face has a narrow l ongitudi nal groove.
About a dozen species occur in North
America north of Mexico.
POL YZONI IDA are very si mi l ar to Pl aty
desmi da but are smooth and l ack the
dorsal groove. The usual col or i s cream.
About 1 5 species occur north of Mexi co.
The orders Pl atydesmi da and Pol yzoni i da,
wi th smal l heads, bel ong i n the super
order Col obognatha.
CALLIPODIDA, wi th up to 89 ri ngs, have a pai r of ti ny
spi nnerets on the l ast body r i ng . Most are cyl i ndr i cal ,
wi th l ateral r i dges an d knobs. I n mal es, t h e second pai r
of l egs on the 7th r i ng ar e modi fi ed as gonopods . Most
produce strong-smel l i ng whi te secret i ons from st i nk
gl ands. About 20 speci es are known nor t h of Mexi co,
one wi despread i n wester n Europe and many i n easter n
Medi terranean. Some ar e fast- r unni ng scavengers and
predators.
1 48 MI LLI PEDES
POLYDESMI DA is an order i n whi ch most
speci es have 20 ri ngs that usual l y bear
promi nent keel s whi ch gi ve them a fl at
l ooki ng bac k. Al l l ack eyes; the fi rst pai r
of l egs on t he 7t h r i ng are gonopods .
Many are bri ghtl y col ored; al most al l have
sti nk gl ands . Of 2700 speci es, about 250
occur nort h of Mexi co, fewer i n Europe.
d
Pseudopolydesmus
serratus
GREENHOUSE
MI LLI PEDE
Oxidus gracilis
2.5 em ( 1
"
)
cosmopol i tan i n
greenhouses and
subtropics
3 em ( 1 .2
"
)
eastern N.A.
Pachydesmus crassicutis
7 em (2
.
8")
Motyxia sp . .
4 cm ( 1 .5
"
)
southern Cal iforni a;
bi ol umi nescent
southeastern U.S.
Sigmoria aberrans
4 cm ( 1 . 5")
N. Carol i na, Vi rgi ni a
MILLIPEDES 1 49
J ULIDA is one of four orders of cyl i ndri cal mi l l i pedes
that have both pai rs of l egs on the 7th ring of the mal e
modi fed as copul atory organs (gonopods) . More than
1 00 species are found north of Mexi co. The numerous
members of the fami l y Parai ul i dae, or wi reworms, meas
ure 1 5-90 mm (. 7 -3. 5") l ong. Most are smooth, with up to
74 body ri ngs and the gonopods are outsi de the body.
I n mal es, the frst pair of l egs are greatl y enl arged. I n
the fami ly J ul i dae, the gonopods are i n a pouch, and
t he mal e's frst pai r of l egs are hook-shaped. They ar e
native t o Europe and western Asia; i ntroduced to parks
and gardens i n N.A.
SPI ROSTREPTI DA is an order of l arge (to 28 em, or 1 1 ")
cyl i ndrical mi l l i pedes found mai nl y in the tropi cs. There
i s onl y one pai r of gonopods, the anteri or. In southwest
ern U. S. and adj acent Mexico, mi l l i pedes of the genus
Ortho
p
orus often congregate i n l arge numbers. Three
speci es occur in states adj acent to Mexi co.
Orthoporus pontis
1 2 Cm (4. 5")
western Texas
1 50 MI LLI PEDES
pr
amentum
stipes
mentum
Q
l amel l ae l i ngual i s
mauthparts
c
l l um
Texas; caves
ead
CAMBALI DA have no l egs on the 4th ri ng; other mi l l i
pedes have one pai r. Cambala (fami l y Cambal i dae) are
easi l y recogni zed by the l arge cover (col l um) of the frst
body segment and by the promi nent l ongi tudi nal ri dges
on the body of most speci es. Cambal i da are rare i n the
Great Pl ai ns and semi -deserts. The l argest, to 60 HH
(2. 5"), occur i n the Appal achi an Mountai ns.
