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The Skin
- protects underlying cells against damage by ultraviolet sunlight - In endothermic animals, it is concerned with temperature regulation including insulation and cooling. - contains sensory receptors. - excretory functions and sometimes respiratory functions - Skin pigmentation may assist in camouflage and signaling - Skin secretions may make the animal sexually attractive or influence other animals behaviors.
Invertebrate Integument
Protozoa have plasma membranes or a protective pellicle. Most multicellular invertebrates have complex tissue coverings. A molluscan epidermis is soft and contains mucous glands; some secrete the calcium carbonate shell. Cephalopods have a more complex integument of cuticle, simple epidermis, connective tissue, reflecting cells (iridocytes), and a thicker layer of connective tissue.
Arthropod Exoskeleton
a. Firmness of exoskeleton & jointed appendages - muscle attachment b. A single-layered epidermis or hypodermis secretes a cuticle with two zones : Procuticle and epicuticle c. Arthropod cuticle may remain soft and flexible as in small crustaceans and insect larvae. Arthropod cuticle is one of the toughest animal materials, yet it is very light.
Two zones 1. Procuticle - thicker, inner, made of protein & chitin layers. 2. epicuticle thin, nonchitinous, made of proteins and lipids, and provides moisture-proofing.
Calcification deposition of calcium carbonate in procuticle - cuticle is stiffened Sclerotization formation of highly resistant protein Sclerotin
Structure
a. An example of vertebrate skin is frog or human skin. b. The thin, outer stratified epithelial layer is epidermis derived from ectoderm. c. The inner, thicker layer is dermis derived from mesoderm. d. The dermis contains blood vessels, collagenous fibers, nerves, pigment cells, fat cells and connective tissue cells, all to support and nourish the epidermis. e. The epidermis is stratified squamous epithelium. f. The basal part is made of cells that undergo frequent mitosis for renewal g. Dermis supports the epidermis; any bony structures derive from dermis tissue.
Dunkleosteus
Bothriolepsis
Scales of contemporary fishes are bony dermal structures evolved from bony armor of early fishes.
In reptiles, dermal bones form the armor of crocodilians, the beaded skin of lizards, and contribute to the shell of turtles.
Teleosts bony scales from dermis retained throughout life Lizards horny scales from epidermis; shed periodically
Claws, beaks, nails and horns are made up of combinations of epidermal and dermal components.
Animal Coloration
a. Structural colors are produced by the physical structure of the surface tissues. b. More common are pigments, varied molecules that reflect specific light rays. c. In crustaceans, these pigments are in large cells with branching processes called chromatophores. d. Pigments may concentrate in the center of a cell or disperse throughout the cell for display.
Melanins are black or brown polymers responsible for earth-colored shades. Carotenoid pigments provide yellow and red colors often contained inside xanthophores. Ommochromes and pteridines are responsible for yellow pigments of molluscs and arthropods. Green is usually produced by yellow pigment overlying blue structural color. Iridophores contain crystals of guanine or another purine rather than pigment; they produce silvery or metallic colors.
Primates have color vision But most mammals are color blind
Mammals are relatively uncolorful, a fact related to their being mostly colorblind. Primates are an exception, and the brilliant skin patches of baboons and mandrills reflect this. Dermal melanophores deposit melanin in growing hair of mammals, providing the color.
Chapter 29
Support, Protection, and Movement
Skeletal Systems
Hydrostatic Skeletons
1. Many invertebrates use their body fluids as an internal hydrostatic skeleton. 2. Muscles in the body wall of an earthworm contract against the coelomic fluids that are incompressible.
Rigid Skeletons
Rigid skeletons provide rigid elements to which muscles can attach. 2 Types : Exo and Endo 1) Exoskeleton - protective skeleton that often must be molted to allow growth - molluscs, arthropods & other inverts
e.g. molluscs - shell grows with the animal
2) endoskeleton - echinoderms and vertebrates. Vertebrate endoskeleton composed of bone and cartilage - protects and supports - major reservoir for calcium and phosphorus - origin of blood cells
Cartilage - major skeletal element of jawless fishes and elasmobranchs Hyaline cartilage - protein gel interlaced with a meshwork of collagenous fibers - joints, support tracheal, laryngeal and bronchial rings
Bone
living tissue with significant deposits of Ca++ Strength of iron yet
1/3 as heavy
Bone Types
Most bone develop from cartilage and is called endochondral or replacement bone. Intramembranous bone - develops from sheets of embryonic cells, mainly the face and cranium.
