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Limiting factors
Limiting factors are elements of an ecosystem that
are in short supply and therefore set limits on
population size
Carrying Capacity (K)-maximum number of
individuals that can occupy an ecosystem
Density-dependent limiting factors-factors whose
limiting effects increase as population density
increases (ex: disease, famine, etc). Some cause an
increase in competition.
Density-independent limiting factors-factors whose
limiting effects are not tied to population density (ex:
natural disasters, climate, etc)
Growth models
Exponential model-also known as a J curve. Assumes
that populations can grow without limit
Logistic model-also known as an S curve. Assumes
that populations know what K is, and will act
accordingly
This changes the reproductive rate equation:
Change N/Change t=rN(K-N/K)
Notice, K is taken into consideration here
Reality-models are only as good as their assumptions,
which means that the graph of real population growth
is slightly different.
Figure 52-7a
Carrying capacity
Later growth
falls to zero
Early growth
is rapid
Growth begins
to slow
Growth and Life History
These growth models are associated with 2
kinds of life-strategies for organisms:
r-selected species=these exhibit rapid,
exponential growth. These are often called
opportunitstic species because they quickly
invade an area, reproduce and die. Offspring
mature quickly and are small. (ex: grasses,
insects)
K-selected species=these are species whose
populations are relatively stable, usually around
K. They produce a small number os offspring
that are large and require lots of care. They
reproduce repeatedly (ex: humans)
Human Population growth
Human population worldwide is reaching 9 billion. It
was just 3 billion 100 years ago.
Why the rapid rise?
Increases in food supply and travel-humans have
domesticated, bred, and fine-tuned agriculture (from
hunter-gatherer to farmer)
Reduction in disease-advances in medicine, like
vaccines, antibiotics, etc have dropped the death rate
and increased the successful birth rate
Reduction in wastes-sewage systems and water
treatment have reduced health hazards
Expansion of habitat-better housing, clothing, etc
have made it easier to live in more places
Figure 52-16-Table 52-2
Figure 52-17
n t
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Cu igh
H
ed iu m
M
Low
Population cycles: Predator/Prey
Predator/prey describes a relationship
between a hunter and an individual that is
eaten.
In general, changes in the prey population
cause similar changes in the predator
population since the predator is dependent
on the prey. Just keep in mind, the prey
are usually predators of the producers, so
their population changes are often due to
seasonal changes in their prey population.
Figure 52-12
Hare Lynx
Community Ecology
Habitat-the area that an organism inhabits within an
ecosystem
Niche-the role of the organism within the environment1
species per niche
Gauses principle of competitive exclusion=when 2 species
try to occupy the same niche, there will be competition
until one species leaves or dies
Resource partitioning=some species can coexist even
though they appear to be competing for the same
resources. They are occupying slightly different niches by
using the resources in different ways.
Character displacement (niche shift)=as a result of
resource partitioning, certain characteristics allow
organisms to obtain their partitioned resources more
successfully
Realized niche=This is the niche that the organism occupies
b/c of resource partitioning. If there were no competitors,
they would otherwise occupy their fundamental niche, but
because of niche overlap, they must adjust
Figure 53-2
These trees are competing for water Space preempted by these barnacles is The large fern has overgrown other individuals
and nutrients. unavailable to competitors. and is shading them.
Few plants are growing under these Grizzly bears drive off black bears. Spotted hyenas and vultures fight over a kill.
Salvia shrubs.
Figure 53-4a
Paramecium
aurelia
Paramecium
caudatum
Figure 53-4b
Higher fitness
Same
Lower fitness fitness
Figure 53-4c
Fundamental
niche
Realized
niche
Figure 53-3
One species eats seeds of a certain size range.
External energy
source, usually
solar energy but
also chemical energy
Abiotic environment
The soil, climate, atmosphere,
and the particulate matter
and solutes in water
Consumers Decomposer
Organisms that eat Organisms that feed
other living organisms on dead organisms or
their waste products
Figure 54-5
Trophic Feeding Decomposer Grazing
level strategy food chain food chain
Quaternary
5
consumer
Coopers hawk
4 Tertiary
consumer
Robin Coopers hawk
Secondary
3
consumer
Earthworm Robin
Primary
2 decomposer
or consumer
Bacteria, archaea Cricket
1 Primary
producer
Dead maple leaves Maple tree leaves
Figure 54-6
Production of biomass
(g/m2/year)
Tertiary consumers 3
10
%
Efficiency of
Secondary consumers 30 15% energy transfer
Primary consumers 20
and decomposers 200 %
Primary
producers 1000
Types of predator/prey
relationships
True predators-kills and eats another
animal
Parasites-are only predatory if they kill
their host
Parasitoid-insects that lay eggs on a host.