SPI ROBOLI DA have on l y one pai r of l egs on the 5th
ri ng; other mi l l i pedes have two pai rs. The mal e's copul a
tory organs (gonopods) are hi dden i n a pouch. The four
cyl i ndri cal groups can al so be separated by the structure
of the mouthparts on the undersi de of the head. About 35
speci es occur north of Mexi co. Narceus pl aces each si ngl e
egg, 1 mm l ong, i n a capsu l e of chewed l eaf l i tter. The
capsul e i s passed posteri orl y by the l egs and i nto the rec
tum where i t i s mol ded and then deposited in a pi l e with
many others.
mouthparts
um
head
Narceus americanus
to 1 0 em
(411)
southeastern U. S. , north to
Ohi o and west to Texas;
i n forest l ogs
MI LLI PEDES 1 5 1
L AND C R US TACE ANS
Crustacea have two pai rs of antennae. Though most are
aquatic (Crayfsh, lobsters, Barnacl es, Shri mps, Water
Fl eas), some have been successful on l and. Beach Fl eas
(Scuds) are found on moi st ocean beaches and i n humi d
tropi cs. Terrestrial Copepods and Ostracods are found
i n the Southern Hemi sphere.
WOODLI CE (order l sopoda) feed on humus and fungi .
North of Mexi co there are nearl y 1 00 speci es. Eggs
and young are carri ed i n a brood pouch by the femal e.
ROCK SLAYERS (li gi i dae) are am
phi bi aus an ocean beaches. They
feed on seaweed, mai nl y at low
tide. The smal l end of the l ong
antennae (second pai r) has more
than ten segments.
PI LL BUGS (Armadi l l i di idae) have
an arched bady and can rol l i nto
a bal l when di sturbed. The two
tai l s ( uropods) are shorter than
the l ast abdomi nal segment .
.
PI LL BUG
Armadi/idium vulgare
1 .4 em (0.61)
cosmopol itan
1 52 LAND CRUSTACEANS
TRI CHONI SCI DS (Tri choni sci dae)
are found in wet 9QDf9. They are
mal l , narrow-bodi ed, and have
onl y four to fve segments i n
smal l end of the l ong antennae
(second pai r). Both the a ntennae
and tai l s ( uropods) are ti pped
by a brush.
SOW BUG is a nome used for
species i n two fami l ies. Those in
the fami ly Oniscidae, represented
here by Oniscus, have three seg
ments i n the small, segmented
end of the l ong antennae (sec
ond pai r); most cannot rol l i nto
a bal l . Members of the fami l y
Porcel l i oni doe have two seg
ments i n the smal l , segmented
end of the l ong antennae (sec
ond pai r).
Metoponorthus
pruinosus
cosmopol itan;
Porcel/io scober
1 .4 em (0.611)
wi despread,
i n northern U.S.;
tubercl es al l over
Trachelipus rathkei
1 . 2 em (0
.
511)
cosmopol i tan, bui l di ngs;
tubercl es on head;
i n , 3rd segment
from end of 7th l eg
has keel
Cylisticus convexus
1 2 mm (0.511 )
Europe, eastern U.S. ;
shi ny,
con rol l up
LAND CRUSTACEANS 1 53
LAND CRABS (Gecarci ni dae), found onl y in the subtrop
i cs and tropi cs, are l and-dwel l i ng crustaceans, but the
femal es return to the ocean to reproduce. They di g tun
nel s 30-40 Lm deep ( 1 2- 1 6"), 1 8 LH (7") i n di ameter,
and come out at ni ght to feed. littl e ones cl i mb wal l s
and trees.
Land Hermit Crabs (Coenobi tidae) can give a good
pi nch with their col orful cl aw if handl ed carel essl y. Most
are scavengers. In the Southwest Pacifc, the Coconut
Crab (Birgus), growi ng to 45 Lm ( 1 8") i n l ength, feeds
on fal l en coconuts, and can be destructive to crops. I t i s
consi dered a del icacy itsel f.
1 54
LAND HERMI T CRAB
Coenobita clypeatus
cl aw to 6 em (2. 3
"
) di ameter
eastern Cari bbean, southern Fl ori da
LAND CRAB
Gecarcinus latera/is
to 9 em (3.511) wi de
Fl ori da Keys, Bermuda,
West I ndi es
MOR E I N F ORMAT I ON
Baker, E . W. et al . , A Manual of Parasi t i c Mi tes, Nat' l Pest Control
Assoc . , 1 956. A l ot of i nformati on on mi tes of economi c i mportance.