Cancellous or spongy bone - has an open, interlacing framework of bony tissue. Some bones proceed to add additional salts to become compact bone.
Skull
Movement from water to land forced dramatic changes in body form. Over time, many skull bones were lost or fused Amphibians have from 50 to 95, mammals have 35 or fewer and humans have 29.
Vertebral Column
main stiffening axis and serves the same function as a notochord. Vertebrae are separated into cervical thoracic, lumbar, sacral and caudal. In birds and humans, the caudal vertebrae are reduced in size and number and sacral vertebrae are fused Humans have 7 cervical or neck vertebrae, 12 thoracic, and five lumbar or back vertebrae. 1st cervical vertebra is the atlas - pivot 2nd cervical vertebra is the axis - turn side-to-side
Ribs
serve as stiffening rods and improve effectiveness of muscle contractions Some vertebrates have reduced ribs; In mammals, the ribs from the thoracic basket prevent collapse of the lungs. Primates other than humans have 13 pairs of ribs; humans have 12 pairs, rarely a 13th pair. Ribs of birds are flattened and the sternum is keeled for the attachment of flight muscles
Appendages
Most vertebrates have paired appendages. Fishes, except agnathans, have pectoral and pelvic girdles supporting pectoral and pelvic fins. In tetrapods, the pelvic girdle is firmly attached to transmit force. The pectoral girdle is more loosely attached to allow greater freedom for manipulation.
Animal Movement
Mechanism Important characteristic of animals, compared to plants Most animal movement relies on a single fundamental mechanism: contractile proteins contract when powered by ATP. The most important protein contractile system is composed of actin and myosin.
The acto-myosin system is almost universal and is found from protozoa to vertebrates.
Ameboid Movement found not only in amebas, but also in wandering cells of metazoans (sperm of nematodes) Ameboid cells change shape by extending and withdrawing pseudopodia on any cell surface. Locomotion is assisted by membrane-adhesion proteins that attach to the substrate to provide traction.
Ciliary Movement
Cilia - minute, hairlike, motile processes that extend from the surfaces of many animal cells. found in all major animal groups (not in nematodes) propel fluids & materials across epithelial surfaces in larger animals. Each cilium contains a peripheral circle of nine double microtubules around two central microtubules. each microtubule has a spiral array of protein subunits called tubulin.
Flagellar movement
The flagellum is whiplike, longer than a cilium and present in fewer numbers. Flagella are found in flagellate protozoans, animal spermatozoa, sponges. Flagella differ more in their beating pattern than in structure. beats symmetrically with snakelike undulations to propel water parallel to the long axis.
A cilium beats asymmetrically with a fast power stroke in one direction followed by a slow recovery; water is propelled parallel to the ciliated surface.
Muscular movement
Contractile Tissue Contractile tissue is most highly developed in muscle cells, called fibers. Can be arranged in so many combinations that any movement is possible.
Smooth Muscle
Smooth or visceral muscle lacks the alternating bands or striations. Cells are long, tapering strands, each containing a single nucleus. muscle cells form sheets of muscle circling the walls of the alimentary canal, blood vessels, respiratory passages, and urinary and genital ducts. Controlled by the autonomic nervous system, contractions are involuntary and unconscious.
Cardiac Muscle
has striations but is uninucleate with branching cells. muscle tissue of the vertebrate heart and is seemingly tireless It is fast acting and striated like skeletal muscle, but contraction is under autonomic control. The heartbeat originates within the specialized muscle; the autonomic nerves merely speed up or slow down this rate.
Thick Filaments Each thick filament is made of myosin molecules packed together in a bundle. Thin Filaments Thin filaments are composed of three different proteins. a. actin - backbone twisted into a double helix b. Tropomyosin 2 strands surrounding the actin c. Troponin - calcium-dependent switch that acts as the control point in contraction