The larvae are parasitic to the host
Herbivores-yes, theyre technically
predators. Some are seed-eaters
(granivores), some eat grasses (grazers),
and some eat other plant material
(browsers)
Avoiding Predation
Organisms have evolved many mechanisms for
avoiding predators.
Secondary Compounds-toxic chemicals produced by
plants that can make herbivores sick
Camouflage (cryptic coloration)-helps the animal
blend into its surroundings (some predators use this
as well to help them hunt)
Aposematic coloration (warning coloration)-a bright
color pattern that advertises that the organism should
be avoided (ex: wasp/bee stripes)
Mimicry-organisms resembling each other (shortens
the predators learning curve)
Mullerian mimicry-dangerous organisms resemble
each other
Batesian mimicry-organisms without a defense
mechanism resemble a dangerous organism
Figure 53-12
Cottonwood tree felled by beavers Resprouted trees have more Survival of beetle larvae
defensive compounds. placed on ant mound
Figure 53-10
Prey and predator
Camouflage: blending into the background Schooling: safety in numbers Weaponry: fighting back
B
C
Species
D
E
Species richness: 6 6 5
Species diversity: 0.59 0.78 0.69
Figure 55-4
Hotspots in terms of species richness of birds
Invasive species increase competition. Invasive species introduce disease. Invasive species increase predation.
Purple loosestrife is crowding out native An introduced fungus has virtually wiped out The brown tree snake has extinguished dozens
organisms in North American marshes. the American chestnut. of bird species on Guam.
Figure 53-19
P. ochraceous
(keystone predator)
present
P. ochraceous
(keystone predator)
absent
Community Change
Succession-a series of more-or-less orderly changes
in an ecosystem over time. Begins with a pioneer
species (usually an r-selected species) and ends with
a climax community (stable)
In general, as organisms inhabit the area, they
changes the texture, pH, and water potential of the
soil, as well as establish competition for resources as
the area becomes crowded
Primary Succession-occurs on a substrate that has
never before supported life.
Secondary Succession-occurs on a substrate that has
gone through some sort of disturbance
Succession on land
In primary succession, generally starts with
lava flow or sand.
Pioneer for lava-lichens
Pioneer for sand-grasses
In secondary succession, you can start at
any point in the successional process,
usually a field.
These used to be thought of as a
predictable series of changes, however,
there are instances that are more random
and less orderly b/c theyre affected by
climate, which species happen to arrive
first, etc.
Figure 53-21-1
Old field
Pioneering species
Figure 53-21-2
Early successional
community
Mid-successional
community
Figure 53-21-3
Late-successional
community
Long-lived tree
species mature.
Climax community
Succession in Water
This happens when you start with a
lake or pond that changes to a
marsh-like state.
The marsh is followed by meadow,
with lots of grasses.
Finally, theres a climax community of
native vegetation
Ecosystem Ecology
Biogeochemical Cycles-this is the flow of important elements through the
ecosystem.
Hydrologic cycle-mainly an abiotically-driven cycle.
Reservoirs-ocean, air, groundwater, glaciers
Assimilation-plants absorb water from the soil, animals drink water
and eat organisms
Release-transpiration, evaporation, etc.
Carbon cycle-mainly biotically-driven and tied to atmospheric CO2 levels
that cause the greenhouse effect
Reservoirs-atmosphere, fossil fuel, peat, organic material (like
cellulose)
Assimilation-photosynthesis, animals eating plants and each other
Release-respiration and decomposition
Nitrogen cycle-also a biotically driven cycle dependent heavily on
bacteria
Reservoirs-atmospheric N2, soil (nitrates, nitrites, ammonium,
amonia)
Assimilation-nitrogen fixation by bacteria, nitrification by bacteria
Release-denitrification by bacteria, decomposition, animal waste
Phosphorous cycle-again, a biotically-driven cycle
Reservoirs-rocks and ocean sediments
Assimilation-plants absorb phosphates and are eaten by animals
Release-decomposition, animal waste
Figure 54-13
Percolation
o undwater: 36
Runoff and gr Water table
(saturated soil)
Figure 54-14
Organisms, soil,
litter, peat: 2190
Rivers (erosion):
Organisms, chemical 0.8
processes in ocean:
40,000
Industrial
Protein and fixation: 100
Bacteria in mud nucleic acid
synthesis
use N-containing
molecules as energy Lightning
sources, excrete N2: and rain: 3
310 Internal Nitrogen-fixing bacteria
Nitrogen-fixing cycling: in roots and soil: 202
Internal cyanobacteria: 1200
Runoff: 36
cycling: 15
8000
Mud Decomposition of
Permanent burial: 10 detritus into ammonia
Biomes
These are ecosystems that have
characteristic biotic and abiotic factors
Land Biomes are largely determined by
latitude (except desert which is determined
by climate)
As you move from the poles toward the equator,
biodiversity and biomass increase. The length of
the growing season also increases.