Barnes, R . D. , Invertebrate Zool ogy, Saunders, Phi l adel phi a, 1 987. A
good textbook gi vi ng background on Crustacea, Myri apods and Arach
ni ds.
Bonnet, P. , Bi bl i ographi c Araneorum, Toul ouse, 7 Vol . , 1 945-1 962. An
i ndi spensabl e reference work l i sti ng al l spi der l i terature up to 1 938. I n
l arger l i brari es onl y.
Cl oudsl ey-Thompson, J . l . , Spi ders, Scorpi ons, Centi pedes and Mi tes,
Pergamon Press, london, 1 968. A very useful , accurate book.
Foel i x, R . F. , Bi ol ogy of Spi ders. Harvard Uni versi ty Press, Cambri dge,
1 982. Spi der bi ol ogy wi th emphasi s on physi ol ogy and behavi or.
Gertsch, W. J . , Ameri can Spi ders, 2nd ed. Van Nostrand- Rei nhol d, New
York, 1 978 . Natural hi story of Ameri can spi ders.
Grasse, P. , edi t . , Trai te de Zool ogi e, val . 6, Masson & Ci e, Pari s, 1 949.
One of t he best summari es of bi ol ogy, anatomy and systemati cs of spi ders
and al l i ed groups; i n French.
Jones, D. The Larousse Gui de to Spi ders, Larousse, N. Y. , 1 983. Pi ct ures
of European spi ders.
Kaestner, A. , adapt. levi , H. W. and l . R. levi , Invertebrate Zool ogy
val . 2: Art hropod Rel ati ves, Chel i cerata, and Myri apoda. R. E .
Kri eger Publ . Co. , Mel bourne, F l , 1 980. A summary o f our knowl edge
of arachni ds and myri apods; i n Engl i sh .
Kaston, B. J . , How t o Know the Spi ders, 3rd ed. W. C. Brown Co . ,
Dubuque, I owa, 1 978. Keys t o common spi der genera.
Kaston, B. J . , Spi ders of Connecti cut, rev. ed. Bul l . Conn . Geol . Nat .
Hi s! . Surv. 70, 1 98 1 . The most useful reference book on spi ders of
eastern U. S .
Krantz, G. W. , A Manual o f Acarol ogy. Oregon State Uni v. Bookstore,
Corval l i s , rev. ed. 1 978.
locket, G. H . and A. F. Mi l l i dge, Bri t i sh Spi ders, Ray Soc . , london,
1 95 1 - 1 953. Speci al i zed, separati ng spi der speci es of Great Bri tai n .
Roth, V. D. , Spi der Genera of North Ameri ca. Publ i shed by author,
Portal , AZ, 1 986. Key to fami l i es and genera found in North Ameri ca.
Shuttl esworth , D. E . and S. . Swai n, The Story of Spi ders, Garden Ci ty,
New York, 1 959. An excel l ent chi l dren's book.
Yagi numa, T. , Spi ders of Japan i n Col or, Osaka, Japan, new ed . 1 986.
I n Japanese; wel l i l l ustrated, the names of spi ders i n lat i n.
