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Thorsten Wohland Dep. Of Chemistry S8-03-06 Tel.: 6516 1248 E-mail: chmwt@nus.edu.sg
Seeing the light, Fig. 10.11
Textbook
Color vision: Perspective from different disciplines, Backhaus Light Vision Color, A. Valberg
Spatial summation
Illuminate spots on the retina of different size and determine the number of photons needed before the spot can be seen
T.N. Cornsweet, Fig. 2.5 Sensitivity constant Sensitivity decreases 1st spot: only few rods on average 2nd spot: smaller than summation area 3rd spot: larger than summation area
Temporal summation
How many photons have to arrive in a certain time interval so that the eye sees a flash?
time (ms) 0 10 20
Lateral Inhibition
One ganglion cell receives signal from many receptors, excitatory or inhibitory signals.
+- - -- - + - - - -- - +- - + --
One cone/rod can contribute to some ganglion cells excitatory to others inhibitory.
Lateral Inhibition
rest excitation inhibition No difference -> rest Strong excitation No difference -> rest STL Fig. 7.12
Lateral Inhibition
Afterimages
You can have negative and positive afterimages. The effect comes from the fact that when a cone/rod is stimulated for a long time it desensitizes.
1) The cones perceiving the black square are not excited, the cones perceiving the white surrounding are excited and desensitize with time. 2) When looking at the white surface on the right, the desensitized cones are less excited than the rested cones in the middle and thus you see a white square.
No image
The exposed cones are desensitized, give lower signal than surrounding rested cones.
Afterimages
Positive afterimages. You can sensitize your retina by closing your eyes and resting your cones (remember when you close eyes a long time and open them you seem to be blinded first). When you open your eyes shortly (seconds) and look at some bright object the cones get excited. When you close your eyes again the cones will not desensitize and will stay stimulated longer and give you a positive afterimage.
Now lets recall what we know about the CIE system and then lets see whether there are any facts left unexplained. Can we perhaps resolve some of these issues with our new knowledge of the retina and its organization?
Take one cone; shine light of constant intensity on the cone; measure the light transmitted; calculate absorption
Color mixing
Sensitivity
1.0 P1 x P2 P3 If P1, P2, and x have same intensity we have too much red. Since P3 excites the red cone 4 times less P2, we can subtract 4 times P3 to get our mixture: x = P1 + P 2 4 P 3
x: equal excitation of blue and green cone by 30%, no excitation of red P1: excitation of blue cone by 30%, no excitation of green and red P2: excitation of green cone by 30%, excitation of red by 80%, no excitation of blue P3: excitation only of red by 20%.
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Complementarity
Do all spectral hues have a complementary spectral hue?
Hue discrimination
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STL, Fig. 10.9 Experiment done by asking a person to estimate how much blue, yellow, green, and red is contained in a hue represented by a pure wavelength.
Hue cancellation
Opponent processing
Can we connect the cones in a fashion, so that the signal at the ganglion cells will correspond to the four opponent colors red, green, blue, and yellow?
Possible combinations
S M + L L M
Perceived Brightness
Red-Green
Possible combinations
M S +
Yellow
L
So we have constructed 3 new signals from the original three cones: 1. Black White 2. Green Red 3. Blue Yellow
based on 4 colors
Blue - Yellow
Double opponency
Opponency of location (inside versus outside)
One ganglion cell receives signal from many receptors, excitatory or inhibitory signals.
Opponency of color
Double Opponency
1. Individual L and M cone signals are pooled by a ganglion cell to give a Red-Green opponent signal
L -
M -
Red - Green
Green - Red
2. Depending on the position of the cones on your retina the Red-Green opponent signal can work as excitatory (+) some as inhibitory (-) signals.
+ +- + + +
Temporal Processing
Benham disk: White parts excite all three cones. However, the three cones recover from activation differently. When black falls onto the excited cones, some are still stimulated (e.g. the blue one) while others (red and green) have already recovered. Thus one sees blue.
Temporal Processing
Cones No excitation, no color perception Flash of white light All cone excited, white is seen Red cone de-excites fast, a blue/green (cyan) color is seen Green cone de-excites next. Blue is seen
Summary
Color Perception Mechanisms Tristimulus Theory Color naming, hue cancellation Opponent processing Spatial Processing of Color Temporal Processing of Color