Water Biomes are determined by salinity
and depth
Land Biomes
In the tropics: Northern Coniferous
Rain forest-200-400 forest (Taiga)-30-100
cm of rain annually cm of rain annually
Savanna-30-50 cm of Tundra-20-60 cm of
rain annually rain annually
In the temperate zone: Desert-occurs at any
Deciduous forest-70- latitude-less than 30
200 cm of rain cm of rain annually
annually
Grassland-30-100 cm
of rain annually
Chaparal-30-50 cm of
rain annually (coastal
region)
Figure 50-9
Barrow
Dawson
Chicago
Konza Prairie
Yuma
Belm
Figure 50-23
Low angle of
incoming sunlight
Moderate angle of
incoming sunlight
Sunlight directly
overhead
Large amount of
sunlight per unit area
Figure 54-11
Organic
matter
Organic
matter
Figure 50-12
Cascade
Mountains This area is in
a rain shadow
Figure 50-14
Bogs are stagnant and acidic. Marshes have nonwoody plants. Swamps have trees and shrubs.
Figure 50-3
Littoral
Limnetic
zo n e
zone
Photic
zone
Be
Aphotic
nt
h
zone
ic
zo
ne
Primary Productivity
This is the amount of light energy
converted to chemical energy in an
ecosystem.
Gross Primary productivity-is the total
primary productivity of the ecosystem
Net Primary productivity-is the gross
primary productivity- the energy used by
producers for respiration (R)
Formula for net primary productivity:
NPP=GPP-R
Figure 54-3
Aquatic
Terrestrial
Human Impact on the Biosphere
Human activity over time has been damaging to the
biosphere.
Ecology is really a study of balance, and as the
human population has grown, our wastes and
byproducts have thrown off that balance.
There have been, in recent years, efforts to
reestablish the balance and conserve our resources.
Ex: Prior to the 1980s, CFCs were common
propellants used in household goods. As a result of
human use of CFCs, they built up in the atmosphere,
reacting with ozone (O3), and causing holes in the
ozone layer. This layer surrounds the planet and
shields us from damaging UV radiation. As a result
of banning CFCs, and using safer alternatives, the
holes in the ozone layer are repairing themselves.
Climate change
Human activities have caused a buildup of
CO2 in the atmosphere. Of course, CO2 is
a greenhouse gas, which means that we
have additional heat building up in our
atmosphere
Data indicates that the world is getting
warmer, which sparks many problems like
raising sea levels, changing weather
patterns that could decrease agricultural
output, change the trophic structure of our
oceans and land
Figure 54-15b
Cold-water copepods are declining in the North Atlantic. Flowering times for some species in midwestern
North America are earlier in the year.
Great
Britain
Cold-water Warm-water
copepods copepods
Baptista flowers
Figure 54-21
Much of the ocean is stratified by density and temperature. Global warming increases the density gradient, making it
less likely for layers to mix.
Surface layer: Water is warm, less dense Surface layer: Water is much warmer, less dense
Benthic zone: Water is 4C, highest density Benthic zone: Water is 4C, highest density
Pollution
Obviously, this causes destruction of water, air, and
land resources, as they become fouled by waste.
This has obvious negative side-effects for orgnisms on
the planet, but there are 2 important issues
associated with pollution that arent always discussed.
Biological magnification-while energy decreases as it
moves up the food chain, pollutants and toxins, like
DDT, concentrate.
Eutrophication-believe it or not, this IS a bad thing.
Over-fertilization nourishes the water, causing algal
blooms. Not only can these be toxic to animals, but
they are r-strategists, so they die, sink, and are
decomposed at the bottom of bodies of water (mainly
freshwater). Decomposition is an oxygen-consuming
process, which leaves the bottom waters anoxic,
causing fish-kills
Other Environmental Problems
Acid Rain-Sulphur-containing compounds
belched from smokestacks turn into
sulphuric acid in the atmosphere
Deforestation-clear-cutting of forests for
logging and expanding human population.
This is particularly bad b/c the nutrients in
tropical rain forests are in the canope.
Desertification-overgrazing of grasslands
bordering deserts turns these into deserts.
All of these can lead to endangerment of or
reduction in species and biodiversity.
Figure 55-10
1975 2001
Figure 55-6
Terrestrial
Freshwater
Marine
Sustainable Practices
These are things that humans can do
to reduce our impact on the
biosphere.
Examples include: reforestation,
smoke-stack filters, reduction of fossil
fuel consumption, smart use of
fertilizers, plowing and strip cropping
to reduce erosion, use of biological
methods for controling pests,
establishing protected areas, etc