1 5 5
Abaci on, 1 48
abdomen, spider, 8, 9,
1 3
Acacesi a, 6 1
Acanthepei ra, 67
Acanthocteni dae, 1 1 2
Acanthoctenus, 1 1 2
Acari , 1 34- 1 39
accessory cl aws, 1 3
Achoearaneo, 40, 50
Acrographinotus, 1 30
Aculepei ra, 64
Ael uri l l us, 1 00
Agel ena, 73
Agel eni dae, 72-76
Agelenopsi s, 72, 73
Amaurobi i dae, 1 1 1
Amaurobi us, 1 06, 1 1 1
Amblyomma, 1 39
Amblypygi , 1 1 7
American House
Spider, 36
Ammotrechel l a, 1 1 9
Ammotrechidoe, 1 1 8
Amphipoda, 1 52
anal tubercl e, 9, 1 3, 3 1
Androctonus, 1 26
Anelosi mus, 38
Anoples, 1 2
ant mi mi cs, 89
Antrodi aeti da, 23
Antrodiaetus, 23
Anuroctonus, 1 25
Anyphaenidae, 88
Aphonopel ma, 2 1
Apochthoni us, 1 2 1
Arachni da, 4
"
Araneidae, 52-69
Araneus, 53, 56-59
Aroniello, 59
Archaea, 51
Archaeidoe, 51
Arctoso, 85
Arenophi l us, 1 44
Argas, 1 38
Argasi dae, 1 38
Argiope, 1 4, 53, 68-69
Argi opi dae, 52-69
Argyrades, 39
Argyroneto, 76
1 56
I NDX
Femal e,
Argyronetidoe, 76
Ari adna, 27, 1 08
Armadi l l i di i dae, 1 52
Armadi l l i di um, 1 52
Arrenurus, 1 36
Arrowshaped
Micratheno, 67
Arthropoda, 4, 6
Assomi i dae, 1 29
Atopethol us, 1 46
Atrax, 1 6, 24
Atypidae, 25
Atypus, 25
Aysha, 88
bal l ooni ng, 1 5
Banana Spider, 92
Banded Argiape, 68
Barn Spider, 58
Barychel i dae, 2 1
Basi l i ca Spider, 64
Bathyphantes, 49
Beach Fleas, 1 52
Beetle Mites, 1 37
Bi rd Spi ders, 20, 2 1
Bi rgus, 1 54
Bl ack & Yel low
Argiope, 68
Bl ack Widow, 1 7, 36, 42
Blue Bug, 1 38
Bolas Spider, 66
book l ung, 9, 1 2- 1 3
Boophi l us, 1 39
Bothriocyrtum, 23
Bothri uri doe, 1 24
Bothri urus, 5, 1 24
Bowl & Doi l y
Spi der, 47
Brachycybe, 1 48
Brown Centruroides,
1 27
Brown Dog Ti ck, 1 39
Brown Recluse
Spider, 1 7
Brown Spi ders, 1 6, 29
Brown Wi dow, 43
bul b, 9
Burrowing Wolf
Spiders, 85
Buthidae, 1 26- 1 27
Mal e
Buthus, 1 26, 1 27
Caddi dae, 1 32
Caddo, 1 32
cal ami strum, 1 06
Cal l i lepi s, 87
Cal l i podi dae, 1 48
Cal l obi us, 1 1 1
Cambol a, 1 5 1
Cambol i da, 1 5 1
Caponi i dae, 26
carapace, 8, 9, 1 6
Carol i na Wolf Spi der, 83
Casti anei ra, 89
Castor Bean Tick, 1 39
Cattle Ti ck, 1 39
Cave Orb-weaver, 60
Cave Spi ders, 32, 5 1
Centipedes. 1 42- 1 45
Centruroi des, 1 23, 1 26- 1 27
cephalothorax, 8, 9, 1 6
Ceraticel us, 45
Ceratolasmatidae, 1 3 1
Chacti dae, 1 24
Choronti dae, 1 1 7
Cheese mi tes, 1 37
Chei racanthi um, 1 6, 89
chel i cera, 8, 9, 1 6
Chel icerata, 4 , 6
Chel i ler, 1 2 1
Chi ggers, 1 35
Chi lognatha, 1 47
Chi lopoda, 1 42- 1 45
Chi trel l a, 1 2 1
Chryssa, 38
Ci curi na, 75
claws, 1 3 , 1 29, 1 30
Cl ubi ona, 88-89
Cl ubi noni dae, 88-89
Cobweb Weavers,
36-43, 5 1
'
74. 1 1 0
Coelotes, 73
Coenobita, 1 54
Coenobi ti dae, 1 54
Coleosomo, 39
col l ecti ng spi ders, 1 8
col l um, 1 5 1
Col obagnatha, 1 48
col ul us, 8, 9, 1 3, 5 1
comb, 36, 1 26
Combfooted Spi ders, Drapeti sca, 48 Gecarci ni dae, 1 54
36-43 Drassodes, 87 Gecarci nus, 1 54
Copepoda, 1 52 Dwarf Spiders, 44-45 geni tal pl ate, 1 26
Caras, 74 Dysdera, 27 Geol ycosa, 85
Cori arachne, 96 Dysderi dae, 27 Geophi l omorpho, 1 44
Corythol i a, 1 04 Gi ant Crab Spi ders, 92
Cosmetidae, 1 30 egg sac, 1 0, 1 4
Gi ant Hai ry
courtshi p, spider, 1 0 embol us, 9 Hadrurus, 1 25
coxa, 8, 9, 1 6, 1 29, endi te, 8, 9, 1 6, 25
Gl omeri da, 1 47
1 30, 1 3 1 Enoplognatha, 4 1 , 5 1 Gl omeri s, 1 47
Crab Spi ders, 92, 93 epi gynum, 8, 9, 1 3, 70 Gnaphosa, 86
94-97 Epi si nus, 38 Gnaphosi dae, 86-87,
Cri bel l ate Spi ders, Eremobates, 1 1 9 88, 89, 90, 1 1 2
1 06- 1 1 5 Eremobatidae, 1 1 8 gnathochi l ari um, 1 5 1
cri bel l um, 3, 8, 1 06 Eresi dae, 1 09 Gol den lynx Spi der, 77
Crosbycus, 1 3 1 Eresus, 1 09 Gol den Si l k Spi der, 65
Cross Spider, 56 Eri go'e, 45 gonopods, 1 46
Crustaceans, 4, 5, Eri s, 1 0 1 gonopore, 8
1 52-54 Ero, 50 Gonyl epti doe, 1 30
Cryphoeca, 75 Euagrus, 24 gossamer, 1 5
Cryptopidae, 1 45 Eukoeneni a, 1 40 Grass Spi der, 73
Cryptocel l us, 1 40 European Greenhouse
Cryptothel e, 3 1 Buthus, 1 26 Mi l l i pede, 1 49
Cteni dae, 1 4, 9"t House Spider, 74 Green lynx Spi der, 77
Cteni zi dae, 22 Tarant do, 83 Grocer
'
s I tch Mi tes,
Cupi ennus, 91 Water Spider, 76 1 37
Cyclocosmi a, 22 Euryopi s, 39 growth, spi der, 1 1
Cycl ase, 63 Euscorpi us, 1 22, 1 24
Cyl i sti cus, 1 53 Eustal a, 62 Habrocestum, 1 00
cymbi um, 9 Evarcha, 99 hackl edthreads, 1 06
Cyphophthal mi , 1 28 eyes, 8, 1 6 Hadrobunus, 1 33
Cyrtophol i s, 2 1 Hadrurus, 1 25
Cyrtophora, 64 fang, 1 6, 1 43, 1 44 Haemaphysal i s, 1 39
Featherlegged Spider, 1 1 4 Hahni a, 75
Daddy-l ong-l egs, femur, 9 Hahni i dae, 75
1 28, 1 32- 1 33 Fi l istata, 1 1 , 1 08 Hai ry Mygal omorphs,
Daddy-l ong-l egs Fi l istatidae, 1 08 20-2 1
Spiders, 32 Fi shi ng Spiders, 80-81 Hamatal i wa, 77
Dermacentor, 1 38, 1 39 Floricomus, 45 Hammock Spi der, 46
Di ctyna, 1 1 0 Fl ori nda, 48 Hapl odrassus, 87
Di ctyni dae, 1 1 0 Fl ower Spiders, 94 Hapl ogyne Spi ders,
di ggi ng rake, 23 Fol di ng Door 8, 26-30
Oi gueti a, 30 Spider, 23 Hard Ti cks, 1 38, 1 39
Di gueti dae, 30 Fronti nel l o, 47 Harvestmen, 1 28- 1 33
Oi nopi dae, 1 1 3 Funnel Weavers, Harvest Mi tes, 1 35
Di nopsi s, 1 1 3 72-76, 1 09, 1 1 1 Hasari us, 1 04
Di pl ocentri doe, 1 22 Funnelweb Hoseth
'
s
Di plocentrus, 1 22 Mygalomorphs, Centruroi des, 1 27
Di plopoda, 1 46- 1 5 1 1 6, 24 Hel ophora, 48
Di plosphyroni da, 1 2 1 Furrow Spider, 58 Hentz
'
s
Di pl uri dae, 24 Centruroi des, 1 27
Di poena, 39 Garden Centipede, 1 4 1 Herpyl l us, 87
Dol i omal us, 93 Garden Spider, 56 Hersi l i a; 35
Dolomedes, 78, 80- 81 Gasteracantha, 66 Hersi l i i dae, 35
dragl i nes, 1 0, 1 5 Gea, 69 Heteropoda, 92
1 57
z
Heterosphyroni da, 1 2 1 li gi i dae, 1 52 Mi crommata, 92
Hexuro, 24 li mnochares, 1 36 Mi crowhi pscorpi ons,
Homol onychi dae, 90 li mul us, 6 1 40
Homol onychus, 90 li nyphi a, 47 Mi cryphanti nae, 44
House Pseudoscorpi on, li nyphi i dae, 44-49 Mi gi doe, 22
1 2 1 li nyphi i nae, 46-49 Mi l l i pedes, 1 46- 1 5 1
House Spi der, 36, 74 li phi sti i doe, 5, 7 Mi meti dae, 50
Huntsman Spi der, 92 li thobi omorpho, 1 44 Mi metus, 50
Hydrachnel l ae, 1 36 li thobi us, 1 44 Mi sumena, 94
Hypochi l i dae, 1 07 lobed Argi ape, 69 Mi sumenops, 94, 95
Hypochi l us, 1 06, 1 07 lone Star Ti ck, 1 39 Mi te Harvestmen, 1 28
Hypsel i stes, 45 long-bodi ed Cel l ar Mi tes, 1 34- 1 39
Hypti otes, 1 1 5 Spi der, 33 Mi topus, 1 32
long- jawed mol ti ng, 1 1
l ci us, 1 0 Orb-weavers, 72 Monkey Spi ders, 20- 21
lschyropsal i di dae, 1 32 Loxosceles, 1 1 , 1 7, 29 Monosphyroni do, 1 2 1
lschyropsol i s, 1 32 loxoscel i dae, 29 Mordant Uroctonus, 1 25
lsometrus, 1 26 l ung covers, 25 Moss Mi tes, 1 37
l sopoda, 1 52- 1 53 l ung sl i ts, 27, 1 06 Motyxi a, 1 46, 1 49
lsoxyo, 67 lycoso, 1 6, 82, 83 Mygal omorpha, 20
I tch Mi te, 1 37 lycosi dae, 82-85 Mygal omorphs,
I xodes, 1 39 lynx Spi ders, 77 20, 1 07, 1 09
l xodi des, 1 38- 1 39 lyssomanes , 1 05 Myri apoda, 5, 1 4 1
I xodi dae, 1 38- 1 39 Myrmeki aphi l a, 22
mal l et-shaped organs, Myrmonyssus, 1 34
v
jaws, spi der, 8, 1 0, 26 1 1 8- 1 1 9
o
Jul i da, 1 50 Mal mi gnotte, 42 Norceus, 1 5 1
g u
r
Jul i dae, 1 50 Monge Mi tes, 1 37 Nebo, 1 22
u
=
Si x-eyed Crab
Spi ders, 28
Si x-spotted Fi shi ng
Spi der, 80
Soft Ti cks, 1 38
Soi l Centi pedes, 1 44
Sol i fugae, 1 1 8- 1 1 9
Sol pugi dae, 1 1 9
Sosi ppus, 85
Sow Bug, 1 53
Sporassi dae, 92
spermatophore, 1 22
Spermophora, 32, 33
Sphodros, 25
Spi der Mi tes, 1 35
spi der rel ati ves, 1 1 6
Spi ned Mi crathena, 67
spi nnefets, 8, 9, 1 3, 1 48
Spi ntharus, 38
z
Spi nturni x, 1 34
Spi robol i da, 1 46, 1 5 1
L
Spi rostrepti da, 1 50
Spi tti ng Spi ders, 28
Spotted l sometrus, 1 26
Stri gomi o, 1 44
Z
v Stri ped Centruroi des,
1 27
M
l '
AGO DEN - GUI DE